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Adams, T. L., Polcyn M. J., Mateus O., Winkler D. A., & Jacobs L. L. (2011).  First occurrence of the long-snouted crocodyliform Terminonaris (Pholidosauridae) from the Woodbine Formation (Cenomanian) of Texas. Journal of Vertebrate Paleontology. 31, 712-716., Number 3 Abstract
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Adams, T. L., Polcyn M. J., Mateus O., Winkler D. A., & Jacobs L. L. (2010).  New occurrence of the long-snouted crocodyliform,Terminonaris cf.T. robusta, from Woodine Formation (Cenomanian) ot Texas. 52A., Jan Abstract

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Adams, T. L., Polcyn M. J., Mateus O., Winkler D. A., & Jacobs L. L. (2011).  First occurrence of the long-snouted crocodyliform Terminonaris (Pholidosauridae) from the Woodbine Formation (Cenomanian) of Texas. Journal of Vertebrate Paleontology. 31, 712-716., Jan: So Methodist Univ, Univ Nova Lisboa Abstractadams_polcyn_mateus_et_al_2011_terminonaris_crocodile_pholidosauridae.pdf

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Agnolin, F. L., Mateus O., Milàn J., Marzola M., Wings O., Adolfssen J. S., & Clemmensen L. B. (2018).  Ceratodus tunuensis, sp. nov., a new lungfish (Sarcopterygii, Dipnoi) from the Upper Triassic of central East Greenland. Journal of Vertebrate PaleontologyJournal of Vertebrate Paleontology. e1439834., 2018: Taylor & Francis Abstractagnolin_et_al_2018_ceratodus_tunuensis_greenland.pdfWebsite

ABSTRACTThe fossil record of post-Paleozoic lungfishes in Greenland is currently restricted to a few brief reports of isolated and undetermined tooth plates coming from the uppermost Fleming Fjord Formation (late Norian) in Jameson Land, central East Greenland. Here, we describe Ceratodus tunuensis, sp. nov., a new dipnoan from a thin bed of calcareous lake mudstone from the ?rsted Dal Member of the Fleming Fjord Formation. The Ceratodus fossil record indicates that during the Late Triassic, this genus was restricted to the middle latitudes of the Northern Hemisphere. This record matches previous paleobiogeographical analyses and indicates that terrestrial biota during the Late Triassic was strongly influenced by paleolatitude.Citation for this article: Agnolin, F. L., O. Mateus, J. Milàn, M. Marzola, O. Wings, J. Schulz Adolfssen, and L. B. Clemmensen. 2018. Ceratodus tunuensis, sp. nov., a new lungfish (Sarcopterygii, Dipnoi) from the Upper Triassic of central East Greenland. Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2018.1439834.

Agnolin, F. L., Mateus O., Milàn J., Marzola M., Wings O., Adolfssen J. S., & Clemmensen L. B. (2018).  Ceratodus tunuensis, sp. nov., a new lungfish (Sarcopterygii, Dipnoi) from the Upper Triassic of central East Greenland. Journal of Vertebrate Paleontology. e1439834., apr: Informa {UK} Limited AbstractWebsite
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Allain, R., Taquet P., Battail B., Dejax J., Richir P., Veran M., Limon-Duparcmeur F., Vacant R., Mateus O., Sayarath P., Khenthavong B., & Phouyavong S. (1999).  Un nouveau genre de dinosaure sauropode de la formation des Gres superieurs (Aptien-Albien) du Laos. Comptes Rendus de l'Academie des Sciences - Series IIA - Earth and Planetary Science. 329, 609-616., oct, Number 8 Abstractallain_taquet_battail_dejax_richir_mateus_et_al_1999_un_nouveau_genre_de_dinosaure_sauropode_de_la_formation_des_gres_superieurs_aptien-albien_du_laos.pdfWebsite

The partly-articulated postcranial remains of two sauropod skeletons recently found in Tang Vay (Savannakhet Province, Laos) are assigned to the species Tangvayosaurus hoffeti (nov. gen., nov. sp.). The derived characters present in the new material confirm the presence of titanosaurs in South East Asia at the end of the Early Cretaceous, but are not consistent with its placement within Titanosaurus genus as first done by Hoffet in 1942. All of the material relative to this species is therefore referred to a new genus: Tangvayosaurus. Tangvayosaurus and the Thai genus Phuwiangosaurus have strong affinities and are considered as primitive titanosaurs.

Allain, R., Taquet P., Battail B., Dejax J., Richir P., Veran M., Limon-Duparcmeur F., & et al (1999).  A new genus of sauropod dinosaur from the Gres superieurs formation (Aptian-Albian) of Laos. Comptes Rendus de l'Academie des Sciences Series IIA Earth and Planetary Science. 329, 609–616., Number 8 Abstract
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Allain, R., Taquet P., Battail B., Dejax J., Richir P., Véran M., Limon-Duparcmeur F., Vacant R., Mateus O., Sayarath P., Khenthavong B., & Phouyavong S. (1999).  Un nouveau genre de dinosaure sauropode de la formation des Grès supérieurs (Aptien-Albien) du Laos. Comptes Rendus de l{\textquotesingle}Académie des Sciences - Series {IIA} - Earth and Planetary Science. 329, 609–616., oct, Number 8: Elsevier {BV} AbstractWebsite
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Allain, R., Taquet P., Battail B., Dejax J., Richir P., Véran M., Limon-Duparcmeur F., & et al (1999).  Un nouveau genre de dinosaure sauropode de la formation des Grès supérieurs (Aptien-Albien) du Laos. Comptes Rendus de l'Académie des Sciences-Series IIA-Earth and Planetary Science. 329, 609–616., Number 8 Abstract
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Allain, R., Taquet P., Battail B., Dejax J., Richir P., Véran M., Limon-Duparcmeur F., & et al (1999).  A new genus of sauropod dinosaur from the Gres superieurs formation (Aptian-Albian) of Laos | Un nouveau genre de dinosaure sauropode de la formation des Gres superieurs (Aptien-Albien) du Laos. Comptes Rendus de l'Academie de Sciences - Serie IIa: Sciences de la Terre et des Planetes. 329, 609–616., Number 8 Abstract
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Antunes, M. T., & Mateus O. (2003).  Dinosaurs of Portugal. Comptes Rendus Palevol. 2, 77-95., Number 1 Abstractantunes_mateus_2003_dinosaurs_of_portugal.pdfWebsite

A synthesis on the state of art on dinosaur knowledge in Portugal is presented. The following genera have been recognized: Ceratosaurus, Torvosaurus, Lourinhanosaurus, Allosaurus, cf. Compsognathus, Stokesosaurus, cf. Richardoestesia, cf. Archaeopteryx, Euronychodon, cf. Paronychodon, Dinheirosaurus, Lourinhasaurus, Lusotitan, cf. Pleurocoelus, Lusitanosaurus, Dacentrurus, Dracopelta, Phyllodon, Hypsilophodon, Alocodon, Trimucrodon, Draconyx, Iguanodon, and Taveirosaurus. Most are from Late Jurassic localities at the Lourinhã area and Guimarota. A new genus, Lusotitan, is here raised to include the Late Jurassic ‘Brachiosaurus’ atalaiensis. Lower Cretaceous until Cenomanian material is scarce, except for dinosaur footprints. An interesting Late-Cretaceous, mostly small dinosaur association has been collected between Aveiro and Taveiro.

Antunes, M. T., & Mateus O. (2003).  Dinosaurs of Portugal. Comptes Rendus Palevol. 2, 77–95., jan, Number 1: Elsevier {BV} AbstractWebsite
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Araújo, R., Polcyn M. J., Lindgren J., Jacobs L. L., Schulp A. S., Mateus O., Gonçalves O. A., & Morais M. - L. (2015).  New aristonectine elasmosaurid plesiosaur specimens from the Early Maastrichtian of Angola and comments on paedomorphism in plesiosaurs. Netherlands Journal of Geosciences. FirstView, 1–16., 2 Abstractaraujo_et_al_2015_paedomorphism-libre.pdfWebsite

ABSTRACT New elasmosaurid plesiosaur specimens are described from the Early Maastrichtian of Angola. Phylogenetic analyses reconstruct the Angolan taxon as an aristonectine elasmosaurid and the sister taxon of an unnamed form of similar age from New Zealand. Comparisons also indicate a close relationship with an unnamed form previously described from Patagonia. All of these specimens exhibit an ostensibly osteologically immature external morphology, but histological analysis of the Angolan material suggests an adult with paedomorphic traits. By extension, the similarity of the Angolan, New Zealand and Patagonian material indicates that these specimens represent a widespread paedomorphic yet unnamed taxon.

Araújo, R., Mateus O., Walen A., & Christiansen N. (2009).  Preparation techniques applied to a stegosaurian Dinosaur from Portugal. Journal of Paleontological Techniques. 5, 1-24., Jan Abstractarajomateusetal2009.preparationtechn.pdfWebsite

General vertebrate paleontological techniques that have been used in the Museum of Lourinhã (Portugal) are presented here, in particular those applied to a stegosaurian dinosaur skeleton, Miragaia longicollum. A monolith jacket technique using polyurethane foam and plaster is presented. Mechanical preparation
techniques combining the use of an electric grinder and airscribes proved effective during the initial phases of preparation on well-preserved bone embedded in hard matrix. We also present a technique to mould monoliths in the early stages of preparation, creating a thin silicone rubber mould in several contiguous
parts. To mould and cast monoliths before removing individual bones has proven valuable for the preservation of taphonomic data and for display purposes. Polyurethane resin combined with plaster is useful for small casts, while polyester resin applied in four layers is the preferred technique for larger casts.
The four layers are composed of: a first thin layer of polyester resin with bone colour; followed by another layer of polyester resin of sediment colour and containing glass microspheres to make it thicker. The third layer is composed of fibre glass chopped strands, and the fourth is composed of fibre glass mats embedded
in plain polyester resin. 3D scanning and digitization techniques where tested for the storage of osteological information of individual bones and proved very promising.

Araújo, R., Castanhinha R., & Mateus O. (2011).  Evolutionary major trends of ornithopod dinosaurs teeth. (J.Calvo, J. Porfiri, B. Gonzalez Riga y D. Dos Santos (scientific editors), Ed.).Dinosaurios y paleontología desde América Latina. 25-31., Jan: EDIUNC, Editorial de la Universidad Nacional de Cuyo Abstractaraujo_et_al_2011_evolutionary_major_trends_of_ornithopod_dinosaurs_teeth.pdf

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Araujo, R., Castanhinha R., & Mateus O. (2008).  Major trends in the evolution of teeth and mandibles in ornithopod dinosaurs. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 18., Neuquén, Argentina Abstract
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Araújo, R., Polcyn M. J., Schulp A. S., Mateus O., Jacobs L. L., Gonçalves O. A., & Morais M. - L. (2015).  A new elasmosaurid from the early Maastrichtian of Angola and the implications of girdle morphology on swimming style in plesiosaurs. Netherlands Journal of Geosciences. FirstView, 1–12., 1 Abstractaraujo_et_al_2015_a_new_elasmosaurid_from_the_early_maastrichtian_of_angola.pdfWebsite

ABSTRACT We report here a new elasmosaurid from the early Maastrichtian at Bentiaba, southern Angola. Phylogenetic analysis places the new taxon as the sister taxon to Styxosaurus snowii, and that clade as the sister of a clade composed of (Hydrotherosaurus alexandrae (Libonectes morgani + Elasmosaurus platyurus)). The new taxon has a reduced dorsal blade of the scapula, a feature unique amongst elasmosaurids, but convergent with cryptoclidid plesiosaurs, and indicates a longitudinal protraction-retraction limb cycle rowing style with simple pitch rotation at the glenohumeral articulation. Morphometric phylogenetic analysis of the coracoids of 40 eosauropterygian taxa suggests that there was a broad range of swimming styles within the clade.

Araujo, R., Mateus O., Walen A., & Christiansen N. (2009).  Preparation techniques applied to a stegosaurian Dinosaur from Portugal: excavation, laboratory mechanical and chemical preparation, moulding and casting, 3D scanning. Journal of Paleontological Techniques 5: 22.. 22. Abstract
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Araújo, R., Jacobs L., Polcyn M., Mateus O., & Schulp A. (2010).  Plesiosaurs from the Maastrichtian of Bentiaba, Namibe Province, Angola. Society of Vertebrate Paleontology 70th Annual Meeting.. Abstractaraujo_polcyn_mateus__schulp_2010_plesiosaurs_from_maastrichtian_of_angola_svp10abstracts.pdf

Recent excavations at the Maastrichtian locality of Bentiaba, Namibe Province, in the southern part of Angola, have yielded high quality and partially articulated plesiosaur specimens that indicate at least three taxa were present. A new elasmosaurid is the most abundant and well-preserved plesiosaur taxon at Bentiaba. It is known from a complete articulated paddle and other elements of the skeleton, many of which remain to be collected. The most diagnostic elements so far recovered are the complete pelvic and pectoral girdles, which indicate that the Bentiaba elasmosaurid is probably a new genus because it bears an asymmetrical ventral process of the coracoids, a complete cordiform posterior coracoid vacuity, and pronounced excavation of the anterior border of the coracoids, among other characters. A polycotylid is [...]

Araújo, R., Polcyn M. J., Schulp A. S., Mateus O., Jacobs L. L., Gon?alves A. O., & Morais M. - L. (2014).  A new elasmosaurid from the early Maastrichtian of Angola and the implications of girdle morphology on swimming style in plesiosaurs. Geologie en Mijnbouw/Netherlands Journal of Geosciences. 94, 109-120., Number 1 Abstract
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Araujo, R., Castanhinha R., & Mateus O. (2008).  Major trends in the evolution of teeth and mandibles in ornithopod dinosaurs. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 18–18., Neuquén, Argentina Abstract
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Araújo, R., Polcyn M., Jacobs L. L., Mateus O., & Schulp A. S. (2011).  Plesiosaur structural extreme from the Maastrichtian of Angola. 71st Annual Meeting of the Society of Vertebrate Paleontology. 63., Jan: Abstracts of the 71st Annual Meeting of the Society of Vertebrate Paleontology Abstractaraujo_et_al_mateus_2011_plesiosaur_extreme_angola_svp11abstracts.pdf

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Araújo, R., Castanhinha R., Martins R. M. S., Mateus O., Hendrickx C., Beckmann F., Schell N., & Alves L. C. (2013).  Filling the gaps of dinosaur eggshell phylogeny: Late Jurassic Theropod clutch with embryos from Portugal. Scientific Reports. 3, , may: Nature Publishing Group AbstractWebsite
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Araújo, R., Castanhinha R., Mateus O., & Martins R. (2012).  Late Jurassic theropod embryos from Porto das Barcas, Lourinhã formation, Portugal. Journal of Vertebrate Paleontology, Program and Abstracts, 2012, ISSN 1937-2809 . 57. Abstractaraujo_et_al_mateus_2012_dinosaur_eggs_portugal_svp_2012_abstract_book_meeting_abstracts.pdf.pdf

A clutch of several crushed eggs and embryos from the Late Jurassic (near the Kimmeridgian-Tithonian
boundary), Lourinhã Formation, Portugal contains a complete maxilla, erupted and scattered teeth,
and presacral vertebrae. The maxilla bears four teeth separated by individualized interdental plates,
the dorsal process of the maxilla is confluent with the maxillary body, the ventral rim of the antorbital
fossa is parallel to the tooth row, and the anterior border of the maxilla forms a right angle with the
ventral margin. The teeth are conical and recurved distally with carinae on mesial and distal sides. The
vertebrae are amphiplatyan, with a ventral pair of neurovascular foramina and heavily pitted articular
facets. These fossils allow unambiguous association of basal theropod osteology (Megalosauroidea) with
a new eggshell morphotype. Synchrotron micro-computed tomographic scanning (SRμCT), scanning
electron microscopy, and thin-sections under polarized and normal light revealed that the outer
ornamentation of the eggshell is composed of anastomosing ridges and islets, the pores communicate
near the outer region of the eggshells, and in radial section they are irregular canals that ramify towards
the surface. Micro-proto induced x-ray emission (micro-PIXE) analysis of the eggshell (excluding pores)
revealed the presence of Mg, Fe, Mn (0.33%, 0.27% and 0.18%, respectively) and several trace elements,
with a corresponding loss of Ca (39.4% detected but 40.0% expected for calcite), which suggests minimal
eggshell diagenesis. The eggshells do not luminesce, which could imply that no diagenetic alteration
took effect. However, the quenching effect of Fe2+ has to be taken into consideration. Conversely,
luminescence is observed in the pores since they are filled with sediment, composed of phyllosilicates,
as revealed by SRμCT, micro-PIXE and x-ray diffraction analyses.

Araújo, R., Castanhinha R., Martins R. M. S., Mateus O., Hendrickx C., Beckmann F., Schell N., & Alves L. C. (2013).  Filling the gaps of dinosaur eggshell phylogeny: Late Jurassic Theropod clutch with embryos from Portugal. Scientific Reports. 3(1924), , 2013/05/30/onlin: Macmillan Publishers Limited. All rights reserved Abstractaraujo_et_al_2013_filling_the_gaps_of_dinosaur_eggshell_phylogeny_late_jurassic_theropod_clutch_with_embryos_from_portugal.pdf

The non-avian saurischians that have associated eggshells and embryos are represented only by the sauropodomorph Massospondylus and Coelurosauria (derived theropods), thus missing the basal theropod representatives. We report a dinosaur clutch containing several crushed eggs and embryonic material ascribed to the megalosaurid theropod Torvosaurus. It represents the first associated eggshells and embryos of megalosauroids, thus filling an important phylogenetic gap between two distantly related groups of saurischians. These fossils represent the only unequivocal basal theropod embryos found to date. The assemblage was found in early Tithonian fluvial overbank deposits of the Lourinhã Formation in West Portugal. The morphological, microstructural and chemical characterization results of the eggshell fragments indicate very mild diagenesis. Furthermore, these fossils allow unambiguous association of basal theropod osteology with a specific and unique new eggshell morphology.

Araujo, R., Castanhinha R., & Mateus O. (2008).  Major trends in the evolution of teeth and mandibles in ornithopod dinosaurs. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 18., Neuquén, Argentina Abstractaraujo_et_al._2008._major_trends_in_the_evolution_of_teeth_and_mandibles_in_ornithopod_dinosaurs.pdf

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Araújo, R., Polcyn M. J., Lindgren J., Jacobs L. L., Schulp A. S., Mateus O., Gon?alves A. O., & Morais M. - L. (2014).  New aristonectine elasmosaurid plesiosaur specimens from the Early Maastrichtian of Angola and comments on paedomorphism in plesiosaurs. Geologie en Mijnbouw/Netherlands Journal of Geosciences. 94, 93-108., Number 1 Abstract
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Azanza, M. M., Coimbra R., Puértolas-Pascual E., Russo J., Bauluz B., & Mateus O. (2019).  Crystallography of Lourinhanosaurus eggshells (Dinosauria, Theropoda, Allosauroidea). Journal of Vertebrate Paleontology, Program and Abstracts. 156-157.moreno_azanza_et_al_2019_svp_abstract.pdf
Azanza, M. M., Coimbra R., Puértolas-Pascual E., Russo J., Bauluz B., & Mateus O. (2019).  Crystallography of Lourinhanosaurus eggshells (Dinosauria, Theropoda, Allosauroidea). Journal of Vertebrate Paleontology, Program and Abstracts. 156-157. Abstract
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Beccari, V., Mateus O., Wings O., Milàn J., & Clemmensen L. B. (2021).  Issi saaneq gen. et sp. nov.—A New Sauropodomorph Dinosaur from the Late Triassic (Norian) of Jameson Land, Central East Greenland. Diversity. 13, , Number 11 Abstractdiversity-13-00561-v2.pdfWebsite

The Late Triassic (Norian) outcrops of the Malmros Klint Formation, Jameson Land (Greenland) have yielded numerous specimens of non-sauropod sauropodomorphs. Relevant fossils were briefly reported in 1994 and were assigned to Plateosaurus trossingensis. However, continuous new findings of early non-sauropod sauropodomorphs around the globe facilitate comparisons and allow us to now revise this material. Here, the non-sauropod sauropodomorph Issi saaneq gen. et sp. nov. is described based on two almost complete and articulated skulls. The two skulls represent a middle-stage juvenile and a late-stage juvenile or subadult. Issi saaneq differs from all other sauropodomorphs by several unique traits: (1) a small foramen at the medial surface of the premaxilla; (2) an anteroposteriorly elongated dorsoposterior process of the squamosal; (3) a relatively high quadrate relative to rostrum height; (4) a well-developed posterodorsal process of the articular. These features cannot be explained by taphonomy, ontogeny, or intraspecific variation. Issi saaneq shows affinities to Brazilian plateosaurids and the European Plateosaurus, being recovered as the sister clade of the latter in our phylogenetic analysis. It is the northernmost record of a Late Triassic sauropodomorph, and a new dinosaur species erected for Greenland. Issi saaneq broadens our knowledge about the evolution of plateosaurid sauropodomorphs.

Beccari, V., Pinheiro F. L., Nunes I., Anelli L. E., Mateus O., & Costa F. R. (2021).  Osteology of an exceptionally well-preserved tapejarid skeleton from Brazil: Revealing the anatomy of a curious pterodactyloid clade. PLOS ONE. 16(8), e0254789 - ., 2021/08/25: Public Library of Science Abstractbeccari_et_al_2021.pdfWebsite

A remarkably well-preserved, almost complete and articulated new specimen (GP/2E 9266) of Tupandactylus navigans is here described for the Early Cretaceous Crato Formation of Brazil. The new specimen comprises an almost complete skeleton, preserving both the skull and post-cranium, associated with remarkable preservation of soft tissues, which makes it the most complete tapejarid known thus far. CT-Scanning was performed to allow the assessment of bones still covered by sediment. The specimen can be assigned to Tupa. navigans due to its vertical supra-premaxillary bony process and short and rounded parietal crest. It also bears the largest dentary crest among tapejarine pterosaurs and a notarium, which is absent in other representatives of the clade. The new specimen is here regarded as an adult individual. This is the first time that postcranial remains of Tupa. navigans are described, being also an unprecedented record of an articulated tapejarid skeleton from the Araripe Basin.

Belvedere, M., Castanera D., Meyer C. A., Marty D., Mateus O., Silva B. C., Santos V. F., & Cobos A. (2019).  Late Jurassic globetrotters compared: A closer look at large and giant theropod tracks of North Africa and Europe. Journal of African Earth Sciences. 158, 103547. Abstractbelvedere_et_al_2019_jurassic_globetrotters_compared.pdfWebsite

Late Jurassic theropod tracks are very common both in North Africa and Europe. Two recently described ichnotaxa Megalosauripus transjuranicus and Jurabrontes curtedulensis from the Kimmeridgian of Switzerland show the coexistence of two apex predators in the same palaeoenvironment. Similar tracks can be found in tracksites from the Iberian Peninsula and from Morocco. Here, we further explore the similarities among the Swiss ichnotaxa and the other tracks from Germany (Kimmeridgian), Spain (Tithonian-Berriasian), Portugal (Oxfordian-Tithonian) and Morocco (Kimmeridgian) through novel three-dimensional data comparisons. Specimens were grouped in two morphotypes: 1) large and gracile (30 < Foot Length<50 cm) and 2) giant and robust (FL > 50 cm). The analyses show a great morphological overlap among these two morphotypes and the Swiss ichnotaxa (Megalosauripus transjuranicus and Jurabrontes curtedulensis, respectively), even despite the differences in sedimentary environment and age. This suggests a widespread occurrence of similar ichnotaxa along the western margin of Tethys during the Late Jurassic. The new data support the hypothesis of a Gondwana-Laurasia faunal exchange during the Middle or early Late Jurassic, and the presence of migratory routes around the Tethys.

Bianucci, G., Miján I., Lambert O., Post K., & Mateus O. (2013).  Bizarre fossil beaked whales (Odontoceti, Ziphiidae) fished from the Atlantic Ocean floor off the Iberian Peninsula. Geodiversitas. 35(1), 105-153. Abstractbianucci_et_al_2013_fossil_beaked_whales_iberian_peninsula.pdf

Forty partial fossil skulls belonging to beaked whales (Cetacea, Odontoceti, Ziphiidae) were collected by trawling and long-line fishing on Neogene (probably Late Early to Middle Miocene) layers of the Atlantic floor off the coasts of Portugal and Spain (Asturias and Galicia). e systematic study of the most diagnostic Iberian specimens, those preserving the rostrum and the dorsal part of the cranium, led to the recognition of two new genera (Globicetus n. gen. and Imocetus n. gen.) and four new species (Choneziphius leidyi n. sp., G. hiberus n. gen., n. sp., I. piscatus n. gen., n. sp., and Tusciziphius atlanticus n. sp.).
Based on the matrix of a previous work, the phylogenetic analysis places all the new taxa in the subfamily Ziphiinae Gray, 1850. More fragmentary specimens are tentatively referred to the genera Caviziphius Bianucci & Post, 2005 and Ziphirostrum du Bus, 1868. Among these new ziphiids, extremely bizarre skull morphologies are observed. In G. hiberus n. gen., n. sp. the proximal portion of the rostrum bears a voluminous premaxillary spheroid. In T. atlanticus n. sp. a medial premaxillary bulge is present on the rostrum; together with asymmetric
rostral maxillary eminences at the rostrum base, this bulge displays various degrees of elevation in different specimens, which may be interpreted as sexual dimorphism. Specimens of I. piscatus n. gen., n. sp. bear two sets of even crests: spur-like rostral maxillary crests and longitudinal maxillary crests laterally bordering a wide and long facial basin. A preliminary macroscopic observation of these elements indicates very dense bones, with a compactness comparable with that of cetacean ear bones. Questioning their function, the high medial rostral elements (the premaxillary spheroid of G. hiberus n. gen., n. sp. and the medial bulge of T. atlanticus n. sp.) remind the huge rostral maxillary crests of adult males of the extant Hyperoodon ampullatus (Forster, 1770). In the latter, the crests are very likely related to head-butting. However, they are made of much more spongy bone than in the fossil taxa studied here, and therefore possibly better mechanically suited for facing impacts. Other interpretations of these unusual bone specializations, related to deep-diving (ballast) and echolocation (sound reflection), fail to explain the diversity of shapes and the hypothetical sexual dimorphism observed in at least part of the taxa. e spur-like rostral maxillary crests and long maxillary crests limiting the large facial basin in I. piscatus n. gen., n. sp. and the excrescences on the maxilla at the rostrum base in Choneziphius spp. are instead interpreted as areas of origin for rostral and facial muscles, acting on the nasal passages, blowhole, and melon. From a palaeobiogeographic point of view, the newly described taxa further emphasize the differences in the North Atlantic (including Iberian Peninsula) and South African Neogene ziphiid faunal lists. Even if the stratigraphic context is poorly understood, leaving open the question of the geological age for most of the dredged specimens, these differences in the composition of cold to temperate northern and southern hemisphere fossil ziphiid faunas may be explained by a warm-water equatorial barrier.

Bonaparte, J. F., & Mateus O. (1999).  A new diplodocid, Dinheirosaurus lourinhanensis gen. et sp. nov., from the Late Jurassic beds of Portugal. Revista del Museo Argentino de Ciencias Naturales. 5, 13–29., Number 2 Abstract
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Bonaparte, J. F., & Mateus O. (1999).  A new diplodocid, Dinheirosaurus lourinhanensis gen. et sp. nov., from the Late Jurassic beds of Portugal. Rev. Mus. Argent. Cienc. Nat. 5, 13-29., Number 2 Abstractbonapartemateus1999_sauropod_dinheirosaurus_portugal.pdfWebsite

Presacral vertebrae of a new Diplodocidae from the Late Jurassic Amoreira-Porto Novo Formation of Lourinhã, Portugal are described and figured. Dinheirosaurus lourinhanensis gen. et sp. is diagnosed by well developed accessory articulations derived from the hyposphene, exposed in lateral view; and by the smaller dorsoventral length of the lower section of the neural arch. It is considered that the organization of the dorsal neural arch of Dinheirosaurus is more derived than in Diplodocus, except in the dorsoventral development of the lower portion of it, which is higher (more derived) in the latter. Possibly the isolated geography of Portugal in the Late Jurassic gave rise to the distinct characters of this new genus.

Bonaparte, J. F., & Mateus O. (1999).  A new diplodocid, Dinheirosaurus lourinhanensis gen. et sp. nov., from the Late Jurassic beds of Portugal. Revista del Museo Argentino de Ciencias Naturales. 5, 13–29., Number 2 Abstract
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Brandao, J. M., Callapez P. M., Mateus O., & Castro P. (2010).  Colecções e museus de Geologia: missão e gestão. , Jan: Ed. Universidade de Coimbra e Centro de Estudos e Filosofia da História da Ciência. Abstract

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Brusatte, S. L., Butler R. J., Mateus O., & Steyer J. S. (2015).  A new species of Metoposaurus from the Late Triassic of Portugal and comments on the systematics and biogeography of metoposaurid temnospondyls. Journal of Vertebrate Paleontology. 35, , Number 3 Abstract
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Brusatte, S. L., Butler R. J., Mateus O., Steyer J. S., & Whiteside J. H. (2013).  Terrestrial vertebrates from the Late Triassic of Portugal: new records of temnospondyls and archosauriforms from a Pangaean rift sequence. 61st Symposium on Vertebrate Palaeontology and Comparative Anatomy. 15-16., Edinburgh Abstractbrusatte_et_al_2013_triassic_algarve_portugal_temnospondyls_svpca.org_years_2013_edinburgh_abstracts.pdf

The Late Triassic (ca. 237-201.5 million years ago) was a transitional interval in the evolution of terrestrial ecosystems, during which ‘modern’ clades such as archosaurs and mammals were radiating while ‘archaic’ groups such as temnospondyl amphibians and basal synapsids remained abundant. Little is known about the Triassic terrestrial (nonmarine) vertebrates of the Iberian Peninsula. The Algarve Basin of southern Portugal is an extensional rift basin formed during the breakup of Pangaea, which is filled with terrestrial, lacustrine, and marginal marine siliciclastics of the Grés de Silves Formation, interbedded with CAMP basalts that mark the end-Triassic extinction (radioisotopically dated to ~198-201.5 Ma). Since 2009, our field project in the Algarve has discovered numerous vertebrate specimens within the Grés de Silves, including a monodominant bonebed containing hundreds of specimens of metoposaurids, a peculiar group of temnospondyls that filled crocodile-like predatory niches in lacustrine and fluvial environments. These specimens appear to belong to a new species of Metoposaurus, similar to M. diagnosticus and M. krasiejowensis from central Europe but possessing several putative autapomorphies of the braincase and lower jaw. We also discovered a mandible of a phytosaur, the first specimen of these long-snouted, semi-aquatic archosauriforms from the Iberian Peninsula. These discoveries of characteristic Carnian Norian taxa indicate that the fossil-bearing portion of the Grés de Silves is Late Triassic in age, and provide further evidence that metoposaurids and phytosaurs commonly occurred together in low palaeolatitudes during this time.

Brusatte, S. L., Butler R. J., Mateus O., & Steyer S. J. (2015).  A new species of Metoposaurus from the Late Triassic of Portugal and comments on the systematics and biogeography of metoposaurid temnospondyls. Journal of Vertebrate PaleontologyJournal of Vertebrate Paleontology. e912988., 2015: Taylor & Francis Abstractbrusatte_et_al2015metoposaurusportugal.pdfWebsite

ABSTRACTMetoposaurids are a group of temnospondyl amphibians that filled crocodile-like predatory niches in fluvial and lacustrine environments during the Late Triassic. Metoposaurids are common in the Upper Triassic sediments of North Africa, Europe, India, and North America, but many questions about their systematics and phylogeny remain unresolved. We here erect Metoposaurus algarvensis, sp. nov., the first Metoposaurus species from the Iberian Peninsula, based on several new specimens from a Late Triassic bonebed in Algarve, southern Portugal. We describe the cranial and pectoral anatomy of M. algarvensis and compare it with other metoposaurids (particularly other specimens of Metoposaurus from Germany and Poland). We provide a revised diagnosis and species-level taxonomy for the genus Metoposaurus, which is currently represented with certainty by three European species (M. diagnosticus, M. krasiejowensis, M. algarvensis). We also identify cranial characters that differentiate these three species, and may have phylogenetic significance. These include features of the braincase and mandible, which indicate that metoposaurid skulls are more variable than previously thought. The new Portuguese bonebed provides further evidence that metoposaurids congregated in fluvial and lacustrine settings across their geographic range and often succumbed to mass death events. We provide an updated paleogeographic map depicting all known metoposaurid occurrences, which shows that these temnospondyls were globally distributed in low latitudes during the Late Triassic and had a similar, but not identical, paleogeographic range as phytosaurs.http://zoobank.org/urn:lsid:zoobank.org:pub:083C80C6-0AB6-49E1-A636-6A8BDBC06A47ABSTRACTMetoposaurids are a group of temnospondyl amphibians that filled crocodile-like predatory niches in fluvial and lacustrine environments during the Late Triassic. Metoposaurids are common in the Upper Triassic sediments of North Africa, Europe, India, and North America, but many questions about their systematics and phylogeny remain unresolved. We here erect Metoposaurus algarvensis, sp. nov., the first Metoposaurus species from the Iberian Peninsula, based on several new specimens from a Late Triassic bonebed in Algarve, southern Portugal. We describe the cranial and pectoral anatomy of M. algarvensis and compare it with other metoposaurids (particularly other specimens of Metoposaurus from Germany and Poland). We provide a revised diagnosis and species-level taxonomy for the genus Metoposaurus, which is currently represented with certainty by three European species (M. diagnosticus, M. krasiejowensis, M. algarvensis). We also identify cranial characters that differentiate these three species, and may have phylogenetic significance. These include features of the braincase and mandible, which indicate that metoposaurid skulls are more variable than previously thought. The new Portuguese bonebed provides further evidence that metoposaurids congregated in fluvial and lacustrine settings across their geographic range and often succumbed to mass death events. We provide an updated paleogeographic map depicting all known metoposaurid occurrences, which shows that these temnospondyls were globally distributed in low latitudes during the Late Triassic and had a similar, but not identical, paleogeographic range as phytosaurs.http://zoobank.org/urn:lsid:zoobank.org:pub:083C80C6-0AB6-49E1-A636-6A8BDBC06A47

Burigo, A., & Mateus O. (2024).  Notes on the type specimens of Allosaurus atrox and A. Ferox (Theropoda, Allosauridae). Paleolusitana. 2, 18-20. Abstractaburigo__omateus_v2.pdf

Allosaurus Marsh, 1877 is one of the most iconic theropods of the Late Jurassic of USA and Portugal. However,
since the first description the genus was used as taxonomical wastebasket to which several species were
described. We revisit three type specimens, Allosaurus fragilis YPM 1930, A. atrox YPM 1890 and A. ferox USNM
2315. We interpret the vertebral material of A. atrox and support it as a junior synonym of A. fragilis, and A. ferox
as Allosaurus indet. rather than A. fragilis.

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Campos, H., & Mateus O. (2018).  The first record of placodonts in Portugal and its chronological and paleoecological implications. XVI Annual Meeting of the European Association of Vertebrate Palaeontologists. 38.: Abstract book of the XVI Annual Meeting of the European Association of Vertebrate Palaeontologists, Caparica, Portugal June 26th-July 1st, 2018. Abstractcampos__mateus_2018_eavp_abstract.pdf

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Campos, H., Mateus O., & Moreno-Azanza M. (2017).  Preliminary results on the stratigraphy and taphonomy of multiple bonebeds in the Triassic of Algarve. Abstract book of the XV Encuentro de Jóvenes Investigadores en Paleontología/XV Encontro de Jovens Investigadores em Paleontologia, Lisboa, 428 pp.. 83-87. Abstractcampos_et_al._2017_preliminary_results_pn_the_stratigraphy_and_taphonomy_of_multiple_beds_in_the_triassic_of_algarve..pdf

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Campos, H., & Mateus O. (2018).  The first record of placodonts in Portugal and its chronological and paleoecological implications. XVI Annual Meeting of the European Association of Vertebrate Palaeontologists. 38.: Abstract book of the XVI Annual Meeting of the European Association of Vertebrate Palaeontologists, Caparica, Portugal June 26th-July 1st, 2018. Abstract

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Carrano, M., Mateus O., & Mitchell J. (2013).  First definitive association between embryonic Allosaurus bones and prismatoolithus eggs in the Morrison Formation (Upper Jurassic, Wyoming, USA). Annual Meeting of Vertebrate Paleontology. 101.: Journal of Vertebrate Paleontology, Program and Abstracts, 2013 Abstractcarrano_mateus_mitchell_2013_allosaurus_embryos_morrison_svp_abstract.pdf

Despite more than a century of collecting, resulting in one of the best-studied vertebrate fossil records anywhere in the world, the Upper Jurassic Morrison Formation has produced surprisingly few examples of dinosaur eggs associated with embryonic remains. Even more puzzling, none of these seem to pertain to the theropod Allosaurus, one of the most common and best-understood dinosaur taxa in the formation. Here we
report on a dinosaur nest site that has produced both abundant prismatoolithid eggshell and embryonic (or perinatal) bones of Allosaurus from Fox Mesa, Wyoming. This represents the first such discovery for any theropod in the Jurassic of North America. The nest is heavily weathered but contains a few ellipsoid eggshell clusters that suggest an egg size of about 8 x 6.5 cm. Study of the eggshell morphology and microstructure confirms that a single egg type is present throughout, which is indistinguishable from Prismatoolithus coloradensis. All of the identifiable embryonic materials pertain to theropods, and two premaxillae specimens show the five alveoli diagnostic for Allosaurus among Morrison theropods. This confirms the theropod origin of Prismatoolithus eggs and implicates Allosaurus as the specific Morrison parent taxon. As a result, it is now possible to assign several previous discoveries of dinosaur eggs and potential nests to Allosaurus, including the isolated egg from the Cleveland-Lloyd Quarry. This discovery
also calls into question prior assignments of Prismatoolithus eggs to ornithopods, and suggests that more detailed study of such sites is warranted. Prismatoolithus eggshells are also associated with the Upper Jurassic theropod Lourinhanosaurus from Portugal, along with larger embryos that exhibit four premaxillary alveoli.