Publications in the Year: 2017

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Mateus, O.  2017.  Que Dinossauros Existiram em Portugal? : Poster 80x59 cm, as a supplement of newspaper “Correio da Manhã” of 16 September 2017poster_correio_da_manha.jpg

Conference Proceedings

Campos, H, Mateus O, Moreno-Azanza M.  2017.  Preliminary results on the stratigraphy and taphonomy of multiple bonebeds in the Triassic of Algarve. Abstract book of the XV Encuentro de Jóvenes Investigadores en Paleontología/XV Encontro de Jovens Investigadores em Paleontologia, Lisboa, 428 pp.. :83-87. Abstractcampos_et_al._2017_preliminary_results_pn_the_stratigraphy_and_taphonomy_of_multiple_beds_in_the_triassic_of_algarve..pdf

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Pereira, B, Mateus O, Kullberg JC, Rocha R.  2017.  The geotouristic potential of the Oeste Region of Portugal. 14th European Geoparks Conference | Abstracts Book 167. :167., Ponta Delgadapereira_et_al_2017_geotouristic_oeste.pdf
Marzola, M, Mateus O, Milàn J, Clemmensen LB.  2017.  The 2016 Dinosaur Expedition to the Late Triassic of the Jameson Land Basin, East Greenland. Abstract book of the XV Encuentro de Jóvenes Investigadores en Paleontología/XV Encontro de Jovenes Investigadores em Paleontologia, Pombal, 428 pp.. :249–253. Abstract08._ejip_2017_-_marzola_et_al._2017._the_2016_dinosaur_expedition_to_the_late_triassic_of_the_jameson_land_basin_east_greenland.pdf

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Correia, T, Barcelos L, Nunes T, Riff D, Mateus O.  2017.  On a titanosaur scapula from the Marília Formation (Upper Cretaceous, Bauru Group) in Campina Verde Town. XXV Congresso Brasileiro de Paleontologia Boletim de Resumos. :77. Abstractthiago_abstract_brazil_2017.pdf

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Costa, F, Silva T, Fernandes J, Calvo R, Mateus O.  2017.  Retracing the history of a stegosaurian dinosaur discovery in Portugal and the importance of record-keeping in Palaentology. Abstract book of the XV Encuentro de Jóvenes Investigadores en Paleontología/XV Encontro de Jovens Investigadores em Paleontologia, Lisboa, 428 pp.. :119-124. Abstractcosta_et_al_2017_retracing_the_history_-_2017.pdf

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Guillaume, A, Costa F, Mateus O.  2017.  Skin impressions on stegosaur tracks from the Upper Jurassic of Portugal. Abstract book of the XV Encuentro de Jóvenes Investigadores en Paleontología/XV Encontro de Jovens Investigadores em Paleontologia, Lisboa, 428 pp.. :191-195. Abstractguillaume_2017_skin.pdf

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Marzola, M, Mateus O, Milàn J, Clemmensen LB.  2017.  European affinities of the Late Triassic biota from Greenland are related to paleolatitude. 15th Annual Meeting of the European Association of Vertebrate Palaeontologists. 91:61.: Zitteliana Abstract09._eavp_2017_-_marzola_et_al._2017._european_affinities_of_the_late_triassic_biota_from_greenland_are_related_to_paleolatitude.pdf

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Mateus, O, Callapez PM, Puértolas-Pascual E.  2017.  The oldest Crocodylia? a new eusuchian from the Late Cretaceous (Cenomanian) of Portugal Journal of Vertebrate Paleontology, Program and Abstracts. 2017:160. Abstractmateus_et_al_2017_oldest_crocodylia_svp_2017_abstract.pdf

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Saleiro, A, Mateus O.  2017.  Upper Jurassic bonebeds around Ten Sleep, Wyoming, USA: overview and stratigraphy. Abstract book of the XV Encuentro de Jóvenes Investigadores en Paleontología/XV Encontro de Jovens Investigadores em Paleontologia, Lisboa, 428 pp.. :357-361. Abstractsaleiro__mateus_2017._upper_jurassic_bonebeds_around_ten_sleep_wyoming_usa-overview_and_stratigraphy..pdf

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Marzola, M, Mateus O, Milàn J, Clemmensen LB.  2017.  Synrift sedimentary deposition and vertebrate fossil abundance: the tetrapod record from Greenland. Journal of Vertebrate Paleontology, Program and Abstracts. 2017:159-160. Abstractmarzola_et_al_2017_svp_abstract_greenland.pdf

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Journal Article

Myers, TS, Polcyn MJ, Mateus O, Vineyard DP, Gonçalves AO, Jacobs LL.  2017.  A new durophagous stem cheloniid turtle from the lower Paleocene of Cabinda, Angola. Papers in Palaeontology. 2017:1-16. Abstractnew_durophagous_stem_cheloniid_turtle_from_the_lower_paleocene_of_cabinda_angola.pdfWebsite

A new stem cheloniid turtle, Cabindachelys landanensis, gen. et sp. nov., is represented by a nearly complete skull and partial hyoid collected in lower Paleocene shallow marine deposits, equivalent to the offshore Landana Formation, near the town of Landana in Cabinda, Angola. A partial chelonioid carapace previously reported from this locality is referred here to C. landanensis. Cabindachelys landanensis possesses clear synapomorphies of Pan-Cheloniidae, including a rod-like rostrum basisphenoidale, V-shaped basisphenoid crest, and secondary palate, but also retains a reduced foramen palatinum posterius, unlike most other pan-cheloniids. Phylogenetic analysis suggests that C. landanensis forms a weakly-supported clade with Erquelinnesia gosseleti, Euclastes acutirostris, Euclastes wielandi and Terlinguachelys fischbecki, although a close relationship between the protostegid T. fischbecki and these durophagous pan-cheloniids is unlikely. The Paleocene–Eocene strata near Landana have produced a number of turtle fossils, including the holotype specimen of the pleurodire Taphrosphys congolensis. A turtle humerus collected c. 1 m above the holotype skull of C. landanensis differs from humeri of chelonioids and Taphrosphys, indicating that a third turtle taxon is present at Landana. Cheloniid fossil material is rare in the Landana assemblage, in comparison with the abundant remains of Taphrosphys congolensis found throughout the stratigraphic section. This disparity implies that C. landanensis preferred open marine habitats, whereas Taphrosphys congolensis spent more time in nearshore environments. The appearance of new durophagous species such as C. landanensis in the early Paleocene reflects the rapid radiation of pan-cheloniids as they diversified into open niches following the K–Pg extinction.

Marzola, M, Mateus O, Shubin NH, Clemmensen LB.  2017.  Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland), 2017. Journal of Vertebrate Paleontology. :e1303501.: Taylor & Francis Abstractmarzola_et_al_2017_cyclotosaurus_greenland.pdfWebsite

ABSTRACTCyclotosaurus naraserluki, sp. nov., is a new Late Triassic capitosaurid amphibian from lacustrine deposits in the Fleming Fjord Formation of the Jameson Land Basin in Greenland. It is based on a fairly complete and well-preserved skull associated with two vertebral intercentra. Previously reported as Cyclotosaurus cf. posthumus, C. naraserluki is unique among cyclotosaurs for having the postorbitals embaying the supratemporals posteromedially. The anterior palatal vacuity presents an autapomorphic complete subdivision by a wide medial premaxillary-vomerine bony connection. The parasphenoid projects between the pterygoids and the exoccipitals, preventing a suture between the two, a primitive condition shared with Rhinesuchidae, Eryosuchus, and Kupferzellia. Within Cyclotosaurus, the Greenlandic skull has a distinctive combination of circular choanae (shared with C. ebrachensis, C. posthumus, and C. robustus) and a convex posteromedial margin of the tabulars (also present in C. ebrachensis and C. intermedius). A phylogenetic analysis indicates that C. naraserluki is the sister taxon of the middle Norian C. mordax from southern Germany, with which it shares a pair of premaxillary foramina. Cyclotosaurus is one of the most successful and diverse genera of Late Triassic temnospondyls, with at least eight species reported from middle Carnian to late Norian. Cyclotosaurus naraserluki is the largest amphibian ever reported from Greenland and one of the Late Triassic vertebrates with the highest northern paleolatitude currently known.http://zoobank.org/urn:lsid:zoobank.org:pub:43AAA541-031C-4EE1-B819-4846EBBD1BBBSUPPLEMENTAL DATA?Supplemental materials are available for this article for free at www.tandfonline.com/UJVPCitation for this article: Marzola, M., O. Mateus, N. H. Shubin, and L. B. Clemmensen. 2017. Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland). Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2017.1303501.ABSTRACTCyclotosaurus naraserluki, sp. nov., is a new Late Triassic capitosaurid amphibian from lacustrine deposits in the Fleming Fjord Formation of the Jameson Land Basin in Greenland. It is based on a fairly complete and well-preserved skull associated with two vertebral intercentra. Previously reported as Cyclotosaurus cf. posthumus, C. naraserluki is unique among cyclotosaurs for having the postorbitals embaying the supratemporals posteromedially. The anterior palatal vacuity presents an autapomorphic complete subdivision by a wide medial premaxillary-vomerine bony connection. The parasphenoid projects between the pterygoids and the exoccipitals, preventing a suture between the two, a primitive condition shared with Rhinesuchidae, Eryosuchus, and Kupferzellia. Within Cyclotosaurus, the Greenlandic skull has a distinctive combination of circular choanae (shared with C. ebrachensis, C. posthumus, and C. robustus) and a convex posteromedial margin of the tabulars (also present in C. ebrachensis and C. intermedius). A phylogenetic analysis indicates that C. naraserluki is the sister taxon of the middle Norian C. mordax from southern Germany, with which it shares a pair of premaxillary foramina. Cyclotosaurus is one of the most successful and diverse genera of Late Triassic temnospondyls, with at least eight species reported from middle Carnian to late Norian. Cyclotosaurus naraserluki is the largest amphibian ever reported from Greenland and one of the Late Triassic vertebrates with the highest northern paleolatitude currently known.http://zoobank.org/urn:lsid:zoobank.org:pub:43AAA541-031C-4EE1-B819-4846EBBD1BBBSUPPLEMENTAL DATA?Supplemental materials are available for this article for free at www.tandfonline.com/UJVPCitation for this article: Marzola, M., O. Mateus, N. H. Shubin, and L. B. Clemmensen. 2017. Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland). Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2017.1303501.

Ripple, WJ, Wolf C, Newsome TM, Galetti M, Alamgir M, Crist E, Mahmoud MI, Laurance WF, other scientist signatories +15 364.  2017.  World Scientists’ Warning to Humanity: A Second Notice. BioScience. :bix125. Abstractripple_et_al_2017_world_scientists_warning_to_humanity-_a_second_notice_bix125.pdfWebsite

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Waskow, K, Mateus O.  2017.  Dorsal rib histology of dinosaurs and a crocodylomorph from western Portugal: Skeletochronological implications on age determination and life history traits. Comptes Rendus Palevol. 16:425-439. Abstractwaskowmateus2017_histology.pdfWebsite

Abstract Bone histology is an important tool for uncovering life history traits of extinct animals, particularly those that lack modern analogs, such as the non-avian dinosaurs. In most studies, histological analyses preferentially focus on long bones for understanding growth rates and determining age. Here we show, by analyzing ornithischians (a stegosaur and an ornithopod), saurischians (a sauropod and a theropod), and a crocodile, rib histology is a suitable alternative. The estimated age for all sampled taxa ranges between 14 to 17 years for Lourinhanosaurus antunesi and 27 to 31 years estimated for Draconyx loureiroi. The theropod Baryonyx was skeletally mature around 23–25 years of age but showed unfused neurocentral sutures, a paedomorphic feature possibly related to aquatic locomotion. Our results show that ribs can contain a nearly complete growth record, and reveal important information about individual age, point of sexual maturity, and, in some cases, sex. Because ribs are more available than long bones, this method opens new possibilities for studying rare and incomplete fossils, including holotypes.

Lallensack, JN, Klein H, Milàn J, Wings O, Mateus O, Clemmensen LB.  2017.  Sauropodomorph dinosaur trackways from the Fleming Fjord Formation of East Greenland: Evidence for Late Triassic sauropods. Acta Palaeontologica Polonica. 62(4):833-843. Abstractlallensack_et_al_2017_-_sauropodomorph_tracks_greenland.pdf

The Late Triassic (Norian–early Rhaetian) Fleming Fjord Formation of central East Greenland preserves a diverse fossil fauna, including both body and trace fossils. Trackways of large quadrupedal archosaurs, although already reported in 1994 and mentioned in subsequent publications, are here described and figured in detail for the first time, based on photogrammetric data collected during fieldwork in 2012. Two trackways can be referred to Eosauropus, while a third, bipedal trackway may be referred to Evazoum, both of which have been considered to represent sauropodomorph dinosaur tracks. Both the Evazoum and the Eosauropus trackways are distinctly larger than other trackways referred to the respective ichnogenera. The trackmaker of the best preserved Eosauropus trackway is constrained using a synapomorphy-based approach. The quadrupedal posture, the entaxonic pes structure, and five weight-bearing digits indicate a derived sauropodiform trackmaker. Other features exhibited by the tracks, including the semi-digitigrade pes and the laterally deflected unguals, are commonly considered synapomorphies of more exclusive clades within Sauropoda. The present trackway documents an early acquisition of a eusauropod-like pes anatomy while retaining a well-developed claw on pedal digit IV, which is reduced in eusauropods. Although unequivocal evidence for sauropod dinosaurs is no older than the Early Jurassic, the present trackway provides evidence for a possible Triassic origin of the group.

Moreno-Azanza, M, Bauluz B, Canudo JI, Mateus O.  2017.  The conservative structure of the ornithopod eggshell: electron backscatter diffraction characterization of Guegoolithus turolensis from the Early Cretaceous of Spain, 2017. Journal of Iberian Geology. :1-9. Abstractthe_conservative_structure_of_the_ornithopod_eggshell.pdfWebsite

The Spheroolithidae oospecies Guegoolithus turolensis, putatively attributed to non-hadrosauroid styracosterns was first described in the Barremian of the Iberian Basin, and later reported in the Valanginian–Hauterivian of the Cameros Basin, with both occurrences separated by a few hundred kilometres but by over 10 million years.

Mateus, O, Marzola M, Schulp AS, Jacobs LL, Polcyn MJ, Pervov V, Gonçalves AO, Morais ML.  2017.  Angolan ichnosite in a diamond mine shows the presence of a large terrestrial mammaliamorph, a crocodylomorph, and sauropod dinosaurs in the Early Cretaceous of Africa. Palaeogeography, Palaeoclimatology, Palaeoecology. 471:220-232. Abstractmateus_et_al_2017_angolan_ichnosite_catoca.pdfWebsite

Abstract We report here new and the first mammaliamorph tracks from the Early Cretaceous of Africa. The tracksite, that also bears crocodylomorph and sauropod dinosaurian tracks, is in the Catoca diamond mine, Lunda Sul Province, Angola. The mammaliamorph tracks have a unique morphology, attributed to Catocapes angolanus ichnogen. et ichnosp. nov. and present an anterolateral projection of digit I and V. The tracks with an average length of 2.7 cm and width of 3.2 cm are the largest mammaliamorph tracks known from the Early Cretaceous unmatched in size in the skeletal fossil record. The crocodylomorph trackways and tracks are attributed to Angolaichnus adamanticus ichnogen. et ichnosp. nov. (‘ichnofamily’ Batrachopodidae) and present a functionally pentadactyl pes, an extremely outwardly rotated handprint, and an unusual tetradactyl and plantigrade manus. One medium-sized sauropod dinosaur trackway preserved skin impressions of a trackmaker with stride length of 1.6 m; a second is that of a small-sized sauropod trackmaker with a pace length of 75 cm.

Tschopp, E, Tschopp FA, Mateus O.  2017.  The Overlap Index, a tool to quantify the amount of anatomical overlap among groups of incomplete terminal taxa in phylogenetic analyses. Journal of Vertebrate Paleontology, Program and Abstracts. 2017:205-206. Abstract
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Tschopp, E, Mateus O.  2017.  Osteology of Galeamopus pabsti sp. nov. (Sauropoda: Diplodocidae), with implications for neurocentral closure timing, and the cervico-dorsal transition in diplodocids. PeerJ. 5:e3179. Abstracttschopp__mateus_2017_osteology_of_galeamopus_pabsti_sp.__nov._sauropoda__diplodocidae_.pdfWebsite

Diplodocids are among the best known sauropod dinosaurs. Numerous specimens of currently 15 accepted species belonging to ten genera have been reported from the Late Jurassic to Early Cretaceous of North and South America, Europe, and Africa. The highest diversity is known from the Upper Jurassic Morrison Formation of the western United States: a recent review recognized 12 valid, named species, and possibly three additional, yet unnamed ones. One of these is herein described in detail and referred to the genus \textit{Galeamopus}. The holotype specimen of \textit{Galeamopus pabsti} sp. nov., SMA 0011, is represented by material from all body parts but the tail, and was found at the Howe-Scott Quarry in the northern Bighorn Basin in Wyoming, USA. Autapomorphic features of the new species include a horizontal canal on the maxilla that connects the posterior margin of the preantorbital and the ventral margin of the antorbital fenestrae, a vertical midline groove marking the sagittal nuchal crest, the presence of a large foramen connecting the postzygapophyseal centrodiapophyseal fossa and the spinopostzygapophyseal fossa of mid- and posterior cervical vertebrae, a very robust humerus, a laterally placed, rugose tubercle on the concave proximal portion of the anterior surface of the humerus, a relatively stout radius, the absence of a distinct ambiens process on the pubis, and a distinctly concave posteroventral margin of the ascending process of the astragalus. In addition to the holotype specimen SMA 0011, the skull USNM 2673 can also be referred to \textit{Galeamopus pabsti}. Histology shows that the type specimen SMA 0011 is sexually mature, although neurocentral closure was not completed at the time of death. Because SMA 0011 has highly pneumatized cervical vertebrae, the development of the lamination appears a more important indicator for individual age than neurocentral fusion patterns. SMA 0011 is one of very few sauropod specimens that preserves the cervico-dorsal transition in both vertebrae and ribs. The association of ribs with their respective vertebrae shows that the transition between cervical and dorsal vertebrae is significantly different in \textit{Galeamopus pabsti} than in \textit{Diplodocus carnegii} or \textit{Apatosaurus louisae}, being represented by a considerable shortening of the centra from the last cervical to the first dorsal vertebra. Diplodocids show a surprisingly high diversity in the Morrison Formation. This can possibly be explained by a combination of geographical and temporal segregation, and niche partitioning.

Russo, J, Mateus O, Marzola M, Balbino A.  2017.  Two new ootaxa from the late Jurassic: The oldest record of crocodylomorph eggs, from the Lourinhã Formation, Portugal, 03. PLOS ONE. 12:1-23., Number 3: Public Library of Science Abstractrusso_et_al_2017_two_new_ootaxa.pdfWebsite

The Late Jurassic Lourinhã Formation is known for its abundant remains of dinosaurs, crocodylomorphs and other vertebrates. Among this record are nine localities that have produced either dinosaur embryos, eggs or eggshell fragments. Herein, we describe and identify the first crocodiloid morphotype eggs and eggshells from the Lourinhã Formation, from five occurrences. One clutch from Cambelas, composed of 13 eggs, eggshell fragments from Casal da Rola and Peralta, one crushed egg and eggshells from Paimogo North, and four crushed eggs as well as eggshell fragments from Paimogo South. We observed and confirmed diagnostic morphological characters for crocodiloid eggshells and which are consistent with a crocodylomorph affinity, such as the ellipsoidal shape, wedge-shaped shell units, triangular extinction under cross-polarized light, and tabular ultrastructure. This material is distinctive enough to propose two new ootaxa within the oofamily Krokolithidae, Suchoolithus portucalensis, oogen. and oosp. nov., for the material from Cambelas, the most complete clutch known for crocodiloid eggs, and Krokolithes dinophilus, oosp. nov., for the remaining material. These are the oldest crocodylomorph eggs known, extending the fossil record for this group to the Late Jurassic. Furthermore, except for the clutch from Cambelas, the material was found with theropod eggs and nests, in the other four occurrences, which seem to suggest some form of biological relationship, still unclear at this point.

Tschopp, E, Tschopp FA, Mateus O.  2017.  Overlap Indices: Tools to quantify the amount of anatomical overlap among groups of incomplete terminal taxa in phylogenetic analyses. Acta Zoologica. 99(2):169-176. Abstracttschopp_et_al-2017-acta_zoologica_overlap_indices_tools_to_quantify_the_amount.pdfWebsite

Phylogenetic analyses of morphological data are often characterized by missing data due to incomplete operational taxonomic units, as in fossils. This incomplete knowledge derives from various reasons, including—in the case of fossils—the numerous filters an organism has to pass through during taphonomy, fossilization, weathering and collecting. Whereas several methods have been proposed to address issues raised by the inclusion of incomplete terminal taxa, until recently no tool existed to easily quantify the amount of anatomical overlap within a particular clade. The Overlap Indices provide such values and might prove useful for comparative cladistics. We herein describe these new indices and their applications in detail and provide an example file for their calculation. A case study of diplodocid sauropod dinosaurs shows how the Overlap Indices will help to explore and quantify, which one of a number of conflicting tree topologies is supported by more anatomical traits, which skeletal regions are underrepresented in a particular phylogenetic matrix, and which taxon would improve character state score completeness.