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Adams, T. L., Polcyn M. J., Mateus O., Winkler D. A., & Jacobs L. L. (2010).  New occurrence of the long-snouted crocodyliform,Terminonaris cf.T. robusta, from Woodine Formation (Cenomanian) ot Texas. 52A., Jan Abstract

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Allain, R., Taquet P., Battail B., Dejax J., Richir P., Veran M., Limon-Duparcmeur F., & et al (1999).  A new genus of sauropod dinosaur from the Gres superieurs formation (Aptian-Albian) of Laos. Comptes Rendus de l'Academie des Sciences Series IIA Earth and Planetary Science. 329, 609–616., Number 8 Abstract
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Allain, R., Taquet P., Battail B., Dejax J., Richir P., Véran M., Limon-Duparcmeur F., & et al (1999).  A new genus of sauropod dinosaur from the Gres superieurs formation (Aptian-Albian) of Laos | Un nouveau genre de dinosaure sauropode de la formation des Gres superieurs (Aptien-Albien) du Laos. Comptes Rendus de l'Academie de Sciences - Serie IIa: Sciences de la Terre et des Planetes. 329, 609–616., Number 8 Abstract
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Araújo, R., Polcyn M. J., Lindgren J., Jacobs L. L., Schulp A. S., Mateus O., Gonçalves O. A., & Morais M. - L. (2015).  New aristonectine elasmosaurid plesiosaur specimens from the Early Maastrichtian of Angola and comments on paedomorphism in plesiosaurs. Netherlands Journal of Geosciences. FirstView, 1–16., 2 Abstractaraujo_et_al_2015_paedomorphism-libre.pdfWebsite

ABSTRACT New elasmosaurid plesiosaur specimens are described from the Early Maastrichtian of Angola. Phylogenetic analyses reconstruct the Angolan taxon as an aristonectine elasmosaurid and the sister taxon of an unnamed form of similar age from New Zealand. Comparisons also indicate a close relationship with an unnamed form previously described from Patagonia. All of these specimens exhibit an ostensibly osteologically immature external morphology, but histological analysis of the Angolan material suggests an adult with paedomorphic traits. By extension, the similarity of the Angolan, New Zealand and Patagonian material indicates that these specimens represent a widespread paedomorphic yet unnamed taxon.

Araújo, R., Polcyn M. J., Schulp A. S., Mateus O., Jacobs L. L., Gonçalves O. A., & Morais M. - L. (2015).  A new elasmosaurid from the early Maastrichtian of Angola and the implications of girdle morphology on swimming style in plesiosaurs. Netherlands Journal of Geosciences. FirstView, 1–12., 1 Abstractaraujo_et_al_2015_a_new_elasmosaurid_from_the_early_maastrichtian_of_angola.pdfWebsite

ABSTRACT We report here a new elasmosaurid from the early Maastrichtian at Bentiaba, southern Angola. Phylogenetic analysis places the new taxon as the sister taxon to Styxosaurus snowii, and that clade as the sister of a clade composed of (Hydrotherosaurus alexandrae (Libonectes morgani + Elasmosaurus platyurus)). The new taxon has a reduced dorsal blade of the scapula, a feature unique amongst elasmosaurids, but convergent with cryptoclidid plesiosaurs, and indicates a longitudinal protraction-retraction limb cycle rowing style with simple pitch rotation at the glenohumeral articulation. Morphometric phylogenetic analysis of the coracoids of 40 eosauropterygian taxa suggests that there was a broad range of swimming styles within the clade.

Araújo, R., Polcyn M. J., Lindgren J., Jacobs L. L., Schulp A. S., Mateus O., Gon?alves A. O., & Morais M. - L. (2014).  New aristonectine elasmosaurid plesiosaur specimens from the Early Maastrichtian of Angola and comments on paedomorphism in plesiosaurs. Geologie en Mijnbouw/Netherlands Journal of Geosciences. 94, 93-108., Number 1 Abstract
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Araújo, R., Polcyn M. J., Schulp A. S., Mateus O., Jacobs L. L., Gon?alves A. O., & Morais M. - L. (2014).  A new elasmosaurid from the early Maastrichtian of Angola and the implications of girdle morphology on swimming style in plesiosaurs. Geologie en Mijnbouw/Netherlands Journal of Geosciences. 94, 109-120., Number 1 Abstract
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Bonaparte, J. F., & Mateus O. (1999).  A new diplodocid, Dinheirosaurus lourinhanensis gen. et sp. nov., from the Late Jurassic beds of Portugal. Revista del Museo Argentino de Ciencias Naturales. 5, 13–29., Number 2 Abstract
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Bonaparte, J. F., & Mateus O. (1999).  A new diplodocid, Dinheirosaurus lourinhanensis gen. et sp. nov., from the Late Jurassic beds of Portugal. Revista del Museo Argentino de Ciencias Naturales. 5, 13–29., Number 2 Abstract
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Bonaparte, J. F., & Mateus O. (1999).  A new diplodocid, Dinheirosaurus lourinhanensis gen. et sp. nov., from the Late Jurassic beds of Portugal. Rev. Mus. Argent. Cienc. Nat. 5, 13-29., Number 2 Abstractbonapartemateus1999_sauropod_dinheirosaurus_portugal.pdfWebsite

Presacral vertebrae of a new Diplodocidae from the Late Jurassic Amoreira-Porto Novo Formation of Lourinhã, Portugal are described and figured. Dinheirosaurus lourinhanensis gen. et sp. is diagnosed by well developed accessory articulations derived from the hyposphene, exposed in lateral view; and by the smaller dorsoventral length of the lower section of the neural arch. It is considered that the organization of the dorsal neural arch of Dinheirosaurus is more derived than in Diplodocus, except in the dorsoventral development of the lower portion of it, which is higher (more derived) in the latter. Possibly the isolated geography of Portugal in the Late Jurassic gave rise to the distinct characters of this new genus.

Brusatte, S. L., Butler R. J., Mateus O., & Steyer J. S. (2015).  A new species of Metoposaurus from the Late Triassic of Portugal and comments on the systematics and biogeography of metoposaurid temnospondyls. Journal of Vertebrate Paleontology. 35, , Number 3 Abstract
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Brusatte, S. L., Butler R. J., Mateus O., & Steyer S. J. (2015).  A new species of Metoposaurus from the Late Triassic of Portugal and comments on the systematics and biogeography of metoposaurid temnospondyls. Journal of Vertebrate PaleontologyJournal of Vertebrate Paleontology. e912988., 2015: Taylor & Francis Abstractbrusatte_et_al2015metoposaurusportugal.pdfWebsite

ABSTRACTMetoposaurids are a group of temnospondyl amphibians that filled crocodile-like predatory niches in fluvial and lacustrine environments during the Late Triassic. Metoposaurids are common in the Upper Triassic sediments of North Africa, Europe, India, and North America, but many questions about their systematics and phylogeny remain unresolved. We here erect Metoposaurus algarvensis, sp. nov., the first Metoposaurus species from the Iberian Peninsula, based on several new specimens from a Late Triassic bonebed in Algarve, southern Portugal. We describe the cranial and pectoral anatomy of M. algarvensis and compare it with other metoposaurids (particularly other specimens of Metoposaurus from Germany and Poland). We provide a revised diagnosis and species-level taxonomy for the genus Metoposaurus, which is currently represented with certainty by three European species (M. diagnosticus, M. krasiejowensis, M. algarvensis). We also identify cranial characters that differentiate these three species, and may have phylogenetic significance. These include features of the braincase and mandible, which indicate that metoposaurid skulls are more variable than previously thought. The new Portuguese bonebed provides further evidence that metoposaurids congregated in fluvial and lacustrine settings across their geographic range and often succumbed to mass death events. We provide an updated paleogeographic map depicting all known metoposaurid occurrences, which shows that these temnospondyls were globally distributed in low latitudes during the Late Triassic and had a similar, but not identical, paleogeographic range as phytosaurs.http://zoobank.org/urn:lsid:zoobank.org:pub:083C80C6-0AB6-49E1-A636-6A8BDBC06A47ABSTRACTMetoposaurids are a group of temnospondyl amphibians that filled crocodile-like predatory niches in fluvial and lacustrine environments during the Late Triassic. Metoposaurids are common in the Upper Triassic sediments of North Africa, Europe, India, and North America, but many questions about their systematics and phylogeny remain unresolved. We here erect Metoposaurus algarvensis, sp. nov., the first Metoposaurus species from the Iberian Peninsula, based on several new specimens from a Late Triassic bonebed in Algarve, southern Portugal. We describe the cranial and pectoral anatomy of M. algarvensis and compare it with other metoposaurids (particularly other specimens of Metoposaurus from Germany and Poland). We provide a revised diagnosis and species-level taxonomy for the genus Metoposaurus, which is currently represented with certainty by three European species (M. diagnosticus, M. krasiejowensis, M. algarvensis). We also identify cranial characters that differentiate these three species, and may have phylogenetic significance. These include features of the braincase and mandible, which indicate that metoposaurid skulls are more variable than previously thought. The new Portuguese bonebed provides further evidence that metoposaurids congregated in fluvial and lacustrine settings across their geographic range and often succumbed to mass death events. We provide an updated paleogeographic map depicting all known metoposaurid occurrences, which shows that these temnospondyls were globally distributed in low latitudes during the Late Triassic and had a similar, but not identical, paleogeographic range as phytosaurs.http://zoobank.org/urn:lsid:zoobank.org:pub:083C80C6-0AB6-49E1-A636-6A8BDBC06A47

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Clemmensen, L. B., Lindström S., Mateus O., Mau M., Milàn J., & Kent D. V. (2021).  A new vertebrate fossil-bearing layer in the Rhætelv Formation (Kap Stewart Group) of central East Greenland: evidence of a Hettangian marine incursion into the continental Jameson Land Basin. Lethaia. n/a, , Number n/a Abstractlet.12449.pdfWebsite

The Kap Stewart Group (Rhaetian-Sinemurian, Triassic–Early Jurassic) of the Jameson Land Basin in central East Greenland has traditionally been regarded as a strictly continental unit with delta and perennial lake sediments. New finds of plesiosaur bone remain in a thin storm deposited sandstone bed in the middle part of the Rhætelv Formation of the Kap Stewart Group, however, indicates a likely period of marine influence. At the study area at the eastern margin of the basin, the Rhætelv Formation is 300-m thick and overlies unconformably the Norian Fleming Fjord Group. The bone-bearing sandstone occurs 190 m above the base of the group and is closely associated with black laminated mudstones; palynological investigation of three samples from these mudstones indicates that they are of a younger Hettangian age. The Hettangian was a relatively short stage (201.3–199.5 Ma) and elsewhere characterized by two episodes of sea-level highstands. Assuming that the marine incursion in the Jameson land Basin evidenced by the plesiosaur fossil remains took place during the youngest of these sea-level highstands, the bone-bearing bed of the Rhætelv Formation can be dated to 200 Ma and thereby gives the first numerical age constraint of this hitherto poorly dated succession.

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Fernandes, A. E., Beccari V., Kellner A. W. A., & Mateus O. (2023).  A new gnathosaurine (Pterosauria, Archaeopterodactyloidea) from the Late Jurassic of Portugal. PeerJ. 11, e16048. Abstractfernandes_et_al_2023_lusognathus_peerj-16048.pdfWebsite

An incomplete, yet remarkably-sized dentated rostrum and associated partial cervical vertebrae of a pterosaur (ML 2554) were recently discovered from the Late Jurassic (Late Kimmeridgian-Early Tithonian) Lourinhã Formation of Praia do Caniçal, of central west Portugal. This specimen exhibits features such as a spatulated anterior expansion of the rostrum, robust comb-like dentition, and pronounced rims of the tooth alveoli, indicating gnathosaurine affinities. Based on its further unique tooth and dentary morphology, a new genus and species, \textit{Lusognathus almadrava} gen. et spec. nov., is proposed, making this the first named pterosaur species found within Portugal. The presence of this taxon adds yet another element to the fluvio-deltaic lagoonal environment that has been suggested as representative of the Lourinhã Formation in the Late Jurassic, further contributing to the diversity and distribution of gnathosaurines worldwide.

Fernandes, A. F., Beccari V., Kellner A. W. A., & Mateus O. (2022).  A new Gnathosaurine (Archaeopterodactyloidea, Pterosauria) from the Late Jurassic of Portugal. XIX Annual Conference of the European Association of Vertebrate Palaeontologists. 56-57. Abstractfernandes_et_al_2022_pterosaur_eavp_2022_abstract_volume.pdf

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Ferrari, L., Rotatori F. M., Camilo B., Moreno-Azanza M., & Mateus O. (2022).  New specimen of dryomorphan (Ornithischia, Iguanodontia) remains from the Upper Jurassic of Portugal. Abstract book of the XIX Annual Conference of the European Association of Vertebrate Palaeontologists, Benevento/Pietraroja, Italy, 27th June-2nd July 2022.. 61-62.: PalaeoVertebrata, Special Volume 1- 2022, 224. Doi: 10.18563/pv.eavp Abstractferrarietal_2022.pdf

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Foth, C., Evers S. W., Pabst B., Mateus O., Flisch A., Patthey M., & Rauhut O. W. M. (2015).  New insights into the lifestyle of \\textitAllosaurus (Dinosauria: Theropoda) based on another specimen with multiple pathologies. PeerJ. 3, e940., 5 AbstractWebsite

Adult large-bodied theropods are often found with numerous pathologies. A large, almost complete, probably adult \\textitAllosaurus specimen from the Howe Stephens Quarry, Morrison Formation (Late Kimmeridgian–Early Tithonian), Wyoming, exhibits multiple pathologies. Pathologic bones include the left dentary, two cervical vertebrae, one cervical and several dorsal ribs, the left scapula, the left humerus, the right ischium, and two left pedal phalanges. These pathologies can be classified as follows: the fifth cervical vertebra, the scapula, several ribs and the ischium are probably traumatic, and a callus on the shaft of the left pedal phalanx II-2 is probably traumatic-infectious. Traumatically fractured elements exposed to frequent movement (e.g., the scapula and the ribs) show a tendency to develop pseudarthroses instead of a callus. The pathologies in the lower jaw and a reduced extensor tubercle of the left pedal phalanx II-2 are most likely traumatic or developmental in origin. The pathologies on the fourth cervical are most likely developmental in origin or idiopathic, that on the left humerus could be traumatic, developmental, infectious or idiopathic, whereas the left pedal phalanx IV-1 is classified as idiopathic. With exception of the ischium, all as traumatic/traumatic-infectious classified pathologic elements show unambiguous evidences of healing, indicating that the respective pathologies did not cause the death of this individual. Alignment of the scapula and rib pathologies from the left side suggests that all may have been caused by a single traumatic event. The ischial fracture may have been fatal. The occurrence of multiple lesions interpreted as traumatic pathologies again underlines that large-bodied theropods experienced frequent injuries during life, indicating an active predatory lifestyle, and their survival perhaps supports a gregarious behavior for \\textitAllosaurus. Alternatively, the frequent survival of traumatic events could be also related to the presence of non-endothermic metabolic rates that allow survival based on sporadic food consumption or scavenging behavior. Signs of pathologies consistent with infections are scarce and locally restricted, indicating a successful prevention of the spread of pathogens, as it is the case in extant reptiles (including birds).

Foth, C., Evers S., Pabst B., Mateus O., Flisch A., Patthey M., & Rauhut O. W. M. (2015).  New insights into the lifestyle of Allosaurus (Dinosauria: Theropoda) based on another specimen with multiple pathologies. PeerJ PrePrints. 3, e824v1., 2015 Abstractfoth_et_al_2015_peerj-preprints-824.pdfWebsite

Adult large-bodied theropods are often found with numerous pathologies. A large, almost complete, probably adult Allosaurus specimen from the Howe Stephens Quarry, Morrison Formation (Late Kimmeridgian–Early Tithonian), Wyoming, shows multiple pathologies. Pathologic bones include the left dentary, two cervical vertebrae, one cervical and several dorsal ribs, the left scapula, the left humerus, right ischium, and two left pedal phalanges. These pathologies can be classified as follows: the fifth cervical vertebra, the scapula, several ribs and the ischium are traumatic, and a callus on the shaft of the left pedal phalanx II-2 is traumatic-infectious. Traumatically fractured elements exposed to frequent movement (e.g. the scapula and the ribs) show a tendency to develop pseudarthroses instead of callus healing. The pathologies in the lower jaw and a reduced flexor tubercle of the left pedal phalanx II-2 are most likely traumatic or developmental in origin. The pathologies on the fourth cervical are most likely developmental in origin or idiopathic, that on the left humerus is infectious or idiopathic, whereas left pedal phalanx IV-1 is classified as idiopathic. With exception of the ischium, all traumatic / traumatic-infectious pathologic elements show unambiguous evidences of healing, indicating that the respective pathologies did not cause the death of this individual. Alignment of the scapula and rib pathologies from the left side suggests that all may have been caused by a single traumatic event. The ischial fracture may have been fatal. The occurrence of multiple traumatic pathologies again underlines that large-bodied theropods experienced frequent injuries during life, indicating an active predatory lifestyle, and their survival perhaps supports a gregarious behavior for Allosaurus. Signs of infections are scarce and locally restricted, indicating a successful prevention of the spread of pathogens, as it is the case in extant reptiles (including birds).

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Graf, J., Jacobs L. L., Polcyn M. J., Mateus O., & Schulp A. S. (2011).  New fossil whales from Angola. 71st Annual Meeting of the Society of Vertebrate Paleontology. 119., Jan: Abstracts of the 71st Annual Meeting of the Society of Vertebrate Paleontologygraf_et_al_mateus_2011_fossil_whales_from_angola_svp11abstracts.pdf
Guillaume, A. R. D., Natário C., Moreno-Azanza M., & Mateus O. (2021).  New occurences of Celtedens (Lissamphibia: Albanerpetontidae) from the Lourinhã Formation. XVIII Conference of the European Association of Vertebrate Palaeontologists. 87. Abstractguillaume_et_al_2021_celtedens.pdf

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Guillaume, A. R., Moreno-Azanza M., & Mateus O. (2019).  New lissamphibian material from the Lourinhã Formation (Late Jurassic, Portugal). Journal of Vertebrate Paleontology, Program and Abstracts. 112. Abstract
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Guillaume, A. R. D., Natário C., Moreno-Azanza M., & Mateus O. (2021).  New material ascribed to Albanerpetontidae (Lissamphibia; Allocaudata) from the Lourinhã Formation: evidence for intraspecific variation. XIX Encontro de Jovens Investigadores em Paleontologia. 50-51. Abstractguillaume_et_al_2021_albanerpetontidae.pdf

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Guillaume, A. R., Moreno-Azanza M., & Mateus O. (2019).  New lissamphibian material from the Lourinhã Formation (Late Jurassic, Portugal). Journal of Vertebrate Paleontology, Program and Abstracts. 112.guillaume_et_al_2019_svp_abstract.pdf
Guillaume, A. R. D., Moreno-Azanza M., & Mateus O. (2022).  New species of Albanerpetontidae (Lissamphibia) from the Lourinhã Formation. XXXVII Jornadas SEP y V Congreso Ibérico de Paleontología. 94., Cuenca: ISBN 13: 978-84-09-44478-6 Abstractguillaume_et_al_2022_albanerpetontidae_jornadas_sep_cuenca_abstract_2022.pdf

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Hendrickx, C., Araújo R., & Mateus O. (2014).  The nonavian theropod quadrate II: systematic usefulness, major trends and cladistic and phylogenetic morphometrics analyses. PeerJ PrePrints. 2, e380v2., 2014 AbstractWebsite

The skull-bone quadrate in nonavian theropods is very diverse morphologically alongside the disparity of the group as a whole. However this disparity has been underestimated for taxonomic purposes. In order to evaluate the phylogenetic potential and investigate the evolutionary transformations of the quadrate, we conducted a Catalano-Goloboff phylogenetic morphometric analysis as well as a cladistic analysis using 98 discrete quadrate related characters. The cladistic analysis provides a fully resolved tree mirroring to some degree the classification of nonavian theropods. The quadrate morphology by its own provides a wealth of data with strong phylogenetic signal and allows inference of major trends in the evolution of this bone. Important synapomorphies include: for Abelisauroidea, a lateral ramus extending to the ectocondyle; for Tetanurae, the absence of the lateral process; for Spinosauridae, a medial curvature of the ventral part of the pterygoid ramus occurring just above the mandibular articulation; for Avetheropoda, an anterior margin of the pterygoid flange formed by a roughly parabolic margin; and for Tyrannosauroidea, a semi-oval pterygoid flange shape in medial view. The Catalano-Goloboff phylogenetic morphometric analysis reveals two main morphotypes of the mandibular articulation of the quadrate linked to function. The first morphotype, characterized by an anteroposteriorly broad mandibular articulation with two ovoid/subcircular condyles roughly subequal in size, is found in Ceratosauria, Tyrannosauroidea and Oviraptorosauria. This morphotype allows a very weak displacement of the mandible laterally. The second morphotype is characterized by an elongate and anteroposteriorly narrow mandibular articulation and a long and parabolic/sigmoid ectocondyle. Present in Megalosauroidea, Carcharodontosauridae and Dromaeosauridae, this morphotype permits the lower jaw rami to be displaced laterally when the mouth opened.

Hendrickx, C., Aráujo R., & Mateus O. (2015).  The non-avian theropod quadrate I: Standardized terminology with an overview of the anatomy and function. PeerJ. 2015, , Number 9 Abstract
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Hendrickx, C., Araújo R., & Mateus O. (2012).  The nonavian theropod quadrate: systematics usefulness, major trends and phylogenetic morphometrics analysis. : Journal of Vertebrate Paleontology, Program and Abstracts, 2012, p.110. ISSN 1937-2809 Abstracthendrickx_araujo__mateus_2012_quadrate_theropods_svp_2012_abstract_book_meeting_abstracts.pdf.pdf

The quadrate in nonavian theropods is incredibly diverse morphologically; however this morphological disparity has been underestimated for taxonomic purposes. The quadrate topological homologies and anatomy, as well as the terminology, among nonavian theropod clades are reviewed. In order to evaluate the phylogenetic potential and investigate the evolutionary transformations of the quadrate, we conducted a Catalano-Goloboff phylogenetic morphometric analysis using 3 morphometric characters, a total of 28 landmarks coded for 23 taxa, as well as a cladistic analysis using 115 discrete quadrate-related characters coded for 43 taxa. The cladistic analysis provides a fully resolved tree mirroring the current classification of nonavian theropods. The quadrate morphology by its own provides a wealth of data with strong phylogenetic signal. Several unambiguous synapomorphies support nonavian theropod relationships and the resulting consensus tree allows inference of major trends in the evolution of this bone. Important synapomorphies include: for Abelisauridae, a lateral ramus extending to the ectocondyle; for Tetanurae, the absence of the lateral process; for Spinosauridae, a medial curvature of the ventral part of the pterygoid ramus occurring just above the mandibular articulation; for Neotetanurae, an anterior margin of the pterygoid flange formed by a roughly parabolic margin; and for Tyrannosauroidea, a semi-oval pterygoid flange shape in medial view. The Catalano-Goloboff phylogenetic morphometric analysis reveals two main morphotypes of the mandibular articulation of the quadrate linked to function. The first morphotype, characterized by an anteroposteriorly broad mandibular articulation with two ovoid/subcircular condyles roughly subequal in size, is found in Ceratosauria, Tyrannosauroidea and Oviraptorosauria. This morphotype allows a very weak displacement of the mandible laterally. The second morphotype is characterized by an elongate and anteroposteriorly narrow mandibular articulation and a long and parabolic/sigmoid ectocondyle. Present in Megalosauroidea, Allosauroidea and Dromaeosauridae, this morphotype permits the lower jaw rami to be displaced laterally when the mouth opened.

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López-Rojas, V., Clemmensen L. B., Milàn J., Wings O., Klein N., & Mateus O. (2023).  A new phytosaur species (Archosauriformes) from the Upper Triassic of Jameson Land, central East Greenland. Journal of Vertebrate PaleontologyJournal of Vertebrate Paleontology. e2181086., 2023: Taylor & Francis Abstracta_new_phytosaur_species_archosauriformes_from_the_upper_triassic_of_jameson_land_central_east_greenland.pdfWebsite

ABSTRACTHerein we describe phytosaurs from thin fluvial overbank sandstones of the Upper Triassic Malmros Klint Formation of the Fleming Fjord Group (central East Greenland). The new sample includes more than 150 disarticulated bones and teeth from small to large specimens belonging to at least four individuals. The fossils mostly consist of teeth and postcranial elements and permit the recognition of a new species of Mystriosuchus, M. alleroq, diagnosed by an L-shaped quadratojugal whose anterior suture trends anterodorsally and a tripartite degree of heterodonty. Humeral diaphyseal histology of one specimen reveals a fairly compact cortex that surrounds a cancellous medullary region followed by a remodeling zone containing scattered secondary osteons. Primary bone tissue is parallel-fibred with a moderate to low vascular density. The cortex is cyclically interrupted by distinct growth marks indicating a seasonal environment. A change in growth rate from moderate to low is documented within the outer cortex, indicating that at least this individual was close to somatic maturity. Mystriosuchus has formerly been known as an exclusively European taxon. The new findings support the European faunal influence in East Greenland during the Late Triassic inferred from other taxa such as temnospondyls and archosaurs. The mid-late Norian age of European Mystriosuchus suggests an additional age constraint for the vertebrate-bearing portion of the Malmros Klint Formation.

López-Rojas, V., Mateus S., Marinheiro J., Mateus O., & Puértolas-Pascual E. (2024).  A new goniopholidid crocodylomorph from the Late Jurassic of Portugal. Palaeontologia Electronica. 27(1), 1-33.: Paleontological Society Abstractgoniopholididae_1316_compressed.pdf

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López-Rojas, V., Puértolas-Pascual E., Marinheiro J., Mateus O., & Mateus S. (2022).  A new goniopholidid skull from the Late Jurassic of Lourinhã, Portugal. XXVII Jornadas SEP e V Congreso Ibérico de Paleontologia. 101., Cuencalopez-rojas_et_al_2022_goniopholis_jornadas_sep_cuenca_abstract_2022.pdf
López-Rojas, V., Mateus O., Milàn J., Wings O., Klein N., & Clemmensen L. B. (2021).  A new phytosaur from the Late Triassic of Jameson Land, Greenland. 3rd Palaeontological Virtual Congress. 207.: ISBN 978-84-09-36657-6 Abstractlopez_rojas_2021_pvc3_greenland.pdf

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Mannion, P. D., Upchurch P., Mateus O., Barnes R. N., & Jones M. E. H. (2012).  New information on the anatomy and systematic position of Dinheirosaurus lourinhanensis (Sauropoda: Diplodocoidea) from the Late Jurassic of Portugal, with a review of European diplodocoids. Journal of Systematic Palaeontology. 10(3), 521–551., Jan Abstractmannion_et_al_2012_new_information_on_the_anatomy_and_systematic_position_of_dinheirosaurus_lourinhanensis_sauropoda_-_diplodocoidea_from_the_late_jurassic_of_portugal_with_a_review_of_european_diplodocoids.pdf

Although diplodocoid sauropods from Africa and the Americas are well known, their European record remains largely neglected. Here we redescribe Dinheirosaurus lourinhanensis from the Late Jurassic of Portugal. The holotype comprises two posterior cervical vertebrae, the dorsal series and a caudal centrum. Redescription demonstrates its validity on the basis of three autapomorphies: (1) posteriorly restricted ventral keel on posterior cervical vertebrae; (2) three small subcircular fossae posterior to the lateral coel on posterior cervical neural spines; (3) accessory lamina linking the hyposphene with base of the posterior centrodiapophyseal lamina in middle-posterior dorsal vertebrae. Phylogenetic analysis places Dinheirosaurus as the sister taxon to Supersaurus, and this clade forms the sister taxon to other diplodocines. However, this position should be treated with caution as Dinheirosaurus displays several plesiomorphic features absent in other diplodocids (including unbifurcated presacral neural spines, and dorsolaterally projecting diapophyses on dorsal vertebrae) and only four additional steps are required to place Dinheirosaurus outside of Flagellicaudata. We identify Amazonsaurus as the basal-most rebbachisaurid and recover Zapalasaurus outside of the South American Limaysaurinae, suggesting the biogeographic history of rebbachisaurids is more complex than previously proposed. Review of the European diplodocoid record reveals evidence for the earliest known diplodocid, as well as additional diplodocid remains from the Late Jurassic of Spain. A Portuguese specimen, previously referred to Dinheirosaurus, displays strong similarities to Apatosaurus from the contemporaneous Morrison Formation of North America, indicating the presence of a second Late Jurassic Portuguese diplodocid taxon. Along with Dinheirosaurus, these Portuguese remains provide further evidence for a Late Jurassic palaeobiogeographic connection between Europe and North America. No dicraeosaurids are currently known from Europe, but rebbachisaurids are present in the Early Cretaceous, with weak evidence for the earliest known representative from the Late Jurassic of Spain; however, more complete material is required to recognize early members of this clade.

Mannion, P. D., Upchurch P., Mateus O., Barnes R. N., & Jonese M. E. H. (2012).  New information on the anatomy and systematic position of Dinheirosaurus lourinhanensis (Sauropoda: Diplodocoidea) from the Late Jurassic of Portugal, with a review of European diplodocoids. Journal of Systematic Palaeontology. 10, 521-551., Number 3 Abstract
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Marinheiro, J., Mateus O., Alaoui A., Amani F., Nami M., & Ribeiro C. (2014).  New Quaternary fossil sites from the Middle Atlas of Morocco. Comunica\\c cões Geológicas. 101, Especial I, 485-488. Abstract

The paleontological richness of Morocco has been scientifically known since at least the early 20th century. The region of the Middle Atlas, more specifically the Boulemane area, has been however only sparsely studied since the 1960s when it provided vertebrate fossils from the Middle Jurassic. In September 2013, a Moroccan-Portuguese expedition to the village of Taghrout, Boulemane, made excavations in a Pleistocene fossil site that once was a small high-altitude sedimentary basin, uncharted in previous geological maps. The excavations yielded bone material from large mammals, the most common findings are elephants ascribed to Elephas, but artiodactyls, turtles, and in-situ Acheulean tools were also collected. This represents a new and important paleontological and archeological site. In addition to the discoveries of Taghrout, the expedition also retrieved Quaternary vertebrate material from a nearby cave and found new Jurassic localities, with arcosaur bones and dinosaur footprints, in El Mers.

Marinheiro, J., Mateus O., Alaoui A., Amani F., Nami M., & Ribeiro C. (2014).  New Quaternary fossil sites from the Middle Atlas of Morocco. Comunicações Geológicas. 101, Especial I, 485-488. Abstractmarinheiro_et_al_2014_new_quaternary_fossil_sites_from_the_middle_atlas_of_morocco.pdf

The paleontological richness of Morocco has been scientifically known since at least the early 20th century. The region of the Middle Atlas, more specifically the Boulemane area, has been however only sparsely studied since the 1960s when it provided vertebrate fossils from the Middle Jurassic. In September 2013, a Moroccan-Portuguese expedition to the village of Taghrout, Boulemane, made excavations in a Pleistocene fossil site that once was a small high-altitude sedimentary basin, uncharted in previous geological maps. The excavations yielded bone material from large mammals, the most common findings are elephants ascribed to Elephas, but artiodactyls, turtles, and in-situ Acheulean tools were also collected. This represents a new and important paleontological and archeological site. In addition to the discoveries of Taghrout, the expedition also retrieved Quaternary vertebrate material from a nearby cave and found new Jurassic localities, with arcosaur bones and dinosaur footprints, in El Mers.

Marinheiro, J., Mateus O., Alaoui A., Amani F., Nami M., & Ribeiro C. (2014).  New Quaternary fossil sites from the Middle Atlas of Morocco. Comunicações Geológicas. 101, 485–488., Number Especial I: Laboratório Nacional de Energia e Geologia Abstract
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Marinheiro, J., Mateus O., Alaoui A., Amani F., Nami M., & Ribeiro C. (2014).  New quaternary fossil sites from the middle atlas of morocco,Novas jazidas fossilíferas do Quaternário do Médio-Atlas de Marrocos. Comunicacoes Geologicas. 101, 485-488. Abstract
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Martino, R., Ríos M., Mateus O., Rook L., & Pandolf L. (2022).  New insights into the hippopotamid (Mammalia, Hippopotamidae) from the Casino Basin (Tuscany, Italy). XIX Annual conference of the European Association of Vertebrate Palaeontologists (19th EAVP)At: Benevento, Italy. 119. Abstract
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Mateus, O., & Antunes M. T. (2003).  A New Dinosaur Tracksite in the Lower Cretaceous of Portugal. Ciências da Terra. 15, 253-262. Abstractmateus__antunes_2003-_early_k_dino_tracks_portugal.pdfWebsite

A new Lower Cretaceous (Aptian-Albian) dinosaur tracksite at the Olhos de Água beach is described. It is the first vertebrate fossil finding ever found in the concerned unit, and yielded 128 tracks in 17 trackways within an area of ca. 80 square metres.
Three tridactyl footprint morphotypes have been recognized: - Type 1 (“Iguanodontipus-like”) - trackways D, F, K, J and P; - Type 2 (large theropod), although larger in size, typically from a Grallator-like theropod footprint, i.e. A, B, G, H and O trackways; - Type 3 (medium size theropod); M is the only track of this type. There are other, poorly preserved, unidentified trackways. The theropod, swinging trackway B was produced by an animal that was limping. The theropod track M starts eastwards but drastically changes westwards, speeding up at the same time; this dinosaur decided to turn around and run in the opposite direction.
This site shows three main trackway directions: to the South, to the East, and westwards. Except for the trackway O, large theropods A, B, G and H walked southwards. Perpendicularly to these, ornithopods, small theropods and unidentified trackmakers walked towards East (5) and West (7). The segregation of trackmakers and directions, with large theropod trackways southwards and other dinosaurs’ west or eastwards, may mean that large theropods patrolled a walkway area to an important resource, most probably water, often frequented by ornithopods and smaller theropods. There is no evidence of social behavior or gregarism: footprints’ overposition shows that the large, southwards walking theropods passed on different occasions. Three trackway sequences can be established by chronologic order.

Mateus, O. (2009).  New specimens of Angolasaurus bocagei and comments on the early radiations of plioplatecarpine mosasaurs. Journal of Vertebrate Paleontology. Journal of Vertebrate Paleontology. 29, 165A., 1, Number 3: Taylor & Francis Abstract

NEW SPECIMENS OF ANGOLASAURUS BOCAGEI AND COMMENTS ON THE EARLY RADIATIONS OF PLIOPLATECARPINE MOSASAURS POLCYN, Michael, SMU, Dallas, TX, USA; JACOBS, Louis, SMU, Dallas, TX, USA; MATEUS, Octávio, Museu da Lourinhã, Lourinhã, Portugal; SCHULP, Anne, Natuurhistorisch Museum Maastricht, Maastricht, Netherlands New, well preserved material of the Turonian mosasaur Angolasaurus bocagei from the Tadi Beds of the Itombe Formation in northern Angola, allows detailed redescription of its morphology and reassessment of its phylogenetic relationships. Angolasaurus had been previously referred to the genus Platecarpus; however, phylogenetic analysis confirms the valid taxonomic status of A. bocagei, and reconstructs that taxon within a clade that also includes the genera Selmasaurus and Ectenosaurus. These forms are united by an elaborated infrastapedial process of the quadrate and a unique ridge-like descending process of the parietal forming the supraoccipital articulation, but also retain a relatively plesiomorphic configuration of the braincase. That clade is united with all other plioplatecarpines by a number of derived characters including the presence of a novel basicranial circulation pattern. In Africa, North and South America, early plioplatecarpines are known by the Middle Turonian and Angolasaurus and closely related forms appear by the Upper Turonian. Selmasaurus and Ectenosaurus are a rare faunal component of the Santonian and Campanian of North America. Platecarpus planifrons appears in the Coniacian of North America and represents the plesiomorphic condition of the clade containing the remaining species of Platecarpus and Plioplatecarpus, that appears in the Santonian and persist until the end of the Cretaceous, reaching global distribution. The temporal and geographic distribution of these radiations suggest influence of paleogeography and eustatic sea levels.

Mateus, O., & Antunes M. T. (2002).  Novos vestígios de dinossauros saurópodes do Jurássico Superior da Lourinhã (Portugal).. Congresso Ibérico de Herpetologia. , Évora, Portugal Abstract

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Mateus, O., Milàn J., Romano M., & Whyte M. A. (2011).  New finds of stegosaur tracks from the Upper Jurassic Lourinhã formation, Portugal. Acta Palaeontologica Polonica. 56, 651-658., Number 3 Abstract
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Mateus, O., & Antunes M. T. (2002).  Novos vestígios de dinossauros saurópodes do Jurássico Superior da Lourinhã (Portugal). Congresso Ibérico de Herpetologia. , Évora Abstract
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Mateus, O., Araújo R., Natário C., & Castanhinha R. (2011).  A new specimen of the theropod dinosaur Baryonyx from the early Cretaceous of Portugal and taxonomic validity of Suchosaurus. Zootaxa. 2827, 54–68., Jan Abstractmateus_et_al_2011_a_new_specimen_of_the_theropod_dinosaur_baryonyx_from_the_early_cretaceous_of_portugal_and_taxonomic_validity_of_suchosaurus.pdf

Although the Late Jurassic of Portugal has provided abundant dinosaur fossils, material from the Early Cretaceous is scarce. This paper reports new cranial and postcranial material of the theropod dinosaur Baryonyx walkeri found in the Barremian (Papo Seco Formation) of Portugal. This specimen, found at Praia das Aguncheiras, Cabo Espichel, consists of a partial dentary, isolated teeth, pedal ungual, two calcanea, presacral and caudal vertebrae, fragmentary pubis, scapula, and rib fragments. It represents the most complete spinosaurid yet discovered in the Iberian Peninsula and the most complete dinosaur from the Early Cretaceous of Portugal. This specimen is confidently identified as a member of Baryonychinae due to the presence of conical teeth with flutes and denticles in a dentary rosette. The specimen ML1190 shares the following characteristics with Baryonyx walkeri: enamel surface with small (nearly vertical) wrinkles, variable denticle size along the carinae, 6–7 denticles per mm, wrinkles forming a 45 degree angle near the carinae, and tooth root longer than crown. In addition, dubious taxa based on teeth morphology such as Suchosaurus cultridens (Owen, 1840–1845), and Suchosaurus girardi (Sauvage 1897–98; Antunes & Mateus 2003) are discussed, based on comparisons with well-known material such as Baryonyx walkeri Charig & Milner, 1986. Suchosaurus cultridens and S. girardi are considered as nomina dubia due to the lack of diagnostic apomorphies, but both specimens are referred to Baryonychinae incertae sedis.

Mateus, O. (2011).  A new metoposaurid (temnospondyl) bonebed from the Late Triassic of Portugal. 31, , 1 Abstract

The end-Triassic extinction event (ETE), considered one of the ‘Big Five’ mass extinctions, marks a dividing line between early Mesozoic vertebrate assemblages, typically including abundant temnospondyls, basal synapsids and basal archosaurs, and ‘typical’ Mesozoic faunas dominated by dinosaurs, pterosaurs, crocodylomorphs, turtles and mammaliaforms. Recent geochemical work has provided strong evidence that the ETE is synchronous with, and likely caused by, the emplacement of the Central Atlantic magmatic province (CAMP). However, stratigraphic sections containing both terrestrial vertebrates and CAMP basalts are scarce, complicating attempts to examine terrestrial faunal changes during this extinction event. The Triassic–Jurassic Algarve Basin, southern Portugal, is an extensional rift basin to-marginal marine red beds (the ‘Grés de Silves’ Group) interbedded with CAMP basalts.... bonebed from the interval ‘AB1’ of the Grés de Silves. Preliminary excavations yielded at least nine well-preserved temnospondyl individuals represented by partial to nearly complete skulls and disarticulated postcranial elements of juvenile to adult ages. Nearly all material appears to represent a single species of metoposaurid referable to the genus Metoposaurus, well known from the late Carnian–early Norian of Germany and Poland. A number of char- acters of the occiput and mandible suggest that the Algarve material may represent a new species. This new material provides new data on the diversity and paleogeographical distri- bution of the metoposaurids, a highly autapomorphic and peculiar group composed of large aquatic carnivores with a unique elongated but brevirostral skull. This taxon also provides [...] Horizon may be within or close to the late Carnian–early Norian. Additional bone-bearing horizons within the ‘Grés de Silves’ provide a rare opportunity to examine terrestrial faunal change in the lead-up to the ETE.

Mateus, O., Natario C., Araujo R., & Castanhinha R. (2008).  A new specimen of spinosaurid dinosaur aff. Baryonyx from the Early Cretaceous of Portugal. (Universidade de, Coimbra, Ed.).Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 51., Coimbra Abstract
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Mateus, O., & Estraviz-López D. (2022).  A new theropod dinosaur from the early cretaceous (Barremian) of Cabo Espichel, Portugal: Implications for spinosaurid evolution. PLOS ONE. 17, 1-48., 02, Number 2: Public Library of Science Abstractjournal.pone_.0262614.pdfWebsite

Spinosaurids are some of the most enigmatic Mesozoic theropod dinosaurs due to their unique adaptations to aquatic environments and their relative scarcity. Their taxonomy has proven to be especially problematic. Recent discoveries from Western Europe in general, specifically Iberia, provide some of the best specimens for the understanding of their phylogeny, leading to the description of the spinosaurid Vallibonavenatrix cani and the recognition of the Iberian dinosaur Camarillasaurus cirugedae as one of them. Portuguese associated spinosaurid remains (ML1190) from the Papo Seco Formation (early Barremian) were previously assigned to Baryonyx walkeri but new material recovered in 2020 along with new phylogenetic analyses suggests a different phylogenetic placement, making their revision necessary. Here we show that these remains are not attributable to Baryonyx walkeri, but to a new genus and species, Iberospinus natarioi, gen. et sp. nov. The new taxon is characterized by the presence of a single Meckelian foramen in the Meckelian sulcus, a straight profile of the ventral surface of the dentary and a distal thickening of the acromion process of the pubis between other characters. Iberospinus natarioi is recovered as a sister taxon of the clade formed by Baryonyx and Suchomimus, and outside Spinosaurinae when Vallibonaventrix cani is excluded from the analysis. The description of this taxon reinforces Iberia as a hotspot for spinosaur biodiversity, with several endemic taxa for the region. As expected for the clade, the dentary displays a highly vascularized neurovascular network. The morphometric analysis of parts of the skeleton (pedal phalanx and caudal vertebrae, among others) shows an intermediate condition between basal tetanurans and spinosaurines.