Publications

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Conference Proceedings
Ríos, M., Estraviz-López D., Martino R., Lohmann P., Mateus O., & Solounias N. (2022).  The first Sivatheriine (Artiodactyla, Giraffidae) from the Middle Miocene of Portugal. XIX Annual conference of the European Association of Vertebrate Palaeontologists (19th EAVP)At: Benevento, Italy. 167. Abstracteavp_2022_abstractvolume-176.pdf

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Mateus, O., Jacobs L. L., Polcyn M. J., Myers T. S., & Schulp A. S. (2015).  The fossil record of testudines from angola from the turonian to oligocene. Society of Vertebrate Paleontology Annual Meeting. 177., Dallasmateus_et_al_2015_testudines_angola_svp_abstract.pdf
Strganac, C., Jacobs L. L., Ferguson K. M., Polcyn M. J., Mateus O., Schulp A. S., & Morais M. L. (2013).  Late Cretaceous marine reptiles and cooling at the South Atlantic coast inferred through stable oxygen isotopes of Inoceramus from the Namibe Basin, Angola. Geological Society of America Abstracts with Programs. Vol. 45, No. 7, p.0.
Polcyn, M. J., Jacobs L. L., Mateus O., Schulp A. S., Strganac C., Araújo R., Graf J. F., Vineyard D., & Myers T. S. (2013).  A marine vertebrate assemblage from the Campanian-Maastrichtian boundary at Bentiaba, Angola. Geological Society of America Abstracts with Programs. Vol. 45, No. 7, p.0. polcyn_et_al_2013_abstract__a_marine_vertebrate_assemblage_from_the_campanian-maastrichtian_boundary_at_bentiaba_angola_2013__gsa_27-30_october_2013.pdf
Graf, J., Jacobs L. L., Polcyn M. J., Mateus O., & Schulp A. S. (2011).  New fossil whales from Angola. 71st Annual Meeting of the Society of Vertebrate Paleontology. 119., Jan: Abstracts of the 71st Annual Meeting of the Society of Vertebrate Paleontologygraf_et_al_mateus_2011_fossil_whales_from_angola_svp11abstracts.pdf
Steyer, J. S., Mateus O., Butler R. J., Brusatte S. L., & Whiteside J. H. (2011).  A new metoposaurid (temnospondyl) bonebed from the Late Triassic of Portugal. 71st Annual Meeting of the Society of Vertebrate Paleontology. 200., Jan: Abstracts of the 71st Annual Meeting of the Society of Vertebrate Paleontology Abstractsteyer_mateus_et_al_2011_._a_new_metoposaurid_temnospondyl_bonebed_from_the_late_triassic_of_portugal_svp11abstracts.pdf

The end-Triassic extinction event (ETE), considered one of the ‘Big Five’ mass extinctions, marks a dividing line between early Mesozoic vertebrate assemblages, typically including abundant temnospondyls, basal synapsids and basal archosaurs, and ‘typical’ Mesozoic faunas dominated by dinosaurs, pterosaurs, crocodylomorphs, turtles and mammaliaforms.
Recent geochemical work has provided strong evidence that the ETE is synchronous with, and likely caused by, the emplacement of the Central Atlantic magmatic province (CAMP).
However, stratigraphic sections containing both terrestrial vertebrates and CAMP basalts are scarce, complicating attempts to examine terrestrial faunal changes during this extinction event. The Triassic–Jurassic Algarve Basin, southern Portugal, is an extensional rift basin

to-marginal marine red beds (the ‘Grés de Silves’ Group) interbedded with CAMP basalts.

bonebed from the interval ‘AB1’ of the Grés de Silves. Preliminary excavations yielded at least nine well-preserved temnospondyl individuals represented by partial to nearly complete skulls and disarticulated postcranial elements of juvenile to adult ages. Nearly all material appears to represent a single species of metoposaurid referable to the genus Metoposaurus, well known from the late Carnian–early Norian of Germany and Poland. A number of characters of the occiput and mandible suggest that the Algarve material may represent a new species. This new material provides new data on the diversity and paleogeographical distribution of the metoposaurids, a highly autapomorphic and peculiar group composed of large aquatic carnivores with a unique elongated but brevirostral skull. This taxon also provides

Horizon may be within or close to the late Carnian–early Norian. Additional bone-bearing horizons within the ‘Grés de Silves’ provide a rare opportunity to examine terrestrial faunal change in the lead-up to the ETE.

Jacobs, L., Polcyn M., Araújo R., Strganac C., & Mateus O. (2010).  Physical drivers of evolution and the history of the marine tetrapod fauna of Angola. Annual Meeting of the Society of Vertebrate Paleontology. 110A., Jan Abstractjacobs_et_al_mateus_2010_physical_drivers_marine_tetrapod_fauna_of_angola_svp10abstracts.pdf

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Araújo, R., Polcyn M., Jacobs L. L., Mateus O., & Schulp A. S. (2011).  Plesiosaur structural extreme from the Maastrichtian of Angola. 71st Annual Meeting of the Society of Vertebrate Paleontology. 63., Jan: Abstracts of the 71st Annual Meeting of the Society of Vertebrate Paleontology Abstractaraujo_et_al_mateus_2011_plesiosaur_extreme_angola_svp11abstracts.pdf

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Salminen, J., Dinis J., & Mateus O. (2013).  Preliminary magnetostratigraphy for Jurassic/Cretaceous transition in Porto da Calada, Portugal. In: Veikkolainen, T., Suhonen, K., Näränen, J., Kauristie, K., and Kaasalainen, S. (eds.). XXVI Geofysiikan päivät,. 119-122., May 21-22 2013 in Helsinkisalminen.johanna_gfp2013_portugal_preliminary_magnetostratigraphy_for_jurassic_cretaceous_transition_in.pdf
Costa, F., Silva T., Fernandes J., Calvo R., & Mateus O. (2017).  Retracing the history of a stegosaurian dinosaur discovery in Portugal and the importance of record-keeping in Palaentology. Abstract book of the XV Encuentro de Jóvenes Investigadores en Paleontología/XV Encontro de Jovens Investigadores em Paleontologia, Lisboa, 428 pp.. 119-124. Abstractcosta_et_al_2017_retracing_the_history_-_2017.pdf

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Sousa, J., & Mateus O. (2018).  A review of Ichthyosauria from Portugal. XVI Annual Meeting of the European Association of Vertebrate Palaeontologists. 179., Caparica, Portugal June 26th-July 1st, 2018 Abstractsousa_e_mateus_2018_eavp_abstract.pdf

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Rotatori, F. M., Silva B. C., Moreno-Azanza M., & Mateus O. (2021).  A systematic overview of medium and large-sized iguanodontians (Dinosauria, Ornithopoda) from the Upper Jurassic Lourinhã Formation, Portugal. XIX Encontro de Jovens Investigadores em Paleontologia. 62-63. Abstractrotatoriet_l_2021_ejip.pdf

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Brusatte, S. L., Butler R. J., Mateus O., Steyer J. S., & Whiteside J. H. (2013).  Terrestrial vertebrates from the Late Triassic of Portugal: new records of temnospondyls and archosauriforms from a Pangaean rift sequence. 61st Symposium on Vertebrate Palaeontology and Comparative Anatomy. 15-16., Edinburgh Abstractbrusatte_et_al_2013_triassic_algarve_portugal_temnospondyls_svpca.org_years_2013_edinburgh_abstracts.pdf

The Late Triassic (ca. 237-201.5 million years ago) was a transitional interval in the evolution of terrestrial ecosystems, during which ‘modern’ clades such as archosaurs and mammals were radiating while ‘archaic’ groups such as temnospondyl amphibians and basal synapsids remained abundant. Little is known about the Triassic terrestrial (nonmarine) vertebrates of the Iberian Peninsula. The Algarve Basin of southern Portugal is an extensional rift basin formed during the breakup of Pangaea, which is filled with terrestrial, lacustrine, and marginal marine siliciclastics of the Grés de Silves Formation, interbedded with CAMP basalts that mark the end-Triassic extinction (radioisotopically dated to ~198-201.5 Ma). Since 2009, our field project in the Algarve has discovered numerous vertebrate specimens within the Grés de Silves, including a monodominant bonebed containing hundreds of specimens of metoposaurids, a peculiar group of temnospondyls that filled crocodile-like predatory niches in lacustrine and fluvial environments. These specimens appear to belong to a new species of Metoposaurus, similar to M. diagnosticus and M. krasiejowensis from central Europe but possessing several putative autapomorphies of the braincase and lower jaw. We also discovered a mandible of a phytosaur, the first specimen of these long-snouted, semi-aquatic archosauriforms from the Iberian Peninsula. These discoveries of characteristic Carnian Norian taxa indicate that the fossil-bearing portion of the Grés de Silves is Late Triassic in age, and provide further evidence that metoposaurids and phytosaurs commonly occurred together in low palaeolatitudes during this time.

Polcyn, M. J., Jacobs L. L., Schulp A. S., Mateus O., & Araújo R. (2015).  Tethyan and Weddellian biogeographic mixing in the Maastrichtian of Angola. Annual Meeting of the Society of Vertebrate Paleontology. 196., Dallas, TXpolcyn_etal2015_mix_fauna_angola_svp_abstract.pdf
Polcyn, M. J., Jacobs L. L., Schulp A. S., & Mateus O. (2015).  Trolling the Cretaceous Seas: Marine Amniotes of Two West Coast Margins. Geological Society of America Annual Meeting. 55.: Geological Society of America Abstracts with Programs. Vol. 47, No. 4, p.55 Abstract

In this session we review the Upper Cretaceous marine amniote records from the west coasts of North America and Africa. Recent work by our group in Angola, on the west coast of Africa, has opened up new fossiliferous localities, producing well-preserved turtles, plesiosaurs, and mosasaurs, ranging in age from Late Turonian to Late Maastrichtian. These African localities were deposited in arid latitudes and highly productive upwelling zones along the passive margin of a growing South Atlantic Ocean. The fossil record of Cretaceous marine amniotes from the West Coast of North America is relatively meager when compared to the African record and the prolific fossil beds laid down in the epicontinental seas of the Western Interior Seaway and northern Europe. Nonetheless, these localities provide an important glimpse of a marine ecosystem that developed on the active margins of a deep ocean basin. Historically considered to be depauperate and endemic, the west coast fauna was characterized by unusual forms such as Plotosaurus, arguably one of the most derived mosasaurs; however, in recent years, additional taxa have been described, revealing species diversity and ecological partitioning within these communities and in some cases, faunal interchange with other regions. The large quantity of well-preserved fossils from the west coast of Africa is influenced in part by its paleogeographic position, deposited within highly productive areas of Hadley Cell controlled upwelling zones. By contrast, the North American west coast localities have been deposited in temperate and higher latitudes since the Late Cretaceous. Nonetheless, the North American and African faunas share some common characteristics in a possessing a mix of endemic and more cosmopolitan forms. Habitat partitioning reflected in tooth form and body size is comparable between the Angolan and the North American west coast, and there is remarkable convergence in taxa which appear to exploit certain like-niches.

Moreno-Azanza, M., Ezquerro L., Gasca J. M., Díaz-Berenguer E., Nuñez-Lahuerta C., Pérez-Pueyo M., Puértolas-Pascual E., Sellés A. G., Vila B., & Laita E. (2021).  Two latest Cretaceous egg localities in the external sierras (Southern Pyrenees, Huesca Province, NE Spain). Palaeovertebrata - XVIII annual conference of the European Association of Vertebrate Palaeontologists. 44, 120-121. Abstractmoreno-azanza_et_al_2021_eggs_eavp_abstract.pdf

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Saleiro, A., & Mateus O. (2017).  Upper Jurassic bonebeds around Ten Sleep, Wyoming, USA: overview and stratigraphy. Abstract book of the XV Encuentro de Jóvenes Investigadores en Paleontología/XV Encontro de Jovens Investigadores em Paleontologia, Lisboa, 428 pp.. 357-361. Abstractsaleiro__mateus_2017._upper_jurassic_bonebeds_around_ten_sleep_wyoming_usa-overview_and_stratigraphy..pdf

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Silva, T., Costa F., Fernandes J., Calvo R., & Mateus O. (2016).  The use of a portable X-ray fluorescence analyzer in the reconstitution of dinosaur fossils. European Conference on X-Ray Spectrometry. , June 19 – 24, 2016, Gothenburg, Sweden Abstract
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Journal Article
Young, M. T., Hua S., Steel L., Foffa D., Brusatte S. L., Thüring S., Mateus O., Ruiz-Omeñaca J. I., Havlik P., Lepage Y., & De Andrade M. B. (2015).  Addendum to ?Revision of the Late Jurassic teleosaurid genus Machimosaurus (Crocodylomorpha, Thalattosuchia)?. Royal Society Open Science. 2, , Number 2 Abstract
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Young, M. T., Hua S., Steel L., Foffa D., Brusatte S. L., Thüring S., Mateus O., Ruiz-Omeñaca J. I., Havlik P., Lepage Y., & de Andrade M. B. (2015).  Addendum to ‘Revision of the Late Jurassic teleosaurid genus Machimosaurus (Crocodylomorpha, Thalattosuchia)’. Royal Society Open Science. 2, , Number 2: The Royal Society Abstractyoung_et_al_2015_addendum_to_revision_of_the_late_jurassic_teleosaurid_genus_machimosaurus_crocodylomorpha_thalattosuchia.pdfWebsite

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Strganac, C., Ferguson, K.M, Jacobs J. L., Polcyn M. J., & Mateus O. (2012).  Age and paleoecology of mosasaurs and plesiosaurs from the Late Cretaceous South Atlantic margin at Bentiaba, Angola. Journal of Vertebrate Paleontology, Program and Abstracts, 2012. 180.strganac_et_al_mateus_2012_age_bentiaba_angola_2012_svp_abstract.pdf
Mateus, O., Marzola M., Schulp A. S., Jacobs L. L., Polcyn M. J., Pervov V., Gonçalves A. O., & Morais M. L. (2017).  Angolan ichnosite in a diamond mine shows the presence of a large terrestrial mammaliamorph, a crocodylomorph, and sauropod dinosaurs in the Early Cretaceous of Africa. Palaeogeography, Palaeoclimatology, Palaeoecology. 471, 220 - 232. Abstractmateus_et_al_2017_angolan_ichnosite_catoca.pdfWebsite

Abstract We report here new and the first mammaliamorph tracks from the Early Cretaceous of Africa. The tracksite, that also bears crocodylomorph and sauropod dinosaurian tracks, is in the Catoca diamond mine, Lunda Sul Province, Angola. The mammaliamorph tracks have a unique morphology, attributed to Catocapes angolanus ichnogen. et ichnosp. nov. and present an anterolateral projection of digit I and V. The tracks with an average length of 2.7 cm and width of 3.2 cm are the largest mammaliamorph tracks known from the Early Cretaceous unmatched in size in the skeletal fossil record. The crocodylomorph trackways and tracks are attributed to Angolaichnus adamanticus ichnogen. et ichnosp. nov. (‘ichnofamily’ Batrachopodidae) and present a functionally pentadactyl pes, an extremely outwardly rotated handprint, and an unusual tetradactyl and plantigrade manus. One medium-sized sauropod dinosaur trackway preserved skin impressions of a trackmaker with stride length of 1.6 m; a second is that of a small-sized sauropod trackmaker with a pace length of 75 cm.

Mateus, O., Marzola M., Schulp A. S., Jacobs L. L., Polcyn M. J., Pervov V., Gonçalves A. O. {\'ı}mpio, & Morais M. L. (2017).  Angolan ichnosite in a diamond mine shows the presence of a large terrestrial mammaliamorph, a crocodylomorph, and sauropod dinosaurs in the Early Cretaceous of Africa. Palaeogeography, Palaeoclimatology, Palaeoecology. 471, 220–232., apr: Elsevier {BV} AbstractWebsite
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Mateus, O., Jacobs L. L., Schulp A. S., Polcyn M. J., Tavares T. S., Neto A. B., Morais M. L. {\'ı}sa, & Antunes M. T. (2011).  Angolatitan adamastor, a new sauropod dinosaur and the first record from Angola. Anais da Academia Brasileira de Ciências. 83, 221–233., Number 1: {FapUNIFESP} ({SciELO}) AbstractWebsite
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Mateus, O., Mateus O., Jacobs L. L., Schulp A. S., Polcyn M. J., Tavares T. S., Neto A. B., Morais M. L., & Antunes M. T. (2011).  Angolatitan adamastor, a new sauropod dinosaur and the first record from Angola. Anais da Academia Brasileira de Ciências. Abstract
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Mateus, O., Jacobs L. L., Schulp A. S., Polcyn M. J., Tavares T. S., Neto A. B., Morais M. L., & Antunes M. T. (2011).  Angolatitan adamastor, a new sauropod dinosaur and the first record from Angola.. Anais da Academia Brasileira de Ciências. 83, 221-233., Jan Abstractmateus_et_al_2011_angolatitan_adamastor_sauropod.pdfWebsite

A forelimb of a new sauropod dinosaur (Angolatitan adamastor n. gen. et sp.) from the Late Turonian of Iembe (Bengo Province) represents the first dinosaur discovery in Angola, and is one of the few occurrences of sauropod dinosaurs in sub-Saharan Africa collected with good chronological controls. The marginal marine sediments yielding the specimen are reported to be late Turonian in age and, thus it represents a non-titanosaurian sauropod in sub-Saharan Africa at a time taken to be dominated by titanosaurian forms. Moreover, Angolatitan adamastor is the only basal Somphospondyli known in the Late Cretaceous which implies in the existence of relict forms in Africa.

Sander, P. M., Mateus O., Laven T., & Knotschke N. (2006).  Bone histology indicates insular dwarfism in a new Late Jurassic sauropod dinosaur. Nature. 441, 739-741., Jan Abstractsander_mateus_et_al_2006_europasaurus_sauropod_histology_drwarfism_nature.pdf

Sauropod dinosaurs were the largest animals ever to inhabit the land, with truly gigantic forms in at least three lineages1, 2, 3. Small species with an adult body mass less than five tonnes are very rare4, 5, and small sauropod bones generally represent juveniles. Here we describe a new diminutive species of basal macronarian sauropod, Europasaurus holgeri gen. et sp. nov., and on the basis of bone histology we show it to have been a dwarf species. The fossils, including excellent skull material, come from Kimmeridgian marine beds of northern Germany6, 7, and record more than 11 individuals of sauropods 1.7 to 6.2 m in total body length. Morphological overlap between partial skeletons and isolated bones links all material to the same new taxon. Cortical histology of femora and tibiae indicates that size differences within the specimens are due to different ontogenetic stages, from juveniles to fully grown individuals. The little dinosaurs must have lived on one of the large islands around the Lower Saxony basin8. Comparison with the long-bone histology of large-bodied sauropods suggests that the island dwarf species evolved through a decrease in growth rate from its larger ancestor.

Sander, P. M., Mateus O., Laven T., & Knötschke N. (2006).  Bone histology indicates insular dwarfism in a new Late Jurassic sauropod dinosaur. Nature. 441, 739-741. Abstract
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Strganac, C., Salminen J., Jacobs L. L., Ferguson K. M., Polcyn M. J., Mateus O., Schulp A. S., Morais M. L., Tavares T. S., & Gonçalves A. O. (2014).  Carbon isotope stratigraphy and 40Ar/39Ar age of the Cretaceous South Atlantic coast, Namibe Basin, Angola. Journal of African Earth Sciences. onine, 1-11. Abstractstrganac_et_al_2014_carbon_isotope_stratigraphy_magnetostratigraphy_and_40ar_39ar_age_of.pdfWebsite

We present the δ13C and paleomagnetic stratigraphy for marine strata at the coast of southern Angola, anchored by an intercalated basalt with a whole rock 40Ar/39Ar radiometric age of 84.6 ± 1.5 Ma, being consistent with both invertebrate and vertebrate biostratigraphy. This is the first African stable carbon isotope record correlated to significant events in the global carbon cycle spanning the Late Cenomanian to Early Maastrichtian. A positive ∼ 3‰ excursion seen in bivalve shells below the basalt indicates the Cenomanian-Turonian Boundary Event at 93.9 Ma, during Oceanic Anoxic Event 2. Additional excursions above the basalt are correlated to patterns globally, including a negative ∼ 3‰ excursion near the top of the section interpreted as part of the Campanian-Maastrichtian Boundary Events. The age of the basalt ties the studied Bentiaba section to a pulse of Late Cretaceous magmatic activity around the South Atlantic and significant tectonic activity, including rotation, of the African continent.

Strganac, C., Salminen J., Jacobs L. L., Polcyn M. J., Ferguson K. M., Mateus O., Schulp A. S., Morais M. L., Tavares T. S., & Gon?alves A. O. (2014).  Carbon isotope stratigraphy, magnetostratigraphy, and 40Ar/39Ar age of the cretaceous South Atlantic coast, Namibe Basin, Angola. Journal of African Earth Sciences. 99, 452-462., Number PA2 Abstract
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Jacobs, L., Polcyn M., Mateus O., Scott M., Graf J., Kappelman J., Jacobs B., Schulp A., Morais M., & Goncalves O. (2014).  Cenozoic vertebrates of coastal Angola. Journal of Vertebrate Paleontology, Program and Abstracts, 2014. 153.jacobs_et_al._2014_cenozoic_vertebrates_of_coastal_angola.pdf
Stockdale, M., Benton M., & Mateus O. (2014).  Cracking dinosaur endothermy: paleophysiology unscrambled. Journal of Vertebrate Paleontology. Program and Abstracts, 2014, 235-236.stockdale_et_al_2014_eggshells_abstract_svp.pdf
Marx, M. P., Mateus O., Polcyn M. J., Schulp A. S., Gonçalves O. A., & Jacobs L. L. (2021).  The cranial anatomy and relationships of Cardiocorax mukulu (Plesiosauria: Elasmosauridae) from Bentiaba, Angola. PLOS ONE. 16(8), e0255773 - ., 2021/08/17: Public Library of Science Abstractmarx_et_al_2021_cardiocorax_angola.pdfWebsite

We report a new specimen of the plesiosaur Cardiocorax mukulu that includes the most complete plesiosaur skull from sub-Saharan Africa. The well-preserved three-dimensional nature of the skull offers rare insight into the cranial anatomy of elasmosaurid plesiosaurians. The new specimen of Cardiocorax mukulu was recovered from Bentiaba, Namibe Province in Angola, approximately three meters above the holotype. The new specimen also includes an atlas-axis complex, seventeen postaxial cervical vertebrae, partial ribs, a femur, and limb elements. It is identified as Cardiocorax mukulu based on an apomorphy shared with the holotype where the cervical neural spine is approximately as long anteroposteriorly as the centrum and exhibits a sinusoidal anterior margin. The new specimen is nearly identical to the holotype and previously referred material in all other aspects. Cardiocorax mukulu is returned in an early-branching or intermediate position in Elasmosauridae in four out of the six of our phylogenetic analyses. Cardiocorax mukulu lacks the elongated cervical vertebrae that is characteristic of the extremely long-necked elasmosaurines, and the broad skull with and a high number of maxillary teeth (28–40) which is characteristic of Aristonectinae. Currently, the most parsimonious explanation concerning elasmosaurid evolutionary relationships, is that Cardiocorax mukulu represents an older lineage of elasmosaurids in the Maastrichtian.

Mateus, O., Morais M. L., Schulp A. S., Jacobs L. L., & Polcyn M. J. (2006).  The Cretaceous of Angola. Journal of Vertebrate Paleontology. 26, 96-97., Number (Suppl. To 3) Abstract
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Mateus, O., Morais M. L., Schulp A. S., Jacobs L. L., & Polcyn M. J. (2006).  The Cretaceous of Angola. Journal of Vertebrate Paleontology. 26, 96–97., Number (Suppl. T Abstract
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Jacobs, L. L., Mateus O., Polcyn M. J., Schulp A. S., Scotese C. R., Goswami A., Ferguson K. M., Robbins J. A., Vineyard D. P., & Neto A. B. (2009).  Cretaceous paleogeography, paleoclimatology, and amniote biogeography of the low and mid-latitude South Atlantic Ocean. BULLETIN DE LA SOCIETE GEOLOGIQUE DE FRANCE. 180, 333-341., Jan: Univ Agostinho Neto, Univ Nova Lisboa, So Methodist Univ, Univ Texas Arlington, Museu Lourinha, Nat Hist Museum Abstractjacobs_mateus_et_al_2009_cretaceous_paleogeography_paleoclimatology_and_amniote_biogeography_of_the_south_atlantic_ocean_angola_africa_currents.pdf

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Jacobs, L. L., Mateus O., Polcyn M. J., Schulp A. S., Scotese C. R., Goswami A., Ferguson K. M., Robbins J. A., Vineyard D. P., & Neto A. B. (2009).  Cretaceous paleogeography, paleoclimatology, and amniote biogeography of the low and mid-latitude South Atlantic Ocean. Bulletin de la Societe Geologique de France. 180, 333-341., Number 4 Abstract
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Jacobs, L., Polcyn M., Mateus O., Schulp A. S., & Neto A. B. (2009).  The Cretaceous Skeleton Coast of Angola. Journal of Vertebrate Paleontology. 29, 121A., Jan Abstractjacobs_et_al_2009cretaceousskeletoncoas.pdfWebsite

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Jacobs, L., Polcyn M., Mateus O., Schulp, & Neto A. (2009).  The Cretaceous Skeleton Coast of Angola. Journal of Vertebrate Paleontology. 29, 121., Number 3 Abstract
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Jacobs, L. L., Polcyn M. J., Mateus O., Schulp A. S., & Neto A. (2009).  The Cretaceous Skeleton Coast of Angola. Journal of Vertebrate Paleontology. 29, 121–121., Number 3 Abstract
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Marzola, M., Mateus O., Shubin N. H., & Clemmensen L. B. (2017).  Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland). Journal of Vertebrate Paleontology. e1303501., 2017: Taylor & Francis Abstractmarzola_et_al_2017_cyclotosaurus_greenland.pdfWebsite

ABSTRACTCyclotosaurus naraserluki, sp. nov., is a new Late Triassic capitosaurid amphibian from lacustrine deposits in the Fleming Fjord Formation of the Jameson Land Basin in Greenland. It is based on a fairly complete and well-preserved skull associated with two vertebral intercentra. Previously reported as Cyclotosaurus cf. posthumus, C. naraserluki is unique among cyclotosaurs for having the postorbitals embaying the supratemporals posteromedially. The anterior palatal vacuity presents an autapomorphic complete subdivision by a wide medial premaxillary-vomerine bony connection. The parasphenoid projects between the pterygoids and the exoccipitals, preventing a suture between the two, a primitive condition shared with Rhinesuchidae, Eryosuchus, and Kupferzellia. Within Cyclotosaurus, the Greenlandic skull has a distinctive combination of circular choanae (shared with C. ebrachensis, C. posthumus, and C. robustus) and a convex posteromedial margin of the tabulars (also present in C. ebrachensis and C. intermedius). A phylogenetic analysis indicates that C. naraserluki is the sister taxon of the middle Norian C. mordax from southern Germany, with which it shares a pair of premaxillary foramina. Cyclotosaurus is one of the most successful and diverse genera of Late Triassic temnospondyls, with at least eight species reported from middle Carnian to late Norian. Cyclotosaurus naraserluki is the largest amphibian ever reported from Greenland and one of the Late Triassic vertebrates with the highest northern paleolatitude currently known.http://zoobank.org/urn:lsid:zoobank.org:pub:43AAA541-031C-4EE1-B819-4846EBBD1BBBSUPPLEMENTAL DATA?Supplemental materials are available for this article for free at www.tandfonline.com/UJVPCitation for this article: Marzola, M., O. Mateus, N. H. Shubin, and L. B. Clemmensen. 2017. Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland). Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2017.1303501.ABSTRACTCyclotosaurus naraserluki, sp. nov., is a new Late Triassic capitosaurid amphibian from lacustrine deposits in the Fleming Fjord Formation of the Jameson Land Basin in Greenland. It is based on a fairly complete and well-preserved skull associated with two vertebral intercentra. Previously reported as Cyclotosaurus cf. posthumus, C. naraserluki is unique among cyclotosaurs for having the postorbitals embaying the supratemporals posteromedially. The anterior palatal vacuity presents an autapomorphic complete subdivision by a wide medial premaxillary-vomerine bony connection. The parasphenoid projects between the pterygoids and the exoccipitals, preventing a suture between the two, a primitive condition shared with Rhinesuchidae, Eryosuchus, and Kupferzellia. Within Cyclotosaurus, the Greenlandic skull has a distinctive combination of circular choanae (shared with C. ebrachensis, C. posthumus, and C. robustus) and a convex posteromedial margin of the tabulars (also present in C. ebrachensis and C. intermedius). A phylogenetic analysis indicates that C. naraserluki is the sister taxon of the middle Norian C. mordax from southern Germany, with which it shares a pair of premaxillary foramina. Cyclotosaurus is one of the most successful and diverse genera of Late Triassic temnospondyls, with at least eight species reported from middle Carnian to late Norian. Cyclotosaurus naraserluki is the largest amphibian ever reported from Greenland and one of the Late Triassic vertebrates with the highest northern paleolatitude currently known.http://zoobank.org/urn:lsid:zoobank.org:pub:43AAA541-031C-4EE1-B819-4846EBBD1BBBSUPPLEMENTAL DATA?Supplemental materials are available for this article for free at www.tandfonline.com/UJVPCitation for this article: Marzola, M., O. Mateus, N. H. Shubin, and L. B. Clemmensen. 2017. Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland). Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2017.1303501.

Marzola, M., Mateus O., Shubin N. H., & Clemmensen L. B. (2017).  Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland). Journal of Vertebrate Paleontology. e1303501., may: Informa {UK} Limited AbstractWebsite
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Marzola, M., Mateus O., Schulp A., Jacobs L., Polcyn M., & Pervov V. (2014).  Early Cretaceous tracks of a large mammaliamorph, a crocodylomorph, and dinosaurs from an Angolan diamond mine. Journal of Vertebrate Paleontology, Program and Abstracts, 2014. 181.marzola_et_al_2014._cretaceous_tracks_mammaliamorph_a_crocodilomorph_angolan_diamond_mine.pdf
Araújo, R., Castanhinha R., Martins R. M. S., Mateus O., Hendrickx C., Beckmann F., Schell N., & Alves L. C. (2013).  Filling the gaps of dinosaur eggshell phylogeny: Late Jurassic Theropod clutch with embryos from Portugal. Scientific Reports. 3(1924), , 2013/05/30/onlin: Macmillan Publishers Limited. All rights reserved Abstractaraujo_et_al_2013_filling_the_gaps_of_dinosaur_eggshell_phylogeny_late_jurassic_theropod_clutch_with_embryos_from_portugal.pdf

The non-avian saurischians that have associated eggshells and embryos are represented only by the sauropodomorph Massospondylus and Coelurosauria (derived theropods), thus missing the basal theropod representatives. We report a dinosaur clutch containing several crushed eggs and embryonic material ascribed to the megalosaurid theropod Torvosaurus. It represents the first associated eggshells and embryos of megalosauroids, thus filling an important phylogenetic gap between two distantly related groups of saurischians. These fossils represent the only unequivocal basal theropod embryos found to date. The assemblage was found in early Tithonian fluvial overbank deposits of the Lourinhã Formation in West Portugal. The morphological, microstructural and chemical characterization results of the eggshell fragments indicate very mild diagenesis. Furthermore, these fossils allow unambiguous association of basal theropod osteology with a specific and unique new eggshell morphology.

Araújo, R., Castanhinha R., Martins R. M. S., Mateus O., Hendrickx C., Beckmann F., Schell N., & Alves L. C. (2013).  Filling the gaps of dinosaur eggshell phylogeny: Late Jurassic Theropod clutch with embryos from Portugal. Scientific Reports. 3, , may: Nature Publishing Group AbstractWebsite
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Mateus, O., Butler R. J., Brusatte S. L., Whiteside J. H., & Steyer S. J. (2014).  The first phytosaur (Diapsida, Archosauriformes) from the Late Triassic of the Iberian Peninsula. Journal of Vertebrate Paleontology. 34(4), 970-975.mateus_et_al_2014_first_phytosaur_algarve_portugal_jvp.pdfWebsite
Mateus, O., Butler R. J., Brusatte S. L., Whiteside J. H., & Steyer J. S. (2014).  The first phytosaur (Diapsida, Archosauriformes) from the Late Triassic of the Iberian Peninsula. Journal of Vertebrate Paleontology. 34, 970-975., Number 4 Abstract
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Mateus, O., Butler R. J., Brusatte S. L., Whiteside J. H., & Steyer J. S. (2014).  The first phytosaur (Diapsida, Archosauriformes) from the Late Triassic of the Iberian Peninsula. Journal of Vertebrate Paleontology. 34, 970–975., Number 4 Abstract
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Pereira, A. M., Silva M. M., & Mateus O. (2022).  First record of Phymactis papillosa (Lesson, 1830), a Pacific south sea anemone in European shores. Journal of the Marine Biological Association of the United Kingdom. 1–4.: Cambridge University Press Abstractfirst-record-of-phymactis-papillosa-lesson-1830-a-pacific-south-sea-anemone-in-european-shores.pdfWebsite

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Mallison, H., Schwarz-Wings D., Tsai H., Holliday C., & Mateus O. (2014).  Fossil longbone cartilage preserved in stegosaurs?. Journal of Vertebrate Paleontology. Program and Abstracts, 2014, 176.mallison_et_al._2014_fossil_longbone_cartilage_preserved_in_stegosaurs.pdf