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Mateus, O., Neto de Carvalho C., & Klompmaker A. A. (2013).  Decapod crustacean body and ichnofossils from the Mesozoic of Portugal. 5th Symposium on Mesozoic and Cenozoic Decapod Crustaceans. , 25–27 June 2013, Warszawa: Polish Geological Institute − National Research Institute & AGH University of Science and Technologymateus_et_al_2013_crustacea_mesozoic_portugal_5th_decapod_crustaceans_meeting_2013.pdf
Mateus, O. (2014).  Dinosaur taphonomy in the Lourinhã Formation (Late Jurassic, Portugal). International Meeting on Taphonomy and Fossilization. 60–61. Abstract
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Mateus, O. (1998).  Dinossauros de Portugal e um novo terópode do Jurássico Superior da Lourinhã. , Évora: Universidade de Évora Abstract
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Mateus, O. (2014).  Gigantic Jurassic predators. 52 Things You Should Know About Palaeontology. 56-57.: Agile Libremateus_2014_gigantic_jurassic_predators.pdf
Mateus, O. (2009).  The Cretaceous Skeleton Coast of Angola. 29, , 1 Abstract
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Mateus, O., & Jacinto J. J. (1998).  Activity Rithms and habitat of Hemidactylus turcicus (Reptilia, Gekkonidae) in Évora, Portugal. Boletin de ICIJA. 2, 37-43. Abstract
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Mateus, O. (2009).  The sauropod Turiasaurus riodevensis in the the Late Jurassic of Portugal. Journal of Vertebrate Paleontology. 29, 144–144., Number 3 Abstract
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Mateus, O., & Antunes M. T. (2008).  Landmarks in the history of dinosaur paleontology in Portugal, focusing on skeletal remains. Abstract volume, Dinosaurs - A Historical Perspective, 6-7 may 2008. , London Abstract
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Mateus, O., Butler R. J., Brusatte S. L., Whiteside J. H., & Steyer J. S. (2014).  The first phytosaur (Diapsida, Archosauriformes) from the Late Triassic of the Iberian Peninsula. Journal of Vertebrate Paleontology. 34, 970-975., Number 4 Abstract
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Mateus, O., Morais M. L., Schulp A. S., Jacobs L. L., & Polcyn M. J. (2006).  The Cretaceous of Angola. Journal of Vertebrate Paleontology. 26, 96–97., Number (Suppl. T Abstract
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Marzola, M., Mateus O., Russo J., & Milàn J. (2014).  Comparison of modern and fossil Crocodylomorpha eggs and contribution to the oophylogeny of Amniota. XII Annual Meeting of the European Association of Vertebrate Palaeontologists. , p. 192, Regione Piemonte: European Association of Vertebrate Palaeontologists. Museo Regionale di Scienze Naturalimarzola_et_al._2014_comparison_of_modern_and_fossil_crocodylomorpha_eggs_and_contribution_to_the_oophylogeny_of_amniota-_eavp_2014.pdf
Marzola, M., Mateus O., Schulp A. S., Jacobs L. L., Polcyn M. J., Pervov V., Goncalves A. O., & Morais M. L. (2015).  Comparative anatomy and systematics of Cretaceous mammal tracks of Angola. 13th Annual Meeting of the European Association of Vertebrate Palaeontologists - EAVP 2015. , July 2015, Opole, Poland: European Association of Vertebrate Palaeontologistsmarzola_et_al_2015_catoca_tracks_eavp.pdf
Marzola, M., Mateus O., Milàn J., & Clemmensen L. B. (2017).  The 2016 Dinosaur Expedition to the Late Triassic of the Jameson Land Basin, East Greenland. Abstract book of the XV Encuentro de Jóvenes Investigadores en Paleontología/XV Encontro de Jovenes Investigadores em Paleontologia, Pombal, 428 pp.. 249–253. Abstract08._ejip_2017_-_marzola_et_al._2017._the_2016_dinosaur_expedition_to_the_late_triassic_of_the_jameson_land_basin_east_greenland.pdf

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Marzola, M., Mateus O., Wings O., Klein N., M{\`ılan J., & L.B.Clemmensen (2016).  The herpetofauna from the Late Triassic of the Jameson Land Basin (East Greenland): review and updates. XIV EAVP Meeting. 182., Haarlem, The Netherlands: XIV EAVP Meeting, Programme and Abstract Book Abstract
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Marzola, M., Mateus O., Milàn J., & Clemmensen L. B. (2018).  A review of palaeozoic and mesozoic tetrapods from greenland. Bulletin of the Geological Society of Denmark. 66, 21-46. Abstract
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Marzola, M., Russo J., & Mateus O. (2015).  Identification and comparison of modern and fossil crocodilian eggs and eggshell structures. Historical Biology. 27(1), 115-133. Abstractmarzola_et_al_2015_identification_and_comparison_of_modern_and_fossil_crocodilian_eggs_and_eggshell_structures.pdfWebsite

Eggshells from the three extant crocodilian species Crocodylus mindorensis (Philippine Crocodile), Paleosuchus palpebrosus (Cuvier's Smooth-fronted Caiman or Musky Caiman) and Alligator mississippiensis (American Alligator or Common Alligator) were prepared for thin section and scanning electron microscope analyses and are described in order to improve the knowledge on crocodilian eggs anatomy and microstructure, and to find new apomorphies that can be used for identification. Both extant and fossil crocodilian eggs present an ornamentation that vary as anastomo-, ramo- or the here newly described rugosocavate type. The angusticaniculate pore system is a shared character for Crocodylomorpha eggshells and some dinosaurian and avian groups. Previously reported signs of incubated crocodilian eggs were found also on our only fertilised and hatched egg. Paleosuchus palpebrosus presents unique organization and morphology of the three eggshell layers, with a relatively thin middle layer characterised by dense and compact tabular microstructure.

Marzola, M., Mateus O., Milàn J., & Clemmensen L. B. (2017).  The 2016 Dinosaur Expedition to the Late Triassic of the Jameson Land Basin, East Greenland. Abstract book of the XV Encuentro de Jóvenes Investigadores en Paleontolog{\'ıa/XV Encontro de Jovenes Investigadores em Paleontologia, Pombal, 428 pp.. 249–253. Abstract
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Marzola, M., Mateus O., Schulp {A. S. }, Jacobs {L. L. }, Polcyn {M. J. }, Pervov V., Goncalves {A. O. }, & Morais {M. L. } (2015).  Comparative anatomy and systematics of Cretaceous mammal tracks of Angola. 35. Abstract
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Marzola, M., Mateus O., Milàn J., & Clemmensen L. B. (2017).  European affinities of the Late Triassic biota from Greenland are related to paleolatitude. 15th Annual Meeting of the European Association of Vertebrate Palaeontologists. 91, 61.: Zitteliana Abstract09._eavp_2017_-_marzola_et_al._2017._european_affinities_of_the_late_triassic_biota_from_greenland_are_related_to_paleolatitude.pdf

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Marzola, M., Mateus O., Wings O., Klein N., M\{\`ı\}lan J., & L.B.Clemmensen (2016).  The herpetofauna from the Late Triassic of the Jameson Land Basin (East Greenland): review and updates. XIV EAVP Meeting. 182., Haarlem, The Netherlands Abstract
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Marzola, M., Mateus O., & Moreno-Azanza(eds)M. (2018).  Abstract book of the XVI Annual Meeting of the European Association of Vertebrate Palaeontology. XVI Annual Meeting of the European Association of Vertebrate Palaeontologists. , Caparica: Faculdade de Ciências e Tecnologia da Universidade Nova de Lisboa Abstract
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Marzola, M., Mateus O., Milàn J., & Clemmensen L. B. (2017).  European affinities of the Late Triassic biota from Greenland are related to paleolatitude. 15th Annual Meeting of the European Association of Vertebrate Palaeontologists. 91, 61.: Zitteliana Abstract
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Marzola, M., Mateus O., Milàn J., & Clemmensen {L. B. } (2017).  The 2016 Dinosaur Expedition to the Late Triassic of the Jameson Land Basin, East Greenland. 249–253. Abstract
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Marzola, M., Mateus O., & Moreno-Azanza(eds)M. (2018).  Abstract book of the XVI Annual Meeting of the European Association of Vertebrate Palaeontology. XVI Annual Meeting of the European Association of Vertebrate Palaeontologists. , Caparica: Faculdade de Ciências e Tecnologia da Universidade Nova de Lisboaeavp_2018_abstract_book.pdf
Marzola, M., Russo J., & Mateus O. (2015).  Identification and comparison of modern and fossil crocodilian eggs and eggshell structures. Historical Biology. 27, 115-133., Number 1 Abstract
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Marzola, M., Mateus O., Milàn J., & Clemmensen L. B. (2018).  A review of Palaeozoic and Mesozoic tetrapods from Greenland. Bulletin of the Geological Society of Denmark. 66, 21–46. Abstractmarzola_et_al_2018_-_review_of_greenlandic_tetrapods.pdf

This article presents a synthesis of Palaeozoic and Mesozoic fossil tetrapods from Greenland, including an updated review of the holotypes and a new photographic record of the main specimens. All fossil tetrapods found are from East Greenland, with at least 30 different known taxa: five stem tetrapods (Acanthostega gunnari, Ichthyostega eigili, I. stensioi, I. watsoni, and Ymeria denticulata) from the Late Devonian of the Aina Dal and Britta Dal Formations; four temnospondyl amphibians (Aquiloniferus kochi, Selenocara groenlandica, Stoschiosaurus nielseni, and Tupilakosaurus heilmani) from the Early Triassic of the Wordie Creek Group; two temnospondyls (Cyclotosaurus naraserluki and Gerrothorax cf. pulcherrimus), one testudinatan (cf. Proganochelys), two stagonolepids (Aetosaurus ferratus and Paratypothorax andressorum), the eudimorphodontid Arcticodactylus, undetermined archosaurs (phytosaurs and both sauropodomorph and theropod dinosaurs), the cynodont Mitredon cromptoni, and three mammals (Haramiyavia clemmenseni, Kuehneotherium, and cf. ?Brachyzostrodon), from the Late Triassic of the Fleming
Fjord Formation; one plesiosaur from the Early Jurassic of the Kap Stewart Formation; one plesiosaur and one ichthyosaur from the Late Jurassic of the Kap Leslie Formation, plus a previously unreported Late Jurassic plesiosaur from Kronprins Christian Land. Moreover, fossil tetrapod trackways are known from the Late Carboniferous (morphotype Limnopus) of the Mesters Vig Formation and at least four different morphologies (such as the crocodylomorph Brachychirotherium, the auropodomorph Eosauropus and Evazoum, and the theropodian Grallator) associated to archosaurian trackmakers are known from the Late Triassic of the Fleming Fjord Formation. The presence of rich fossiliferous tetrapod sites in East Greenland is linked to the presence of well-exposed continental and shallow marine deposits with most finds in terrestrial deposits from the Late Devonian and the Late Triassic.

Marzola, M., Mateus O., Schulp A. S., Jacobs L. L., Polcyn M. J., Pervov V., Goncalves A. O., & Morais M. L. (2015).  Comparative anatomy and systematics of Cretaceous mammal tracks of Angola. 13th Annual Meeting of the European Association of Vertebrate Palaeontologists - EAVP 2015. , July 2015, Opole, Poland: European Association of Vertebrate Palaeontologists Abstract
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Marzola, M., Mateus O., Shubin N. H., & Clemmensen L. B. (2017).  Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland). Journal of Vertebrate Paleontology. e1303501., 2017: Taylor & Francis Abstractmarzola_et_al_2017_cyclotosaurus_greenland.pdfWebsite

ABSTRACTCyclotosaurus naraserluki, sp. nov., is a new Late Triassic capitosaurid amphibian from lacustrine deposits in the Fleming Fjord Formation of the Jameson Land Basin in Greenland. It is based on a fairly complete and well-preserved skull associated with two vertebral intercentra. Previously reported as Cyclotosaurus cf. posthumus, C. naraserluki is unique among cyclotosaurs for having the postorbitals embaying the supratemporals posteromedially. The anterior palatal vacuity presents an autapomorphic complete subdivision by a wide medial premaxillary-vomerine bony connection. The parasphenoid projects between the pterygoids and the exoccipitals, preventing a suture between the two, a primitive condition shared with Rhinesuchidae, Eryosuchus, and Kupferzellia. Within Cyclotosaurus, the Greenlandic skull has a distinctive combination of circular choanae (shared with C. ebrachensis, C. posthumus, and C. robustus) and a convex posteromedial margin of the tabulars (also present in C. ebrachensis and C. intermedius). A phylogenetic analysis indicates that C. naraserluki is the sister taxon of the middle Norian C. mordax from southern Germany, with which it shares a pair of premaxillary foramina. Cyclotosaurus is one of the most successful and diverse genera of Late Triassic temnospondyls, with at least eight species reported from middle Carnian to late Norian. Cyclotosaurus naraserluki is the largest amphibian ever reported from Greenland and one of the Late Triassic vertebrates with the highest northern paleolatitude currently known.http://zoobank.org/urn:lsid:zoobank.org:pub:43AAA541-031C-4EE1-B819-4846EBBD1BBBSUPPLEMENTAL DATA?Supplemental materials are available for this article for free at www.tandfonline.com/UJVPCitation for this article: Marzola, M., O. Mateus, N. H. Shubin, and L. B. Clemmensen. 2017. Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland). Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2017.1303501.ABSTRACTCyclotosaurus naraserluki, sp. nov., is a new Late Triassic capitosaurid amphibian from lacustrine deposits in the Fleming Fjord Formation of the Jameson Land Basin in Greenland. It is based on a fairly complete and well-preserved skull associated with two vertebral intercentra. Previously reported as Cyclotosaurus cf. posthumus, C. naraserluki is unique among cyclotosaurs for having the postorbitals embaying the supratemporals posteromedially. The anterior palatal vacuity presents an autapomorphic complete subdivision by a wide medial premaxillary-vomerine bony connection. The parasphenoid projects between the pterygoids and the exoccipitals, preventing a suture between the two, a primitive condition shared with Rhinesuchidae, Eryosuchus, and Kupferzellia. Within Cyclotosaurus, the Greenlandic skull has a distinctive combination of circular choanae (shared with C. ebrachensis, C. posthumus, and C. robustus) and a convex posteromedial margin of the tabulars (also present in C. ebrachensis and C. intermedius). A phylogenetic analysis indicates that C. naraserluki is the sister taxon of the middle Norian C. mordax from southern Germany, with which it shares a pair of premaxillary foramina. Cyclotosaurus is one of the most successful and diverse genera of Late Triassic temnospondyls, with at least eight species reported from middle Carnian to late Norian. Cyclotosaurus naraserluki is the largest amphibian ever reported from Greenland and one of the Late Triassic vertebrates with the highest northern paleolatitude currently known.http://zoobank.org/urn:lsid:zoobank.org:pub:43AAA541-031C-4EE1-B819-4846EBBD1BBBSUPPLEMENTAL DATA?Supplemental materials are available for this article for free at www.tandfonline.com/UJVPCitation for this article: Marzola, M., O. Mateus, N. H. Shubin, and L. B. Clemmensen. 2017. Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland). Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2017.1303501.

Marzola, M., Mateus O., Milàn J., & Clemmensen {L. B. } (2017).  European affinities of the Late Triassic biota from Greenland are related to paleolatitude. 61. Abstract
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Marzola, M., Mateus O., Wings O., Klein N., Mìlan J., & L.B.Clemmensen (2016).  The herpetofauna from the Late Triassic of the Jameson Land Basin (East Greenland): review and updates. XIV EAVP Meeting. 182., Haarlem, The Netherlands: XIV EAVP Meeting, Programme and Abstract Book
Marzola, M., Mateus O., Wings O., Klein N., Milan J., & [Unknown] L. B. C. (2016).  The herpetofauna from the Late Triassic of the Jameson Land Basin (East Greenland): review and updates. : XIV EAVP Meeting, Programme and Abstract Book Abstract
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Marzola, M., Mateus O., Shubin N. H., & Clemmensen L. B. (2017).  Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland). Journal of Vertebrate Paleontology. e1303501., may: Informa {UK} Limited AbstractWebsite
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Marzola, M., Mateus O., Milàn J., & Clemmensen {L. B. } (2017).  Synrift sedimentary deposition and vertebrate fossil abundance: the tetrapod record from Greenland. 159–160. Abstract
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Marzola, M., Mateus O., Milàn J., & Clemmensen L. B. (2017).  Synrift sedimentary deposition and vertebrate fossil abundance: the tetrapod record from Greenland. Journal of Vertebrate Paleontology, Program and Abstracts. 2017, 159-160. Abstractmarzola_et_al_2017_svp_abstract_greenland.pdf

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Marzola, M., Mateus O., Schulp A., Jacobs L., Polcyn M., & Pervov V. (2014).  Early Cretaceous tracks of a large mammaliamorph, a crocodylomorph, and dinosaurs from an Angolan diamond mine. Journal of Vertebrate Paleontology, Program and Abstracts, 2014. 181.marzola_et_al_2014._cretaceous_tracks_mammaliamorph_a_crocodilomorph_angolan_diamond_mine.pdf
Marzola, M., Mateus O., Schulp A. S., Jacobs L. L., Polcyn M. J., Pervov V., Goncalves A. O., & Morais M. L. (2015).  Comparative anatomy and systematics of Cretaceous mammal tracks of Angola. 13th Annual Meeting of the European Association of Vertebrate Palaeontologists - EAVP 2015. , Opole, Poland Abstract
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Marx, M. P., Mateus O., Polcyn M. J., Schulp A. S., Gonçalves O. A., & Jacobs L. L. (2021).  The cranial anatomy and relationships of Cardiocorax mukulu (Plesiosauria: Elasmosauridae) from Bentiaba, Angola. PLOS ONE. 16(8), e0255773 - ., 2021/08/17: Public Library of Science AbstractWebsite

We report a new specimen of the plesiosaur Cardiocorax mukulu that includes the most complete plesiosaur skull from sub-Saharan Africa. The well-preserved three-dimensional nature of the skull offers rare insight into the cranial anatomy of elasmosaurid plesiosaurians. The new specimen of Cardiocorax mukulu was recovered from Bentiaba, Namibe Province in Angola, approximately three meters above the holotype. The new specimen also includes an atlas-axis complex, seventeen postaxial cervical vertebrae, partial ribs, a femur, and limb elements. It is identified as Cardiocorax mukulu based on an apomorphy shared with the holotype where the cervical neural spine is approximately as long anteroposteriorly as the centrum and exhibits a sinusoidal anterior margin. The new specimen is nearly identical to the holotype and previously referred material in all other aspects. Cardiocorax mukulu is returned in an early-branching or intermediate position in Elasmosauridae in four out of the six of our phylogenetic analyses. Cardiocorax mukulu lacks the elongated cervical vertebrae that is characteristic of the extremely long-necked elasmosaurines, and the broad skull with and a high number of maxillary teeth (28–40) which is characteristic of Aristonectinae. Currently, the most parsimonious explanation concerning elasmosaurid evolutionary relationships, is that Cardiocorax mukulu represents an older lineage of elasmosaurids in the Maastrichtian.

Martins, R. M. S., Araújo R., Mateus O., Castanhinha R., Pranzas P. K., & Beckmann F. (2012).  Tomography applied to the study of dinosaur fossils from the collection of the museum of Lourinhã (Portugal). I Congresso Internacional GeoCiencias na CPL. , Coimbra: Univ. Coimbramartins_et_al_2012_abstract.pdf
Martins, R. M. S., Beckmann F., Castanhinha R., Mateus O., & Pranzas P. K. (2011).  Dinosaur and crocodile fossils from the mesozoic of Portugal: Neutron tomography and synchrotron-radiation based micro-computed tomography. Materials Research Society Symposium Proceedings. 1319, 319-332. Abstract
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Martins, R. M. S., Castanhinha R. A., Mateus O., Araújo R., Beckmann F., & Schell N. (2012).  {Synchrotron radiation-based techniques applied to the study of dinosaur fossils from the collection of the museum of Lourinhã. Ciencia e Tecnologia dos Materiais. 24, 153-156., Number 3-4 Abstract
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Martins, R. M. S., Beckmann F., Castanhinha R., Mateus O., Araújo R., & Pranzas P. K. (2012).  Synchrotron radiation-based micro-computed tomography applied to the characterization of dinosaur fossils from the Lourinhã Formation. 1 st . Meeting of Synchrotron Radiation Users from Portugal. 1-2., Caparica, Portugalmartins_et_al_2012_tomography_enurs_martins-rui_dinosaurfossils.pdf
Martins, R. M. S., Beckmann F., Castanhinha R., Araújo R., O. Mateus, & Pranzas P. K. (2011).  Tomographic techniques for the study of exceptionally preserved dinosaur and crocodile fossils from the mesozoic of Portugal. VII Encontro Nacional de Biologia Evolutiva, Coimbra, 21 Dezembro 2011. 19. Abstractmartins_et_al_mateus_2011_tomographic_techniques_for_the_study_of_exceptionally_preserved_dinosaur_and_crocodile.pdf

Portugal is ranked within the ten countries with more dinosaur taxa and the Lourinhã Formation is known
by the Late Jurassic findings of dinosaurs and other fossils. Often, studies of the external morphological
characteristics of the fossils are not sufficient and, observations of internal structures, using non-
destructive techniques, are required. The fossils here presented belong to the Museum da Lourinhã
(Portugal) and comprise a lower jaw of a basal crocodilian (possibly a Tomistomidae), eggshells and
several vertebrae from the exceptionally well preserved in ovo remains of Late Jurassic theropod dinosaur
Lourinhanosaurus. Neutron Tomography (NT) experiments with this material has been carried out at the
Geesthacht Neutron Facility in Germany. Additionally, eggshell fragments and several vertebrae have
been studied by Synchrotron-Radiation based Micro-Computed Tomography (SRμCT) at the HARWI II and
BW2 beamlines, respectively. These beamlines are operated by the Helmholtz-Zentrum Geesthacht at the
storage ring DORIS III at the Deutsches Elektronen–Synchrotron DESY in Hamburg, Germany. In both cases
(NT and SRμCT) complete 3D recordings have been obtained using a non-destructive procedure. The high-
quality tomographic datasets can be effectively studied through interactive digital visualization. Hence,
these visualization methods provide precious information about the 3D internal micro morphology of
fossils, like the network of the eggshell pores, often invisible in more traditional techniques, and provide a
direct window into the evolutionary history of organisms.

Martin Sander, P., Mateus O., Laven T., & Knötschke N. (2006).  Bone histology indicates insular dwarfism in a new Late Jurassic sauropod dinosaur. Nature. 441, 739-741., Number 7094 Abstract
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Marinheiro, J., & Mateus O. (2011).  Occurrence of the marine turtle Thalassemys in the Kimmeridgian of Oker, Germany. 71st Annual Meeting of the Society of Vertebrate Paleontology. 151., Jan: Abstracts of the 71st Annual Meeting of the Society of Vertebrate Paleontology Abstractmarinheiro__mateus_2011_turtle_thalassemys_oker_germany_svp11abstracts.pdf

A partial chelonian was collected from the Langenberg Formation quarry of Oker (near Goslar, Lower Saxony, Germany). The amniotes from this formation include other chelonians (possible Plesiochelyidae), the sauropod Europasaurus holgeri, theropods (Velociraptorinae), crocodylomorphs (Atoposauridae, Theriosuchus pusillus, Goniopholis simus, Machimosaurus hugii, Steneosaurus brevirostris) and pterosaurs (Dsungapteridae, ?Ornithocheiroidea, ?Ctenochasmatidae).

The reported specimen, DFMMh/FV 296, includes a skull part (articulated quadrate, squamosal, basisphenoid, and pterygoid), a disarticulated 40 cm long partial carapace, plastron, and one cervical vertebra.

The carapace bears fontanelles, trapezoidal suprapygal with straight edges, small last neurals

[....]
a plicated longitudinal pattern in the proximal end of the costals originating on the posterior side of the scute sulci and dissipating posteriorly, wide central opening in the plastron, xiphiplastra with little or no contact between each other, and both the hyo- and hypoplastron have digit-form buttress projections.

The specimen has a central plastra fontanelle, which is regarded as a feature of the clade including [....], Santanachelys, and Thalassemys. DFMMh/FV 296 differs from
[....]

edges and xiphiplastron participation in the central fontanelle is autapomorphic for the Thalassemys genus. A large central fontanelle with hyo and hypoplastron polygonal medial
[....]
contact between xiphiplastra and reduced size of the two last neural plates is shared between T. hugii and the Oker specimen. DFMMh/FV 296 differs from this species due to the presence of plastral projections. The different shape of the plastron (no polygonal-like margins or hyo- and hypoplastron projections) suggests that T. moseri [....]
ent genus. Therefore, we can assign this specimen to Thalassemys sp.

Marinheiro, J., Mateus O., Alaoui A., Amani F., Nami M., & Ribeiro C. (2014).  New Quaternary fossil sites from the Middle Atlas of Morocco. Comunicações Geológicas. 101, Especial I, 485-488. Abstractmarinheiro_et_al_2014_new_quaternary_fossil_sites_from_the_middle_atlas_of_morocco.pdf

The paleontological richness of Morocco has been scientifically known since at least the early 20th century. The region of the Middle Atlas, more specifically the Boulemane area, has been however only sparsely studied since the 1960s when it provided vertebrate fossils from the Middle Jurassic. In September 2013, a Moroccan-Portuguese expedition to the village of Taghrout, Boulemane, made excavations in a Pleistocene fossil site that once was a small high-altitude sedimentary basin, uncharted in previous geological maps. The excavations yielded bone material from large mammals, the most common findings are elephants ascribed to Elephas, but artiodactyls, turtles, and in-situ Acheulean tools were also collected. This represents a new and important paleontological and archeological site. In addition to the discoveries of Taghrout, the expedition also retrieved Quaternary vertebrate material from a nearby cave and found new Jurassic localities, with arcosaur bones and dinosaur footprints, in El Mers.