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Mateus, O. (2014).  Eggs and eggshells of crocodylomorpha from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology. Program and Abstracts, 2014, 218., Number NA: Taylor & Francis Abstract
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Mateus, O., Jacobs L. L., Polcyn M. J., Schulp A. S., Neto A. B., & Antunes M. T. (2008).  Dinosaur and turtles from the Turonian of Iembe, Angola. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 156–156., Neuquén, Argentina Abstract
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Mateus, O. (2010).  Paleontological collections of the Museum of Lourinhã (Portugal). (Brandao, JM, Callapez, PM, O. Mateus, Castro, P, Ed.).Colecções e museus de Geologia: missão e gestão. 121-126., Jan: Ed. Universidade de Coimbra e Centro de Estudos e Filosofia da História da Ciência Coimbra Abstractmateus_2010_paleontological_collections_of_the_museum_of_lourinha__geocoleccoes_omateus.pdf

Abstract: The paleontological collections of the Museum of Lourinhã, in Portugal, has a rich paleontological collection, particularly of Late Jurassic dinosaurs of the Lourinhã Formation (Kimmeridgian-Tithonian). Most salient highlights comprehend the following dinosaur holotype specimens: stegosaur Miragaia longicollum, theropod Lourinhanosaurus antunesi, sauropod Dinheirosaurus lourinhanensis, ornithopod Draconyx loureiroi, theropod Allosaurus europaeus, and, a mammal, Kuehneodon hahni. Other dinosaur specimens are referred including the nest and eggs and embryos of Lourinhanosaurus. Portugal is very productive in Late Jurassic vertebrates, being the seventh country bearing more dinosaur taxa.

Mateus, O. (2016).  Late Jurassic of Morrison Formation and Portugal tetrapods compared: a model to explain faunal exchange and similarity. Annual Meeting of the Society of Vertebrate Paleontology. 185., Salt Late City: Journal of Vertebrate Paleontology, Program and Abstracts, 2016 Abstract

The precursor of the North Atlantic existed between the North American and Iberian blocks from the earliest Jurassic Hettangian and has been ever expanding since. By the Kimmeridgian and Tithonian, when much of the Morrison Fm rocks were deposited, the proto-Atlantic was more than 300 km wide at 27° paleolatitude between North America and Iberia. Macrovertebrate paleontology reveals a unique story to the isolation of Iberia and instead suggest a paleogeographic land connection between North American and Iberia. Torvosaurus, Allosaurus, Ceratosaurus, Stegosaurus, Supersaurus and others have a distribution restricted to Morrison Formation in North America and Lourinhã Formation in Portugal. A novel paleogeographic model is here suggested: (1) around the Middle–Late Jurassic transition there is a major palaeoceanographic and palaeoclimatic reorganization, coincidental to a major eustatic sea-level drop and uplift associated with the Callovian– Oxfordian Atlantic Regressive Event; (2) creating an ephemeral land bridge presenting a temporary opportunity for terrestrial gateways likely across the Flemish Cap and Galician Bank land masses, allowing large dinosaurian taxa to cross the northern proto-Atlantic in both directions; (3) finally, a Callovian–Oxfordian faunal exchange around the 163 Ma, through latest Kimmeridgian at 152 Ma (the age of equivalent genera in both Morrison and Portugal), is was an interval that allowed speciation, but retaining generic similarity of vertebrates. This model is consistent with the chronology and taxonomy required for speciation of the Iberian and American forms, exemplified by the coeval sister-taxa pairs Torvosaurus tanneri and T. gurneyi, Allosaurus fragilis and A. europaeus, or Supersaurus vivianae and S. lourinhanensis. While some of the smaller animals in the fauna show Morrison/Portugal affinities, most from Iberia have European or even Asian affinities. The larger-bodied fauna are more closely related to Morrison than to mainland Europe (except for dacentrurine stegosaurs). The body size differences and affinities of taxa across paleogeography is comparable to what is observed today across the Wallace Line. Migration may have also occurred in both directions. The closest relative of Torvosaurus is likely the European Bathonian Megalosaurus, thus the presence of the genus in North America represents a European migration. On other hand, Allosaurus and Supersaurus origins are consistent with a North American origin, representing an westto-east migration.

Mateus, O. (2021).  Overview of vertebrate fossils from Portugal, focus on Mesozoic dinosaurs. 3rd Palaeontological Virtual Congress. 15.: ISBN 978-84-09-36657-6 Abstractmateus_2021_pvc3_key_note_speaker.pdf

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Mateus, O. (2013).  Cabinda revisited: age and environment of new Cenozoic vertebrate fossils from northern Angola. 45, , 1 Abstract

In the early 20thcentury, Belgian naturalists reported Paleocene and Eocene sharks, the bothremydid pleurodiran turtleTaphosphrys(formerlyBantuchelys), and a neosuchian and the dyrosaurid crocodyliformCongosaurusfrom coastal outcrops near Landana in the northern province of Cabinda, Angola. In 1935, rare and fragmentary mammals were reported from strata at Malembo Point, south of Landana, and originally considered to be Miocene in age. Subsequent re-identification of the mammal fossils suggested that the Malembo deposits are Lower Oligocene based on the presence of an arsinoithere and hyracoids. An anthropoid canine, originally identified as a carnivore incisor, was also recognized. In four recent expeditions to Cabinda, Projecto PaleoAngola has traced and measured the stratigraphy between Landana and Malembo Point and sampled these deposits for pollen, stable isotopes, detrital zircons, paleomagnetic stratigraphy, and for U/Pb dating of bones, teeth, and coprolites. Although the faunas from all Cabinda localities are dominated by sharks and rays, new discoveries from Landana include a complete cheloniid cryptodire turtle skull, a small snake vertebra, and a bird bone. Discoveries from the Malembo level include a narrow-snouted crocodyliform similar toCongosaurusandEuthecodon, an arsinoithere anterior tooth, an upper molar similar to that of the ptolemaiidanKelba, an unidentified mammalian incisor, and a large primate-like premolar. Recent biostratigraphic advances in East Africa and the new fossil discoveries in Cabinda suggest similarity to late Oligocene faunas in Ethiopia and Kenya, although the large primate-like premolar is unique. To this day, the fossil localities of Malembo provide the only coastal, low latitude, low elevation record of West African Cenozoic terrestrial mammals. https://gsa.confex.com/gsa/2013AM/webprogram/Paper232115.html

Mateus, O., Morais M., Schulp A., Jacobs L., & Polcyn M. (2006).  The Cretaceous of Angola. JOURNAL OF VERTEBRATE PALEONTOLOGY. 26, 96A-97A., Jan Abstractmateus_et_al_2006_svp_abstracts_cretaceous_fo_angola.pdf

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Mateus, O., Mannion P. D., & Upchurch P. (2014).  Zby atlanticus, a new turiasaurian sauropod (Dinosauria, Eusauropoda) from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology. 34(3), 618-634. Abstractmateus_et_al_2014_zby_atlanticus.pdfWebsite

Here we describe a new partial sauropod skeleton from the late Kimmeridgian (Late Jurassic) of the Lourinhã Formation, central west Portugal. The closely associated specimen comprises a complete tooth (with root), a fragment of cervical neural arch, an anterior chevron, and an almost complete right pectoral girdle and forelimb. The new sauropod, Zby atlanticus, n. gen. et sp., can be diagnosed on the basis of four autapomorphies, including a prominent posteriorly projecting ridge on the humerus at the level of the deltopectoral crest. Nearly all anatomical features indicate that Zby is a non-neosauropod eusauropod. On the basis of several characters, including tooth morphology, extreme anteroposterior compression of the proximal end of the radius, and strong beveling of the lateral half of the distal end of the radius, Zby appears to be closely related to Turiasaurus riodevensis from approximately contemporaneous deposits in eastern Spain. However, these two genera can be distinguished from each other by a number of features pertaining to the forelimb. Whereas previously described Late Jurassic Portuguese sauropods show close relationships with taxa from the contemporaneous Morrison Formation of North America, it appears that turiasaurians were restricted to Europe. All adult sauropods recovered in the Late Jurassic of Portugal thus far are very large individuals: it is possible that the apparent absence of small- or medium-sized adult sauropods might be related to the occupation of lower-browsing niches by non-sauropods such as the long-necked stegosaur Miragaia longicollum.

Mateus, O., Milàn J., Romano M., & Whyte M. A. (2011).  New finds of stegosaur tracks from the Upper Jurassic Lourinhã formation, Portugal. Acta Palaeontologica Polonica. 56, 651-658., Number 3 Abstract
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Mateus, O., & Antunes M. T. (2001).  Draconyx loureiroi, a new camptosauridae (Dinosauria, Ornithopoda) from the Late Jurassic of Lourinhã, Portugal. Annales de Paléontologie. 87, 61–73., jan, Number 1: Elsevier {BV} AbstractWebsite
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Mateus, O., & Antunes M. T. (2002).  Novos vestígios de dinossauros saurópodes do Jurássico Superior da Lourinhã (Portugal).. Congresso Ibérico de Herpetologia. , Évora, Portugal Abstract

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Mateus, O. (2009).  New specimens of Angolasaurus bocagei and comments on the early radiations of plioplatecarpine mosasaurs. Journal of Vertebrate Paleontology. Journal of Vertebrate Paleontology. 29, 165A., 1, Number 3: Taylor & Francis Abstract

NEW SPECIMENS OF ANGOLASAURUS BOCAGEI AND COMMENTS ON THE EARLY RADIATIONS OF PLIOPLATECARPINE MOSASAURS POLCYN, Michael, SMU, Dallas, TX, USA; JACOBS, Louis, SMU, Dallas, TX, USA; MATEUS, Octávio, Museu da Lourinhã, Lourinhã, Portugal; SCHULP, Anne, Natuurhistorisch Museum Maastricht, Maastricht, Netherlands New, well preserved material of the Turonian mosasaur Angolasaurus bocagei from the Tadi Beds of the Itombe Formation in northern Angola, allows detailed redescription of its morphology and reassessment of its phylogenetic relationships. Angolasaurus had been previously referred to the genus Platecarpus; however, phylogenetic analysis confirms the valid taxonomic status of A. bocagei, and reconstructs that taxon within a clade that also includes the genera Selmasaurus and Ectenosaurus. These forms are united by an elaborated infrastapedial process of the quadrate and a unique ridge-like descending process of the parietal forming the supraoccipital articulation, but also retain a relatively plesiomorphic configuration of the braincase. That clade is united with all other plioplatecarpines by a number of derived characters including the presence of a novel basicranial circulation pattern. In Africa, North and South America, early plioplatecarpines are known by the Middle Turonian and Angolasaurus and closely related forms appear by the Upper Turonian. Selmasaurus and Ectenosaurus are a rare faunal component of the Santonian and Campanian of North America. Platecarpus planifrons appears in the Coniacian of North America and represents the plesiomorphic condition of the clade containing the remaining species of Platecarpus and Plioplatecarpus, that appears in the Santonian and persist until the end of the Cretaceous, reaching global distribution. The temporal and geographic distribution of these radiations suggest influence of paleogeography and eustatic sea levels.

Mateus, O. (1999).  Monofilia dos dinossauros e Origem das Aves: Serão as aves dinossauros?. Actas do II Congresso de Ornitologia. 184–185., Lisboa Abstract
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Mateus, O. (1998).  Dinossauros Portugueses. Caderno de resumos do I Congresso de Estudantes de Biologia. 13., Évora Abstractmateus_1998_dinossauros_portugueses_i_congresso_estudantes_biologia.pdf.pdf

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Mateus, O., & Azinheira C. (1996).  Passeriformes nidificantes da Ribeira da Viscossa (Évora). (Spea, Ed.).Livro de resumos do I Congresso de Ornitologia. Abstract
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Mateus, O., Maidment S. C. R., & Christiansen N. A. (2008).  A new specimen aff. Dacentrurus armatus (Dinosauria: Stegosauridae) from the Late Jurassic of Portugal. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontologia de Vertebrados. 157., Neuquén, Argentina Abstractmateus_et_al_2008_a_new_specimen_aff._dacentrurus_armatus_dinosauria_stegosauridae_from_the_late_jurassic_of_portugal.pdf

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Mateus, O., Callapez P. M., & Puértolas-Pascual E. (2017).  The oldest Crocodylia? a new eusuchian from the Late Cretaceous (Cenomanian) of Portugal. Journal of Vertebrate Paleontology, Program and Abstracts. 2017, 160. Abstract
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Mateus, O., Dyke G., Motchurova-Dekova N., Ivanov P., & Kamenov G. D. (2008).  The Bulgarian dinosaur: did it exist? European late Cretaceous ornithomimosaurs. 56th Symposium of Vertebrate Palaeontology and Comparative Anatomy. 47., Dublin Abstract
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Mateus, O. (2016).  Exemplos bizarros de evolução em dinossauros e alguns casos portugueses. Do Big Bang ao Homem. 81-95., Porto: U.Porto Edi{\c c}ões Abstract
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Mateus, O. (2012).  Age and paleoecology of mosasaurs and plesiosaurs from the Late Cretaceous South Atlantic margin at Bentiaba, Angola. Journal of Vertebrate Paleontology, Program and Abstracts, 2012. 180.. 180–181., 1 Abstract
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Mateus, O., Mateus O., Jacobs L. L., Schulp A. S., Polcyn M. J., Tavares T. S., Neto A. B., Morais M. L., & Antunes M. T. (2011).  Angolatitan adamastor, a new sauropod dinosaur and the first record from Angola. Anais da Academia Brasileira de Ciências. Abstract
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Mateus, O., & Antunes M. T. (2002).  Novos vestígios de dinossauros saurópodes do Jurássico Superior da Lourinhã (Portugal). Congresso Ibérico de Herpetologia. , Évora Abstract
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Mateus, O. (2009).  Colecções paleontológicas do Museu da Lourinhã (Portugal) / Paleontological collections of the Museum of Lourinhã (Portugal). (Unknown Unknown, Ed.).Journal of Paleontological Techniques. 18–19.., 1 Abstract
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Mateus, O. (1998).  Serão as aves dinossauros?. CiênciaJ. 6, 5. Abstract
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Mateus, I., Mateus H., Antunes M. T., Mateus O., Taquet P., Ribeiro V., & Manuppella G. (1997).  Couvée, oeufs et embryons d'un dinosaure théropode du Jurassique supérieur de Lourinhã (Portugal). Comptes Rendus de l'Académie des Sciences-Series IIA-Earth and Planetary Science. 325, 71–78., Number 1 Abstract
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Mateus, O. (2008).  Checklist for Late Jurassic reptiles and amphibians from Portugal. Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 55., Coimbra Abstractmateus_2008_lista_de_repteis_e_anfibios_do_jurassico_superior_de_portugal__list_congressoherpetolog.pdf

The richness of Late Jurassic vertebrates in Portugal is known since the 19th century by Paul Choffat, Henri Sauvage and other. The Kimmeridgian Guimarota fauna assemblage is the best known, followed by the fauna of Lourinhã formation. Here is presented an attempt to provide a checklist of the reptiles and amphibians of the Late Jurassic. Amphibia: Lissamphibia (Celtedens, cf. Marmorerpeton, Discoglossidae indet.). Chelonia: Eucryptodira (Pleurosternidae indet., Platychelyidae indet., Plesiochelys cf. etalloni, Plesiochelys choffati, Anosteirinae indet.). Squamata: Scincomorpha (Becklesius hoffstetteri; Paramacellodus sp., Saurillodon proraformis, S. henkeli, S. cf. obtusus). Squamata: Anguimorpha (Dorsetisaurus pollicidens, Parviraptor estesi). Crown Lepidosauromorpha (Marmoretta sp.). Choristodera: Cteniogenidae (Ctenogenys reedi). Sauropterygia: Plesiosauria: Cryptoclidoidea: Cryptoclididae indet. Crocodylomorpha (Lisboasaurus estesi, L. mitrocostatus). Crocodyliformes: Neosuchia (Machimosaurus hugii, Goniopholis cf. simus, Goniopholis baryglyphaeus, cf. Bernissartia, Atoposauridae, Theriosuchus guimarotae, cf. Alligatorium, Metriorhynchus sp.). Pterosauria (Rhamphorhynchus sp., Pterodactylus sp.). Dinosauria: Theropoda (Ceratosaurus sp. , Torvosaurus sp., Lourinhanosaurus antunesi, Allosaurus europaeus, Cf. Compsognathus sp., cf. Richardoestesia sp., Dromaeosaurinae indeter., Velociraptorinae indeter., cf. Archaeopteryx sp., aff. Paronychodon). Dinosauria: Sauropoda: Eusauropoda (Dinheirosaurus lourinhanensis, Lourinhasaurus alenquerensis, Lusotitan atalaiensis, Apatosaurus sp.). Dinosauria: Ornithischia: Thyreophora (Dacentrurus armatus, Stegosaurus sp., Dracopelta zbyszewskii). Dinosauria: Ornithischia: Ornithopoda (Phyllodon henkeli, Dryosaurus sp., Hypsilophodon sp., Alocodon kuehnei, Trimucrodon cuneatus, Draconyx loureiroi).

Mateus, O. (2008).  Checklist for Late Jurassic reptiles and amphibians from Portugal. Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 55., Coimbra Abstract
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Mateus, O. (2008).  Fósseis de transição, elos perdidos, fósseis vivos e espécies estáveis. (Levy, et al, Ed.).Evolução: História e Argumentos. 77-96., Lisboa: Esfera do Caosmateus_2008_evolucao_fosseis_de_transicao.pdf
Mateus, O., Mannion P. D., & Upchurch P. (2014).  Zby atlanticus, a new turiasaurian sauropod (Dinosauria, Eusauropoda) from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology. 34, 618-634., Number 3 Abstract
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Mateus, O., & Milan J. (2010).  A diverse Upper Jurassic dinosaur ichnofauna from central-west Portugal. Lethaia. 43, 245–257., Jan Abstractmateus__milan_2010_-_diverse_l_j_ichnofauna_from_lourinha_fm_portugal.pdfWebsite

A newly discovered dinosaur track-assemblage from the Upper Jurassic Lourinha˜ Formation (Lusitanian Basin, central-west Portugal), comprises medium- to large-sized sauropod tracks with well-preserved impressions of soft tissue anatomy, stegosaur tracks and tracks from medium- to large-sized theropods. The 400-m-thick Lourinha˜ Formation consists of mostly aluvial sediments, deposited during the early rifting of the Atlantic Ocean in the Kimmeridgian and Tithonian. The stratigraphic succession shows several shifts between flood-plain mud and fluvial sands that favour preservation and fossilization of tracks. The studied track-assemblage is found preserved as natural casts on the underside of a thin bivalve-rich carbonate bed near the Tithonian–Kimmeridgian boundary. The diversity of the tracks from the new track assemblage is compared with similar faunas from the Upper Jurassic of Asturias, Spain and the Middle Jurassic Yorkshire Coast of England. The Portuguese record of Upper Jurassic dinosaur body fossils show close similarity to the track fauna from the Lourinha˜ Formation.

Mateus, O. (2011).  Plesiosaur structural extreme from the Maastrichtian of Angola. Abstracts of the 71st Annual Meeting of the Society of Vertebrate Paleontology. 63., 1 Abstract
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Mateus, O., & Jacinto J. J. (2002).  Contribuição para o estudo de Hemidactylus turcicus (Reptilia, Gekkonidae): ritmos de actividade e microhabitat em Évora, Portugal. VII Congresso Luso-Espanhol de Herpetologia. 136–136., Évora Abstract
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Mateus, O., Clemmensen L., Klein N., Wings O., Frobøse N., Milàn J., Adolfssen J., & Estrup E. (2014).  The Late Triassic of Jameson Land revisited: new vertebrate findings and the first phytosaur from Greenland. Journal of Vertebrate Paleontology. Program and Abstracts, 2014, 182.mateus_et_al2014-_jameson_land_revisited_-_svp_2014.pdf
Mateus, O. (2014).  Geological and paleoecological setting of a marine vertebrate bonebed from the Lower Maastrichtian at Bentiaba, Angola. Proceedings of the Secondary Adaptation of Tetrapods to Aquatic Life. NA., 1 Abstract

A single, geographically and temporally restricted horizon at Bentiaba, Angola (14.3° S), preserves a concentration of skeletons and isolated elements representing sharks, rays, bony fish, at least three species of turtles, two species of plesiosaurs, at least five species of mosasaurs, and rare volant and terrestrial forms. The concentration, referred to as the Bench 19 Fauna, formed on a narrow continental shelf at paleolatitude 24°S as predicted by paleomagnetic data and confirmed by plate motion models. The shelf evolved as a transform passive margin along faults associated with the opening of the South Atlantic. Latitude 24°S falls today along the coast of northern Namibia, an area of intense upwelling and hyperarid coastal desert. The Namibe Basin in southern Angola is separated from the Walvis Basin of Namibia by the Walvis Ridge, and the continental shelf in northern Namibia is eight times the width of that at Bentiaba. However, the sediment entombing the fossils at Bentiaba is an immature feldspathic sand, shown by detrital zircon ages to be derived from nearby exposed granitic shield rocks, suggesting similar climatic and drainage conditions between the two regions. Temporal control of the Bentiaba section is provided by magnetostratigraphy and stable carbon isotope chemostratigraphy anchored by an Ar40/Ar39radiometric date on basalt. The age of Bench 19 is constrained to chron C32n.1n and thus falls between 71.4 and 71.64 Ma. Massive bedding without hummocky cross-bedding or other sedimentary structures indicates deposition in shallow water below wave base. δ18O analysis of bivalve shells indicates a water temperature of 18° C immediately below Bench 19. Nearest neighbor distance peaks at 5 m (n=19

Mateus, O., Taquet P., Antunes M. T., Mateus H., & Ribeiro V. (1998).  Theropod dinosaur nest from Lourinhã, Portugal. Journal of Vertebrate Paleontology. 18, 61., Number (Suppl. 3) Abstract
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Mateus, O., Maidment S. C. R., & Christiansen N. A. (2008).  A new specimen aff. Dacentrurus armatus (Dinosauria: Stegosauridae) from the Late Jurassic of Portugal. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 157–157., Neuquén, Argentina Abstract
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Mateus, O. (2016).  Late Jurassic of Morrison Formation and Portugal tetrapods compared: a model to explain faunal exchange and similarity. Annual Meeting of the Society of Vertebrate Paleontology. 185., Salt Late City: Journal of Vertebrate Paleontology, Program and Abstracts, 2016 Abstractmateus_2016_late_jurassic_morrison_svp_abstract.pdf

The precursor of the North Atlantic existed between the North American and Iberian blocks from the earliest Jurassic Hettangian and has been ever expanding since. By the Kimmeridgian and Tithonian, when much of the Morrison Fm rocks were deposited, the proto-Atlantic was more than 300 km wide at 27° paleolatitude between North America and Iberia. Macrovertebrate paleontology reveals a unique story to the isolation of Iberia and instead suggest a paleogeographic land connection between North American and Iberia. Torvosaurus, Allosaurus, Ceratosaurus, Stegosaurus, Supersaurus and others have a distribution restricted to Morrison Formation in North America and Lourinhã Formation in Portugal. A novel paleogeographic model is here suggested: (1) around the Middle–Late Jurassic transition there is a major palaeoceanographic and palaeoclimatic reorganization, coincidental to a major eustatic sea-level drop and uplift associated with the Callovian– Oxfordian Atlantic Regressive Event; (2) creating an ephemeral land bridge presenting a temporary opportunity for terrestrial gateways likely across the Flemish Cap and Galician Bank land masses, allowing large dinosaurian taxa to cross the northern proto-Atlantic in both directions; (3) finally, a Callovian–Oxfordian faunal exchange around the 163 Ma, through latest Kimmeridgian at 152 Ma (the age of equivalent genera in both Morrison and Portugal), is was an interval that allowed speciation, but retaining generic similarity of vertebrates. This model is consistent with the chronology and taxonomy required for speciation of the Iberian and American forms, exemplified by the coeval sister-taxa pairs Torvosaurus tanneri and T. gurneyi, Allosaurus fragilis and A. europaeus, or Supersaurus vivianae and S. lourinhanensis. While some of the smaller animals in the fauna show Morrison/Portugal affinities, most from Iberia have European or even Asian affinities. The larger-bodied fauna are more closely related to Morrison than to mainland Europe (except for dacentrurine stegosaurs). The body size differences and affinities of taxa across paleogeography is comparable to what is observed today across the Wallace Line. Migration may have also occurred in both directions. The closest relative of Torvosaurus is likely the European Bathonian Megalosaurus, thus the presence of the genus in North America represents a European migration. On other hand, Allosaurus and Supersaurus origins are consistent with a North American origin, representing an westto-east migration.

Mateus, O., & Castanhinha R. (2008).  PaleoAngola- Predadores de um oceano primitivo. National Geographic Portugal. 8, 26-33., Number 91 Abstract
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Mateus, O. (2007).  Notes and review of the ornithischian dinosaurs of Portugal. Journal of Vertebrate Paleontology. 27(suppl. to 3), 114. Abstract
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Mateus, O., Araújo R., Natário C., & Castanhinha R. (2011).  A new specimen of the theropod dinosaur Baryonyx from the early Cretaceous of Portugal and taxonomic validity of Suchosaurus. Zootaxa. 2827, 54–68., Jan Abstractmateus_et_al_2011_a_new_specimen_of_the_theropod_dinosaur_baryonyx_from_the_early_cretaceous_of_portugal_and_taxonomic_validity_of_suchosaurus.pdf

Although the Late Jurassic of Portugal has provided abundant dinosaur fossils, material from the Early Cretaceous is scarce. This paper reports new cranial and postcranial material of the theropod dinosaur Baryonyx walkeri found in the Barremian (Papo Seco Formation) of Portugal. This specimen, found at Praia das Aguncheiras, Cabo Espichel, consists of a partial dentary, isolated teeth, pedal ungual, two calcanea, presacral and caudal vertebrae, fragmentary pubis, scapula, and rib fragments. It represents the most complete spinosaurid yet discovered in the Iberian Peninsula and the most complete dinosaur from the Early Cretaceous of Portugal. This specimen is confidently identified as a member of Baryonychinae due to the presence of conical teeth with flutes and denticles in a dentary rosette. The specimen ML1190 shares the following characteristics with Baryonyx walkeri: enamel surface with small (nearly vertical) wrinkles, variable denticle size along the carinae, 6–7 denticles per mm, wrinkles forming a 45 degree angle near the carinae, and tooth root longer than crown. In addition, dubious taxa based on teeth morphology such as Suchosaurus cultridens (Owen, 1840–1845), and Suchosaurus girardi (Sauvage 1897–98; Antunes & Mateus 2003) are discussed, based on comparisons with well-known material such as Baryonyx walkeri Charig & Milner, 1986. Suchosaurus cultridens and S. girardi are considered as nomina dubia due to the lack of diagnostic apomorphies, but both specimens are referred to Baryonychinae incertae sedis.

Mateus, O., & Milàn J. (2011).  New dinosaur and pterosaur tracksites from the Late Jurassic of Portugal. Dinosaur Tracks 2011 An International Symposium, . , 14-17 April, 201, Obernkirchen, Germany: Universität Göttingenmateus__milan_2012_new_dinosaur_and_pterosaur_tracksites_from_the_late_jurassic.pdf
Mateus, O., Pereira B., Rocha R., & Kullberg J. C. (2018).  Aspiring Geopark Oeste in Portugal: scientific highlights and importance. 8th International Conference on UNESCO Global Geoparks. , 8-14 Sept., Adamello Brenta Geopark, Trentinomateus_et_al_2018_geopark_oeste.pdf
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