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Mateus, O. (1998).  Serão as aves dinossauros?. CiênciaJ. 6, 5. Abstract
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Mateus, O., & Antunes M. T. (2000).  On the presence of Ceratosaurus sp. (Dinosauria: Theropoda) in the Late Jurassic of Portugal. Abstract volume of the 31st International Geological Congress. , Rio de Janeiro, Brazil Abstract
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Mateus, O., Jacobs L. L., Polcyn M. J., Schulp A. S., Neto A. B., & Antunes M. T. (2008).  Dinosaur and turtles from the Turonian of Iembe, Angola. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontologia de Vertebrados. 156., Neuquén, Argentina Abstractmateus_et_al_2008_dinosaur_and_turtles_from_the_turonian_of_iembe_angola.pdf

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Mateus, O. (2014).  Degradation processes and consolidation of Late Jurassic sandstone dinosaur tracks in museum environment (Museum of Lourinhã, Portugal). Geophysical Research Abstracts. Geophysical Research Abstracts, EGU2014–9026–1, 2014., 1 Abstract
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Mateus, I., Mateus H., Antunes M. T., Mateus O., Taquet P., Ribeiro V., & Manuppella G. (1997).  Couvée, oeufs et embryons d'un dinosaure théropode du Jurassique supérieur de Lourinhã (Portugal). Comptes Rendus de l'Académie des Sciences-Series IIA-Earth and Planetary Science. 325, 71–78., Number 1 Abstract
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Mateus, O., Polcyn M. J., Jacobs L. L., Araújo R., Schulp A. S., Marinheiro J., Pereira B., & Vineyard D. (2012).  Cretaceous amniotes from Angola: dinosaurs, pterosaurs, mosasaurs, plesiosaurs, and turtles. V Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno. 71-105., Salas de los Infantes, Burgos Abstractmateus_et_al_2012_amniotes_from_angola_cretaceous_amniotes_from_angola_dinosaurs_pterosaurs_mosasaurs.pdf

Although rich in Cretaceous vertebrate fossils, prior to 2005 the amniote fossil record of Angola was poorly known. Two horizons and localities have yielded the majority of the vertebrate fossils collected thus far; the Turonian Itombe Formation of Iembe in Bengo Province and the Maastrichtian Mocuio Formation of Bentiaba in Namibe Province. Amniotes of the Mesozoic of Angola are currently restricted to the Cretaceous and include eucryptodire turtles, plesiosaurs, mosasaurs, pterosaurs, and dinosaurs. Recent collecting efforts have greatly expanded our knowledge of the amniote fauna of Angola and most of the taxa reported here were unknown prior to 2005.

Mateus, O., Butler R. J., Brusatte S. L., Whiteside J. H., & Steyer J. S. (2014).  The first phytosaur (Diapsida, Archosauriformes) from the Late Triassic of the Iberian Peninsula. Journal of Vertebrate Paleontology. 34, 970-975., Number 4 Abstract
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Mateus, O. (2010).  Physical drivers of evolution and the history of the marine tetrapod fauna of Angola. –, , 1 Abstract

Modern marine species populations are often evaluated in terms of bottom-up, resource limited structure, or top-down, predator controlled structure. In a larger timeframe, investiga- tion of physical drivers in marine tetrapod evolution relies on the recognition of patterns and the correlation in timing of physical events with biotic change. However, it has been dem- onstrated through the study of fossil cetaceans that a broader deep-time perspective within a top-down or bottom-up framework is informative. Here we examine the fossil record of &UHWDFHRXV PDULQH WHWUDSRGV LQ $QJROD WR GLVFHUQ SDWWHUQV WKDW PD\ UHÀHFW SK\VLFDO GULYHUV RI evolution, and that are also relevant to population structure. In modern marine ecosystems, GLVWULEXWLRQ SDWWHUQV UHÀHFWLQJ SULPDU\ SURGXFWLYLW\ DUH LQGLFDWLYH RI ERWWRP?XS FRQWURO? ,Q the fossil record, productivity-controlled distribution patterns can also be perceived. Physi- cal parameters resulting in environmental stability, sea-level change, oceanic anoxic events, paleoclimate, and paleogeography are examined in comparison with taxonomic diversity and life history patterns. Mosasaurs originated during a time of high global temperatures and shallow temperature gradients. As upper-trophic-level species of modest size and plesiopedal limb structure (capable of terrestrial locomotion), early mosasaurs were subject to both top- down and bottom up pressures. The attainment of larger size coupled with emigration and biogeographic distribution in areas of high primary productivity, and niche differentiation VKRZQ E\ 13C values, indicate bottom-up pressures. Productivity along the African coast since the formation of the Atlantic Ocean facilitated the co-occurrence of diverse marine tetrapods through time, and has culminated today in the Benguela large marine ecosystem. Just as the current Benguela ecosystem has tetrapod species populations dominated by both bottom-up (cetaceans) and top-down strategies (sea birds and pinnipeds), so too did the Cre- taceous community, with mosasaurs and plesiosaurs having predominantly bottom-up popu- lation structure, while sea turtles and pterosaurs were more subject to top-down pressures.

Mateus, O. (2012).  Age and paleoecology of mosasaurs and plesiosaurs from the Late Cretaceous South Atlantic margin at Bentiaba, Angola. Journal of Vertebrate Paleontology, Program and Abstracts, 2012. 180.. 180–181., 1 Abstract
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Mateus, O., Jacobs L., Polcyn M., Schulp A. S., Vineyard D., Buta Neto A., & Telles Antunes M. (2009).  The oldest African eucryptodiran turtle from the Cretaceous of Angola. Acta Palaeontologica Polonica. 54, 581-588., Number 4 Abstract
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Mateus, O. (2010).  Paleontological collections of the Museum of Lourinhã (Portugal). (Brandao, JM, Callapez, PM, O. Mateus, Castro, P, Ed.).Colecções e museus de Geologia: missão e gestão. 121-126., Jan: Ed. Universidade de Coimbra e Centro de Estudos e Filosofia da História da Ciência Coimbra Abstractmateus_2010_paleontological_collections_of_the_museum_of_lourinha__geocoleccoes_omateus.pdf

Abstract: The paleontological collections of the Museum of Lourinhã, in Portugal, has a rich paleontological collection, particularly of Late Jurassic dinosaurs of the Lourinhã Formation (Kimmeridgian-Tithonian). Most salient highlights comprehend the following dinosaur holotype specimens: stegosaur Miragaia longicollum, theropod Lourinhanosaurus antunesi, sauropod Dinheirosaurus lourinhanensis, ornithopod Draconyx loureiroi, theropod Allosaurus europaeus, and, a mammal, Kuehneodon hahni. Other dinosaur specimens are referred including the nest and eggs and embryos of Lourinhanosaurus. Portugal is very productive in Late Jurassic vertebrates, being the seventh country bearing more dinosaur taxa.

Mateus, O., & Castanhinha R. (2008).  PaleoAngola- Predadores de um oceano primitivo. National Geographic Portugal. 8, 26-33., Number 91 Abstract
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Mateus, O., Dyke G., Motchurova-Dekova N., Ivanov P., & Kamenov G. D. (2008).  The Bulgarian dinosaur: did it exist? European late Cretaceous ornithomimosaurs. 56th Symposium of Vertebrate Palaeontology and Comparative Anatomy. 47., Dublin Abstract

For historical and geological reasons the dinosaurian fossil record from central Europe is little known. Here we describe and interpret a portion of a left humerus from the Upper Maastrichtian of Vratsa district in north-western Bulgaria. This bone is the first known record of a dinosaur from Bulgaria; it is certainly a theropod, probably an ornithomimosaur. We discuss the fossil record of other similar fossils of theropod dinosaurs, in particular other problematic remains from the Maastrichtian of Belgium. Rare Earth Element (REE) analysis combined with strontium (Sr) isotope data demonstrate that the Bulgarian dinosaur was initially fossilised in a terrestrial environment and then later re-worked into Late Maastrichtian marine sediments.

Mateus, O. (2014).  Comparison of modern and fossil Crocodylomorpha eggs and contribution to the oophylogeny of Amniota. Annual Meeting of the European Association of Vertebrate Palaeontologists. , p. 192, Regione Piemonte: European Association of Vertebrate Palaeontologists. Museo Regionale di Scienze Naturali. Abstract
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Mateus, O., Taquet P., Antunes M. T., Mateus H., & Ribeiro V. (1998).  Theropod dinosaur nest from Lourinhã, Portugal. Journal of Vertebrate Paleontology. 18, 61., Number (Suppl. 3) Abstract
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Mateus, O., Butler R. J., Brusatte S. L., Whiteside J. H., & Steyer S. J. (2014).  The first phytosaur (Diapsida, Archosauriformes) from the Late Triassic of the Iberian Peninsula. Journal of Vertebrate Paleontology. 34(4), 970-975.mateus_et_al_2014_first_phytosaur_algarve_portugal_jvp.pdfWebsite
Mateus, O., Araújo R., Natário C., & Castanhinha R. (2011).  A new specimen of the theropod dinosaur Baryonyx from the early Cretaceous of Portugal and taxonomic validity of Suchosaurus. Zootaxa. 54-68., Number 2827 Abstract
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Mateus, O., Jacobs L., Polcyn M., Schulp A. S., Vineyard D., Neto A. B., & Antunes M. T. (2009).  The oldest African eucryptodiran turtle from the Cretaceous of Angola. Acta Palaeontologica Polonica. 54, 581-588., Jan: Univ Agostinho Neto, Univ Nova Lisboa, Museu Lourinha, Acad Ciencias Lisboa, Nat Hist Museum Maastricht, So Methodist Univ Abstractmateus_et_al_2009_the_oldest_african_angolachelys_angola_turtle.pdfWebsite

A new Late Cretaceous turtle, Angolachelys mbaxi gen. et sp. nov., from the Turonian (90 Mya) of Angola, represents the oldest eucryptodire from Africa. Phylogenetic analysis recovers Angolachelys mbaxi as the sister taxon of Sandownia harrisi from the Aptian of Isle of Wight, England. An unnamed turtle from the Albian Glen Rose Formation of Texas (USA) and the Kimmeridgian turtle Solnhofia parsonsi (Germany), are successively more distant sister taxa. Bootstrap analysis suggests those four taxa together form a previously unrecognized monophyletic clade of marine turtles, herein named Angolachelonia clade nov., supported by the following synapomorphies: mandibular articulation of quadrate aligned with or posterior to the occiput, and basisphenoid not visible or visibility greatly reduced in ventral view. Basal eucryptodires and angolachelonians originated in the northern hemisphere, thus Angolachelys represents one of the first marine amniote lineages to have invaded the South Atlantic after separation of Africa and South America.

Mateus, O., & The Gigantic dinosaur E. (2006).  The European Enigmatic Dinosaur Evolution (in Japanese). Abstract
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Mateus, O. (1998).  Dinossauros de Portugal e um novo terópode do Jurássico Superior da Lourinhã. , Évora: Universidade de Évora Abstract
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Mateus, O., & Antunes M. T. (2002).  Novos vestígios de dinossauros saurópodes do Jurássico Superior da Lourinhã (Portugal). Congresso Ibérico de Herpetologia. , Évora Abstract
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Mateus, O. (1998).  Dinossauros Portugueses. Caderno de resumos do I Congresso de Estudantes de Biologia. 13–13., Évora Abstract
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Mateus, O. (2009).  The Cretaceous Skeleton Coast of Angola. Journal of Vertebrate Paleontology. 29, 121A., 1, Number 3: Taylor & Francis Abstract

THE CRETACEOUS SKELETON COAST OF ANGOLA JACOBS, Louis, SMU, Dallas, TX, USA; POLCYN, Michael, SMU, Dallas, TX, USA; MATEUS, Octávio, Museu da Lourinhã, Lourinhã, Portugal; SCHULP, Anne, Natuurhistorisch Museum Maastricht, Maastricht, Netherlands; NETO, André , Universidade Agostinho Neto, Luanda, Angola Cretaceous coastal sediments of Angola present a rich and diverse fauna of marine amniotes, including turtles, mosasaurs, and plesiosaurs. The abundance of mosasaurs in particular suggests a highly productive coastal area. Angola today lies at the northern limit of the Namibian Desert, the so-called Skeleton Coast, which results from prevailing southeasterly winds of the descending limb of the southern Hadley Cell sweeping across the African coast. The Benguela upwelling and a highly productive sea are found today off the Namibian Desert coast. However, the Benguela upwelling system, based on results of DSDP studies, is said to have originated in the late Neogene and therefore cannot explain the productivity found along the length of the West African coast. The explanation is found in the northward drift of Africa through the arid climate zone, and is demonstrated by the tracing of the paleogeographic position of fossil localities through time.

Mateus, O. (2011).  New fossil whales from Angola. Abstracts of the 71st Annual Meeting of the Society of Vertebrate Paleontology. 119., 1 Abstract
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Mateus, O., & Campos H. (2018).  Loulé há mais de 220 Milhões de anos: os vertebrados fósseis do Algarve triásico. Loulé: Territórios. Memórias. Identidades. 651-659.: Museu Nacional de Arqueologia | Imprensa Nacionalmateus_campos2018_algarve_triasico.pdf
Mateus, O., Jacobs L. L., Polcyn M. J., Schulp A. S., Neto A. B., & Antunes M. T. (2008).  Dinosaur and turtles from the Turonian of Iembe, Angola. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 156., Neuquén, Argentina Abstract
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Mateus, O., & Milan J. (2010).  A diverse Upper Jurassic dinosaur ichnofauna from central-west Portugal. Lethaia. 43, 245–257., Jan Abstractmateus__milan_2010_-_diverse_l_j_ichnofauna_from_lourinha_fm_portugal.pdfWebsite

A newly discovered dinosaur track-assemblage from the Upper Jurassic Lourinha˜ Formation (Lusitanian Basin, central-west Portugal), comprises medium- to large-sized sauropod tracks with well-preserved impressions of soft tissue anatomy, stegosaur tracks and tracks from medium- to large-sized theropods. The 400-m-thick Lourinha˜ Formation consists of mostly aluvial sediments, deposited during the early rifting of the Atlantic Ocean in the Kimmeridgian and Tithonian. The stratigraphic succession shows several shifts between flood-plain mud and fluvial sands that favour preservation and fossilization of tracks. The studied track-assemblage is found preserved as natural casts on the underside of a thin bivalve-rich carbonate bed near the Tithonian–Kimmeridgian boundary. The diversity of the tracks from the new track assemblage is compared with similar faunas from the Upper Jurassic of Asturias, Spain and the Middle Jurassic Yorkshire Coast of England. The Portuguese record of Upper Jurassic dinosaur body fossils show close similarity to the track fauna from the Lourinha˜ Formation.

Mateus, O., & Jacinto J. J. (2002).  Contribuição para o estudo de Hemidactylus turcicus (Reptilia, Gekkonidae): ritmos de actividade e microhabitat em Évora, Portugal. VII Congresso Luso-Espanhol de Herpetologia. 136., Évora Abstract

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Mateus, O., & Jacinto J. J. (2002).  Contribuição para o estudo de Hemidactylus turcicus (Reptilia, Gekkonidae): ritmos de actividade e microhabitat em Évora, Portugal. VII Congresso Luso-Espanhol de Herpetologia. 136., Évora Abstract
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Mateus, O., & Milan J. (2008).  Ichnological evidence for giant ornithopod dinosaurs in the Upper Jurassic Lourinhã Formation, Portugal. Oryctos. 8, 47-52. Abstractmateus_and_milan_2008_ichnological_evidence_for_giant_ornithopod_big_ornithopod_track_from_u_j_lourinha_fm_portugal.pdfWebsite

The Upper Jurassic Lourinhã Formation (Lusitanian Basin, Portugal) contains a diverse dinosaur fauna comprising theropods, sauropods, stegosaurs, ankylosaurs and several genera of ornithopods. The sedimentology in the area favours preservation of tracksways, and tracks from most of the dinosaurs are also represented by skeletal remains. During fieldwork in the summer of 2003 a new, large, tridactyl track was found at the beach of Vale Frades, approximately 6 km north of Lourinhã (central west Portugal). The track was found together with a stegosaur track on a clay bed exposed within the intertidal zone. Due to the immediate danger of erosion, the track was collected and is now on display at Museu da Lourinhã. The track is 70 cm long and 69 cm wide, the toes are short and broad, with indications of short blunt claws, and there is a high angle of divarication between the outer digits. The shape and dimensions of the track identifies it as deriving from an ornithopod dinosaur with an estimated hip height around three metres. Although very large ornithopods are known from the Cretaceous, the largest known Jurassic ornithopod is Camptosaurus from North America, and the largest known from Portugal is the camptosaurid Draconyx loureiroi. Neither of these reached the body size suggested by the new track. So far the track described herein is the only evidence for a Jurassic ornithopod of that size.

Mateus, O., & Jacinto J. J. (1998).  Activity Rithms and habitat of Hemidactylus turcicus (Reptilia, Gekkonidae) in Évora, Portugal. Boletin de ICIJA. 2, 37-43. Abstract
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Mateus, O., Walen A., & Antunes M. T. (2006).  The large theropod fauna of the Lourinhã Formation (Portugal) and its similarity to the Morrison Formation, with a description of a new species of Allosaurus. New Mexico Museum of Natural History and Science Bulletin. 36, 123-129. Abstractmateus_walen_antunes_-_2006_-_the_large_theropod_fauna_of_the_lourinha_formation__portugal__and_its_similarity_to_the_morrison_formation__with_a_description_of_a_new_species_of_allosaurus.pdf

Late Jurassic theropod dinosaurs have been known in Portugal since 1863 but only now are they being fully understood, with the recognition of genera such as Allosaurus, Aviatyrannis, Ceratosaurus, Lourinhanosaurus, and Torvosaurus from the Lourinhã and Alcobaça Formations (Kimmeridgian/Tithonian). Ceratosaurus dentisulcatus can now be reported from Portugal. It represents the only occurrence of this species outside the Morrison Formation.
New cranial elements confirm the presence of Torvosaurus tanneri, in Portugal. Torvosaurus was the largest Late Jurassic land carnivore. New postcranial and cranial elements allow the erection of a new species from Portugal, Allosaurus europaeus n.sp. The theropod assemblage of Portugal is similar to that of the Morrison Formation.

Mateus, O. (2014).  Eggs and eggshells of crocodylomorpha from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology. Program and Abstracts, 2014, 218., Number NA: Taylor & Francis Abstract
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Mateus, O. (2013).  Crocodylomorphs from the Mesozoic of Portugal and a new skull of eusuchian from the Late Cretaceous. 2013 Hwaseong International Dinosaurs Expedition Symposium, pp.66-67.. , Hwaseong, South Korea Abstractmateus_2013_crocodylomorphs_portugal_new_skull.pdf

The diversity of fossil crocodylomorphs in Portugal is high, with occurrence as old as Mystriosaurus (=Steneosaurus) bollensis from the Lower Jurassic. The Late Jurassic forms are the better documented, and include the following taxa: Machimosaurus hugii, Lisboasaurus estesi Seiffert, 1973, Lusitanisuchus mitrocostatus Seiffert, 1975; Schwarz & Fechner 2004, Theriosuchus guimarotae Schwarz and Salisbury 2005, Cf. Alligatorium, Goniopholis baryglyphaeus, and a crocodylomorph-like eggs in dinosaur nests (Mateus et al., 1998; Ricqlès et al., 2001). From the Lower Cretaceous were reported a few dinosaurs but its record is strangely scarce in crocodylomorphs (Mateus et al., 2011). The Upper Cretaceous crocodiles show a large diversity, but it is mostly based in fragmentary material that require revision, such as “Crocodylus” blavieri? Grey from the Upper Campanian - Maastrichtian of Viso, near Aveiro (initially reported by Sauvage 1897-98), Goniopholis cf. crassidens Owen 1841 and Oweniasuchus pulchelus Jonet 1981. Moreover there is a fascinating, but poorly understood, crocodylomorph diversity in the Cenomanian of Portugal, documented by fragmentary specimens that have been doubtfully assigned to Thoracosaurus Leidy 1852 of the Middle Cenomanian of Cacém, to the nomen dubium Oweniasuchus lusitanicus Sauvage 1897-98 (interpreted as a mesosuchian goniopholid) based in a fragmentary mandible from the Campanian-Maastrichtian, and also from the Middle Cenomanian of Portugal, Buffetaut and Lauverjat (1978) report an fragmentary unidentified possible dyrosaurid from Nazaré. All this specimens are too incomplete to be compared with the specimen here described. In contrast, Cenozoic crocodiles of Portugal are often known after complete skulls and several individuals. The taxa list include Iberosuchus macrodon (Lower to Middle Eocene), Tomistoma calaritanus (Early Miocene) and T. lusitanica (Burdigalian-Helvetian), and Diplocynodon sp. (Antunes, 1961, 1987, 1994).
At least, two different morphotypes of crocodylomorph eggs from the Late Jurassic of Lourinhã Formation are also known.
A new specimen here reported of crocodile based in a partial skull and mandible (ML1818) from the Uppermost Middle Cenomanian platform carbonates of Baixo Mondego, west central Portugal (Tentúgal Fm., Callapez, 2004). The taxon is phylogenetically positioned as a basal Eusuchia, due to the choanae enclosed by the pterygoid, and closely related with stem Crocodylia and Borealosuchus. This specimen represents the only well documented and valid eusuchian species in the Cenomanian of Europe and is the oldest representative of an eusuchian crocodylomorph, with the exception for the Barremian Hylaeochampsa vectiana.

Mateus, O. (2013).  Cathetosaurus as a valid sauropod genus and comparisons with Camarasaurus. Journal of Vertebrate Paleontology. 173., 1 Abstract
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Mateus, O. (2016).  Exemplos bizarros de evolução em dinossauros e alguns casos portugueses. Do Big Bang ao Homem. 81-95., Porto: U.Porto Ediçõesmateus_2016_capitulo_livro_dinosaurs.pdf
Mateus, O. (2009).  New specimens of Angolasaurus bocagei and comments on the early radiations of plioplatecarpine mosasaurs. Journal of Vertebrate Paleontology. Journal of Vertebrate Paleontology. 29, 165A., 1, Number 3: Taylor & Francis Abstract

NEW SPECIMENS OF ANGOLASAURUS BOCAGEI AND COMMENTS ON THE EARLY RADIATIONS OF PLIOPLATECARPINE MOSASAURS POLCYN, Michael, SMU, Dallas, TX, USA; JACOBS, Louis, SMU, Dallas, TX, USA; MATEUS, Octávio, Museu da Lourinhã, Lourinhã, Portugal; SCHULP, Anne, Natuurhistorisch Museum Maastricht, Maastricht, Netherlands New, well preserved material of the Turonian mosasaur Angolasaurus bocagei from the Tadi Beds of the Itombe Formation in northern Angola, allows detailed redescription of its morphology and reassessment of its phylogenetic relationships. Angolasaurus had been previously referred to the genus Platecarpus; however, phylogenetic analysis confirms the valid taxonomic status of A. bocagei, and reconstructs that taxon within a clade that also includes the genera Selmasaurus and Ectenosaurus. These forms are united by an elaborated infrastapedial process of the quadrate and a unique ridge-like descending process of the parietal forming the supraoccipital articulation, but also retain a relatively plesiomorphic configuration of the braincase. That clade is united with all other plioplatecarpines by a number of derived characters including the presence of a novel basicranial circulation pattern. In Africa, North and South America, early plioplatecarpines are known by the Middle Turonian and Angolasaurus and closely related forms appear by the Upper Turonian. Selmasaurus and Ectenosaurus are a rare faunal component of the Santonian and Campanian of North America. Platecarpus planifrons appears in the Coniacian of North America and represents the plesiomorphic condition of the clade containing the remaining species of Platecarpus and Plioplatecarpus, that appears in the Santonian and persist until the end of the Cretaceous, reaching global distribution. The temporal and geographic distribution of these radiations suggest influence of paleogeography and eustatic sea levels.

Mateus, O. (2008).  Checklist for Late Jurassic reptiles and amphibians from Portugal. Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 55., Coimbra Abstract
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Mateus, O., Araújo R., Natário C., & Castanhinha R. (2011).  A new specimen of the theropod dinosaur Baryonyx from the early Cretaceous of Portugal and taxonomic validity of Suchosaurus. Zootaxa. 2827, 54–68., Jan Abstractmateus_et_al_2011_a_new_specimen_of_the_theropod_dinosaur_baryonyx_from_the_early_cretaceous_of_portugal_and_taxonomic_validity_of_suchosaurus.pdf

Although the Late Jurassic of Portugal has provided abundant dinosaur fossils, material from the Early Cretaceous is scarce. This paper reports new cranial and postcranial material of the theropod dinosaur Baryonyx walkeri found in the Barremian (Papo Seco Formation) of Portugal. This specimen, found at Praia das Aguncheiras, Cabo Espichel, consists of a partial dentary, isolated teeth, pedal ungual, two calcanea, presacral and caudal vertebrae, fragmentary pubis, scapula, and rib fragments. It represents the most complete spinosaurid yet discovered in the Iberian Peninsula and the most complete dinosaur from the Early Cretaceous of Portugal. This specimen is confidently identified as a member of Baryonychinae due to the presence of conical teeth with flutes and denticles in a dentary rosette. The specimen ML1190 shares the following characteristics with Baryonyx walkeri: enamel surface with small (nearly vertical) wrinkles, variable denticle size along the carinae, 6–7 denticles per mm, wrinkles forming a 45 degree angle near the carinae, and tooth root longer than crown. In addition, dubious taxa based on teeth morphology such as Suchosaurus cultridens (Owen, 1840–1845), and Suchosaurus girardi (Sauvage 1897–98; Antunes & Mateus 2003) are discussed, based on comparisons with well-known material such as Baryonyx walkeri Charig & Milner, 1986. Suchosaurus cultridens and S. girardi are considered as nomina dubia due to the lack of diagnostic apomorphies, but both specimens are referred to Baryonychinae incertae sedis.

Mateus, O. (2008).   Fósseis de transição, elos perdidos, fósseis vivos e espécies estáveis . Evolução: História e Argumentos. 77-96., Lisboa Abstract
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