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Mateus, O., & Jacinto J. J. (2008).  Hemidactylus turcicus. (A Loureiro, N F de Almeida, M.A Carretero, O S Paulo, Ed.).Atlas dos Anfíbio e Répteis de Portugal. 134-135. Abstract
Mateus, O., Maidment S., & Christiansen N. (2009).  A new long-necked 'sauropod-mimic' stegosaur and the evolution of the plated dinosaurs. Proceedings of the Royal Society of London B. 276, 1815-1821., Jan Abstractmateus_et_al_2009_stegosaur_miragaia_complete_with_suppl.pdfWebsite

Stegosaurian dinosaurs have a quadrupedal stance, short forelimbs, short necks, and are generally considered to be low browsers. A new stegosaur, Miragaia longicollum gen. et sp. nov., from the Late Jurassic of Portugal, has a neck comprising at least 17 cervical vertebrae. This is eight additional cervical vertebrae when compared with the ancestral condition seen in basal ornithischians such as Scutellosaurus.
Miragaia has a higher cervical count than most of the iconically long-necked sauropod dinosaurs. Long neck length has been achieved by ‘cervicalization’ of anterior dorsal vertebrae and probable lengthening of centra. All these anatomical features are evolutionarily convergent with those exhibited in the necks of
sauropod dinosaurs. Miragaia longicollum is based upon a partial articulated skeleton, and includes the only known cranial remains from any European stegosaur. A well-resolved phylogeny supports a new clade that unites Miragaia and Dacentrurus as the sister group to Stegosaurus; this new topology challenges the common view of Dacentrurus as a basal stegosaur.

Mateus, O. (2008).  Two ornithischian dinosaurs renamed: Microceratops Bohlin 1953 and Diceratops Lull 1905. Journal of Paleontology. 82, , Number 2 Abstract
Mateus, O., Marzola M., Schulp A. S., Jacobs L. L., Polcyn M. J., Pervov V., Gonçalves A. O. {\'ı}mpio, & Morais M. L. (2017).  Angolan ichnosite in a diamond mine shows the presence of a large terrestrial mammaliamorph, a crocodylomorph, and sauropod dinosaurs in the Early Cretaceous of Africa. Palaeogeography, Palaeoclimatology, Palaeoecology. 471, 220–232., apr: Elsevier {BV} AbstractWebsite
Mateus, O., & Antunes T. M. (2001).  Draconyx loureiroi, a new camptosauridae (Dinosauria, Ornithopoda) from the Late Jurassic of Lourinhã, Portugal. Annales de Paleontologie. 87, 61-73. Abstractmateus_antunes_2001_draconyx_loureiroi_a_new_camptosauridae_dinosauria_ornithopoda_from_the_late_jurassic_of_lourinha_portugal.pdfWebsite

A new ornithopod dinosaur is described here under the name of Draconyx loureiroi n. gen., n. sp. on teeth, caudal vertebrae, forelimb, hindlimb, and foot material that were found in association in the Late Jurassic-Tithonian of Lourinhã, Portugal. Draconyx is a Camptosauridae related to Camptosaurus.

Mateus, O. (2012).  Evidence for presence of clavicles and interclavicles in sauropod dinosaurs and its implications on the furcula-clavicle homology. Journal of Vertebrate Paleontology. 184–185., 1 Abstract

Clavicles and interclavicles are plesiomorphically present in Reptilia. However, several groups show reduction or even loss of these elements. Crocodylimorpha, e.g., lost the clavicles, whereas dinosaurs are generally interpreted to only preserve the clavicles, the theropod furcula representing an unique case of fused clavicles. In sauropods, reports of clavicles are relatively frequent in non-titanosauriforms. These elements are elongated, curved, and rather stout bones with a spatulate and a bifurcate end. However, they were always found as single bones, and differ from the relatively short and unbifurcated clavicles found articulated with the scapulae of basal sauropodomorphs. Elements from the Howe Quarry (Late Jurassic; Wyoming, USA) shed new light on these interpretations. Besides the elongated, curved bones (herein named morphotype A), also pairs of symmetric, L-shaped bones were recovered (morphotype B), associated with diplodocid dorsal and cervical vertebrae. Elements resembling morphotype B - articulated between the scapulae - have recently been reported from a diplodocid found near Tensleep, Wyoming. Taphonomic evidence, as well as the fact that they were preserved in symmetrical pairs, therefore implies that morphotype B represents the true sauropod clavicles. Contrary to earlier reports, morphotype A elements from the Howe Quarry, as well as of previously reported specimens show a symmetry plane following the long axis of the elements. It is thus possible that the morphotype A elements were single bones from the body midline. The only such element present in the pectoral girdle of tetrapods are the interclavicle and the furcula. Comparison with crocodilian and lacertiform interclavicles indicates that the bifurcate end of the sauropod elements might represent the reduced transverse processes of the anterior end, and the spatulate end would have covered the coracoids or sternal plates ventrally. The presence of both clavicles and interclavicles in the pectoral girdle stiffens the anterior trunk, and enhances considerably its stability. Such an enforcement might have been needed in diplodocids due to the strong lateral forces induced to the fore-limbs by the posteriorly placed center of mass (due to shorter fore- than hind-limbs), as well as lateral movements of the enormously elongated necks and tails. The absence of clavicles and interclavicles in titanosauriforms coincides with the development of wide-gauge locomotion style. The presence of interclavicles in sauropods supports the recently proposed homology of the furcula with the interclavicle, instead of representing fused clavicles. Interclavicles were thus not lost, but may have remained cartilaginous or have yet to be found in basal dinosauriforms.

Mateus, O., & Antunes M. T. (2000).  Ceratosaurus sp. (Dinosauria: Theropoda) in the Late Jurassic of Portugal. Abstract volume of the 31st International Geological Congress. , Rio de Janeiro, Brazil Abstractmateus__antunes_2000_-_ceratosaurus_in_portugal.pdf


Mateus, O., Laven T., & Knotschke N. (2004).  A dwarf between giants?: A new late Jurassic sauropod from Germany. Journal of Vertebrate Paleontology. 23, 90., Number suppl. to 3 Abstract
Mateus, O., & Antunes M. T. (2002).  Novos vestígios de dinossauros saurópodes do Jurássico Superior da Lourinhã (Portugal). Congresso Ibérico de Herpetologia. , Évora Abstract
Mateus, O., Jacobs L. L., Polcyn M. J., Myers T. S., & Schulp A. S. (2015).  The fossil record of testudines from angola from the turonian to oligocene. Society of Vertebrate Paleontology Annual Meeting. 177., Dallasmateus_et_al_2015_testudines_angola_svp_abstract.pdf
Mateus, O. (2009).  DINOSAUR EGGSHELL AND EMBRYO LOCALITIES IN LOURINHA FORMATION, LATE JURASSIC, PORTUGAL. Journal of Vertebrate Paleontology. 29, 76A–76A., 1 Abstract
Mateus, O. (2016).  Late Jurassic of Morrison Formation and Portugal tetrapods compared: a model to explain faunal exchange and similarity. Annual Meeting of the Society of Vertebrate Paleontology. 185., Salt Late City: Journal of Vertebrate Paleontology, Program and Abstracts, 2016 Abstractmateus_2016_late_jurassic_morrison_svp_abstract.pdf

The precursor of the North Atlantic existed between the North American and Iberian blocks from the earliest Jurassic Hettangian and has been ever expanding since. By the Kimmeridgian and Tithonian, when much of the Morrison Fm rocks were deposited, the proto-Atlantic was more than 300 km wide at 27° paleolatitude between North America and Iberia. Macrovertebrate paleontology reveals a unique story to the isolation of Iberia and instead suggest a paleogeographic land connection between North American and Iberia. Torvosaurus, Allosaurus, Ceratosaurus, Stegosaurus, Supersaurus and others have a distribution restricted to Morrison Formation in North America and Lourinhã Formation in Portugal. A novel paleogeographic model is here suggested: (1) around the Middle–Late Jurassic transition there is a major palaeoceanographic and palaeoclimatic reorganization, coincidental to a major eustatic sea-level drop and uplift associated with the Callovian– Oxfordian Atlantic Regressive Event; (2) creating an ephemeral land bridge presenting a temporary opportunity for terrestrial gateways likely across the Flemish Cap and Galician Bank land masses, allowing large dinosaurian taxa to cross the northern proto-Atlantic in both directions; (3) finally, a Callovian–Oxfordian faunal exchange around the 163 Ma, through latest Kimmeridgian at 152 Ma (the age of equivalent genera in both Morrison and Portugal), is was an interval that allowed speciation, but retaining generic similarity of vertebrates. This model is consistent with the chronology and taxonomy required for speciation of the Iberian and American forms, exemplified by the coeval sister-taxa pairs Torvosaurus tanneri and T. gurneyi, Allosaurus fragilis and A. europaeus, or Supersaurus vivianae and S. lourinhanensis. While some of the smaller animals in the fauna show Morrison/Portugal affinities, most from Iberia have European or even Asian affinities. The larger-bodied fauna are more closely related to Morrison than to mainland Europe (except for dacentrurine stegosaurs). The body size differences and affinities of taxa across paleogeography is comparable to what is observed today across the Wallace Line. Migration may have also occurred in both directions. The closest relative of Torvosaurus is likely the European Bathonian Megalosaurus, thus the presence of the genus in North America represents a European migration. On other hand, Allosaurus and Supersaurus origins are consistent with a North American origin, representing an westto-east migration.

Mateus, I., Mateus H., Antunes M. T., Mateus O., Taquet P., Ribeiro V., & Manuppella G. (1997).  Couvée, œufs et embryons d'un Dinosaure Théropode du Jurassique supérieur de Lourinhã (Portugal). Comptes Rendus de l'Academie de Sciences - Serie IIa: Sciences de la Terre et des Planetes. 325, 71–78., Number 1 Abstract
Mateus, O., Overbeeke M., & Rita F. (2008).  Dinosaur Frauds, Hoaxes and "Frankensteins": How to distinguish fake and genuine vertebrate fossils. Journal of Paleontological Techniques. 2, 1-5. Abstract
Mateus, O. (2014).  Preliminary Magnetostratigraphy for the Jurassic–Cretaceous Transition in Porto da Calada, Portugal. (Rogério Rocha, João Pais, Kullberg, {José Carlos}, Stanley Finney, Ed.).STRATI 2013:First International Congress on Stratigraphy At the Cutting Edge of Stratigraphy. 873–877., 1: Springer International Publishing Switzerland Abstract

We present a stratigraphic log supporting a preliminary magnetostratigraphy of a Tithonian–Berriasian section in Porto da Calada (Portugal). Based on biostratigraphy and reversed and normal magnetostratigraphy, the location of the Tithonian–Berriasian boundary is tentatively located at ca. 52 m, not in disagreement with former proposals. Due to the occurrence of later remagnetization (diagenesis), the magnetostratigraphic definition of the Tithonian–Berriasian section at the Cabo Espichel (Portugal) location was not able to be established

Mateus, O., Jacobs L. L., Schulp A. S., Polcyn M. J., Tavares T. S., Neto A. B., Morais M. L., & Antunes M. T. (2011).  Angolatitan adamastor, a new sauropod dinosaur and the first record from Angola.. Anais da Academia Brasileira de Ciências. 83, 221-233., Jan Abstractmateus_et_al_2011_angolatitan_adamastor_sauropod.pdfWebsite

A forelimb of a new sauropod dinosaur (Angolatitan adamastor n. gen. et sp.) from the Late Turonian of Iembe (Bengo Province) represents the first dinosaur discovery in Angola, and is one of the few occurrences of sauropod dinosaurs in sub-Saharan Africa collected with good chronological controls. The marginal marine sediments yielding the specimen are reported to be late Turonian in age and, thus it represents a non-titanosaurian sauropod in sub-Saharan Africa at a time taken to be dominated by titanosaurian forms. Moreover, Angolatitan adamastor is the only basal Somphospondyli known in the Late Cretaceous which implies in the existence of relict forms in Africa.

Mateus, O., Dyke G., Motchurova-Dekova N., Ivanov P., & Kamenov G. D. (2008).  The Bulgarian dinosaur: did it exist? European late Cretaceous ornithomimosaurs. 56th Symposium of Vertebrate Palaeontology and Comparative Anatomy. 47–47., Dublin Abstract
Mateus, O., & Milàn J. (2011).  New dinosaur and pterosaur tracksites from the Late Jurassic of Portugal. Dinosaur Tracks 2011 An International Symposium, . , 14-17 April, 201, Obernkirchen, Germany: Universität Göttingenmateus__milan_2012_new_dinosaur_and_pterosaur_tracksites_from_the_late_jurassic.pdf
Mateus, O., and Dinis J., Cunha P. P., & and (2017).  The Lourinhã Formation: the Upper Jurassic to lower most Cretaceous of the Lusitanian Basin, Portugal – landscapes where dinosaurs walked. Ciências da Terra - Earth Sciences Journal. 19, 75–97., sep, Number 1: {NOVA}.{ID}.{FCT} AbstractWebsite
Mateus, O., Mannion P. D., & Upchurch P. (2014).  Zby atlanticus, a new turiasaurian sauropod (Dinosauria, Eusauropoda) from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology. 34, 618–634., Number 3 Abstract
Mateus, O. (2012).  A preliminary report on coprolites from the Late Triassic part of the Kap Stewart Formation, Jameson Land, East Greenland. Bulletin of the New Mexico Museum of Natural History and Science. 57, 203–205., 1, Number NA Abstract

The basal part of the Triassic-Jurassic (Rhaetian-Sinemurian) Kap Stewart Formation, exposed at Jameson Land, East Greenland, yields an extensive coprolite collection from black, parallel-laminated mudstone (“paper shale”), representing an open lacustrine system. Preliminary investigations show three different types of coprolites: elongated cylindrical masses, composed of irregularly wrapped layers; elongated cylindrical masses with constriction marks; and spirally-coiled specimens.

Mateus, O., & Milan J. (2005).  Ichnological evidence for giant ornithopod dinosaurs in the Late Jurassic Lourinhã Formation, Portugal. Abstract Book of the International Symposium on Dinosaurs and Other Vertebrates Palaeoichnology. 60., Fumanya, Barcelona Abstract
Mateus, O., Araújo R., Natário C., & Castanhinha R. (2011).  A new specimen of the theropod dinosaur Baryonyx from the early Cretaceous of Portugal and taxonomic validity of Suchosaurus. Zootaxa. 54-68., Number 2827 Abstract
Mateus, O. (2008).  Two ornithischian dinosaurs renamed: Microceratops Bohlin 1953 and Diceratops Lull 1905. Journal of Paleontology. 82, 423., Number 2 Abstractmateus_2008_two_ornithischians_renamed__microceratops_bohlin_1953_and_diceratops_lull_.pdfWebsite

dinosaur genera Diceratops Lull, 1905 and Microceratops Bohlin, 1953 are preoccupied by the Hymenoptera insects, Diceratops Foerster, 1868 and Microceratops Seyrig, 1952, respectively. Therefore, the name of the ceratopsian dinosaur Diceratops Lull, 1905 from the Late Cretaceous of United States is a junior homonym of the hymenoptera Diceratops Foerster, 1868. Diceratus n. gen. (Greek di ‘‘two,’’ Greek ceratos ‘‘horned’’) is proposed as the replacement name of Diceratops Lull, 1905. Some workers have considered Diceratops synonymous with Triceratops (e.g., Dodson and Currie, 1990) but it was reinstated by Forster (1996) after analysis of the characteristics of all existing ceratopsid skulls, and recent reviews (e.g., Dodson et al., 2004) have considered Diceratops a valid genus.
Due to preoccupation, the name of the ceratopsian dinosaur Microceratops Bohlin, 1953 from the Cretaceous of the Gobi is
a junior homonym of the insect Microceratops Seyrig, 1952. Microceratus n. gen. (Greek micro ‘‘small,’’ Greek ceratos ‘‘horned’’) is proposed as the replacing name of Microceratops Bohlin, 1953.
Sereno (2000:489) has declared Microceratops a nomen dubium since the holotype material lacks any diagnostic features, a
convention followed by You and Dodson (2004:480). However, the name is still used by Le Loeuff et al. (2002), Lucas (2006),
Alifanov (2003) and Xu et al. (2002), and such practice justifies the renaming of the genus.
In order to preserve some stability, the names chosen here deliberately preserve the same prefixes.

Mateus, O., Dyke G. A. J., Motchurova-Dekova N., Kamenov G. D., & Ivanov P. (2010).  The first record of a dinosaur from Bulgaria. Lethaia. 43, 88-94., Number 1 Abstract
Mateus, O., & Antunes M. T. (2003).  A New Dinosaur Tracksite in the Lower Cretaceous of Portugal. Ciências da Terra. 15, 253-262. Abstractmateus__antunes_2003-_early_k_dino_tracks_portugal.pdfWebsite

A new Lower Cretaceous (Aptian-Albian) dinosaur tracksite at the Olhos de Água beach is described. It is the first vertebrate fossil finding ever found in the concerned unit, and yielded 128 tracks in 17 trackways within an area of ca. 80 square metres.
Three tridactyl footprint morphotypes have been recognized: - Type 1 (“Iguanodontipus-like”) - trackways D, F, K, J and P; - Type 2 (large theropod), although larger in size, typically from a Grallator-like theropod footprint, i.e. A, B, G, H and O trackways; - Type 3 (medium size theropod); M is the only track of this type. There are other, poorly preserved, unidentified trackways. The theropod, swinging trackway B was produced by an animal that was limping. The theropod track M starts eastwards but drastically changes westwards, speeding up at the same time; this dinosaur decided to turn around and run in the opposite direction.
This site shows three main trackway directions: to the South, to the East, and westwards. Except for the trackway O, large theropods A, B, G and H walked southwards. Perpendicularly to these, ornithopods, small theropods and unidentified trackmakers walked towards East (5) and West (7). The segregation of trackmakers and directions, with large theropod trackways southwards and other dinosaurs’ west or eastwards, may mean that large theropods patrolled a walkway area to an important resource, most probably water, often frequented by ornithopods and smaller theropods. There is no evidence of social behavior or gregarism: footprints’ overposition shows that the large, southwards walking theropods passed on different occasions. Three trackway sequences can be established by chronologic order.

Mateus, I., Mateus H., Antunes M. T., Mateus O., Taquet P., Ribeiro V., & Manuppella G. (1998).  Upper Jurassic Theropod Dinosaur embryos from Lourinhã (Portugal). Upper Jurassic paleoenvironments in Portugal, Mem. Acad. Ciências de Lisboa. 37, 101-109. Abstract
Mateus, O. (2008).  Checklist for Late Jurassic reptiles and amphibians from Portugal. Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 55–55., Coimbra Abstract
Mateus, O., & Milan J. (2008).  Sauropod forelimb flexibility deduced from deep manus tracks. (University of, Glasgow, Ed.).52th Paleontological Association Annual Meeting. 18th-21st December 2008. 67-68. Abstract
Mateus, O., & Jacinto J. J. (2002).  Contribuição para o estudo de Hemidactylus turcicus (Reptilia, Gekkonidae): ritmos de actividade e microhabitat em Évora, Portugal. VII Congresso Luso-Espanhol de Herpetologia. 136., Évora Abstract


Mateus, O. (2011).  New fossil whales from Angola. Abstracts of the 71st Annual Meeting of the Society of Vertebrate Paleontology. 119., 1 Abstract
Mateus, O. (2010).  Tectonic Drift, Climate, and Paleoenvironment of Angola Since the Cretaceous. Journal of Vertebrate Paleontology. 02., 1 Abstract

Africa is the only continent that now straddles arid zones located beneath the descending limbs of both the northern and southern Hadley cells, and it has done so since it became a distinct continent in the Early Cretaceous. Since that time, Africa has drifted tectonically some 12 degrees north and rotated approximately 45 degrees counterclockwise. This changing latitudinal setting and position of the landmass under the relatively stable Hadley Cells is manifested as southward migration of climatic zones over the past 132 million years. Data from kerogen, X-ray diffraction analysis of sedimentary matrix, carbon isotopes from shell samples and tooth enamel,new 40Ar/39Ar radiometric dates, pollen and plant macrofossils, and fossil vertebrates indicate a productive upwelling system adjacent to a coastal desert since the opening of the South Atlantic Ocean; however, the position of the coastal desert has migrated southward as Africa drifted north, resulting in today's Skeleton Coast and Benguela Current. This migration has had a profound effect on the placement of the West African coast relative to areas of high marine productivity and resulting extensive hydrocarbon deposits, on the placement of arid zones relative to the continent especially the Skeleton Coast desert, on the climatic history of the Congo Basin (which shows a Late Cretaceous decrease in aridity based on the relative abundance of analcime in the Samba core), and in reducing the southern temperate region of Africa from 17{%} of continental area during the Cretaceous to 2{%} today. We show here that these related geographic and environmental changes drove ecological and evolutionary adjustments in southern African floras and faunas, specifically with respect to the distribution of anthropoid primates, the occurrence of modern relicts such as the gnetalean Welwitschia mirabilis, endemism as in the case of ice plants, and mammalian adaption to an open environment as in springhares. Africa's tectonic drift through climate zones has been a first-order environmental determinant since the Early Cretaceous.

Mateus, O., Jacobs L. L., Polcyn M. J., Schulp A. S., Neto A. B., & Antunes M. T. (2008).  Dinosaur and turtles from the Turonian of Iembe, Angola. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontologia de Vertebrados. 156., Neuquén, Argentina Abstractmateus_et_al_2008_dinosaur_and_turtles_from_the_turonian_of_iembe_angola.pdf


Mateus, O., & Antunes M. T. (2000).  On the presence of Ceratosaurus sp. (Dinosauria: Theropoda) in the Late Jurassic of Portugal. Abstract volume of the 31st International Geological Congress. , Rio de Janeiro, Brazil Abstract
Mateus, O., Natario C., Araujo R., & Castanhinha R. (2008).  A new specimen of spinosaurid dinosaur aff. Baryonyx from the Early Cretaceous of Portugal. (Universidade de, Coimbra, Ed.).Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 51., Coimbra Abstract
Mateus, O. (2014).  Comparison of modern and fossil Crocodylomorpha eggs and contribution to the oophylogeny of Amniota. Annual Meeting of the European Association of Vertebrate Palaeontologists. XII Annual Meeting of the European Association of Vertebrate Palaeontologists, 192., 1 Abstract
Mateus, O., & Castanhinha R. (2008).  PaleoAngola- Predadores de um oceano primitivo. National Geographic Portugal. 8, 26–33., Number 91 Abstract
Mateus, O. (2013).  Crocodylomorphs from the Mesozoic of Portugal and a new skull of eusuchian from the Late Cretaceous. 2013 Hwaseong International Dinosaurs Expedition Symposium, pp.66-67.. , Hwaseong, South Korea Abstractmateus_2013_crocodylomorphs_portugal_new_skull.pdf

The diversity of fossil crocodylomorphs in Portugal is high, with occurrence as old as Mystriosaurus (=Steneosaurus) bollensis from the Lower Jurassic. The Late Jurassic forms are the better documented, and include the following taxa: Machimosaurus hugii, Lisboasaurus estesi Seiffert, 1973, Lusitanisuchus mitrocostatus Seiffert, 1975; Schwarz & Fechner 2004, Theriosuchus guimarotae Schwarz and Salisbury 2005, Cf. Alligatorium, Goniopholis baryglyphaeus, and a crocodylomorph-like eggs in dinosaur nests (Mateus et al., 1998; Ricqlès et al., 2001). From the Lower Cretaceous were reported a few dinosaurs but its record is strangely scarce in crocodylomorphs (Mateus et al., 2011). The Upper Cretaceous crocodiles show a large diversity, but it is mostly based in fragmentary material that require revision, such as “Crocodylus” blavieri? Grey from the Upper Campanian - Maastrichtian of Viso, near Aveiro (initially reported by Sauvage 1897-98), Goniopholis cf. crassidens Owen 1841 and Oweniasuchus pulchelus Jonet 1981. Moreover there is a fascinating, but poorly understood, crocodylomorph diversity in the Cenomanian of Portugal, documented by fragmentary specimens that have been doubtfully assigned to Thoracosaurus Leidy 1852 of the Middle Cenomanian of Cacém, to the nomen dubium Oweniasuchus lusitanicus Sauvage 1897-98 (interpreted as a mesosuchian goniopholid) based in a fragmentary mandible from the Campanian-Maastrichtian, and also from the Middle Cenomanian of Portugal, Buffetaut and Lauverjat (1978) report an fragmentary unidentified possible dyrosaurid from Nazaré. All this specimens are too incomplete to be compared with the specimen here described. In contrast, Cenozoic crocodiles of Portugal are often known after complete skulls and several individuals. The taxa list include Iberosuchus macrodon (Lower to Middle Eocene), Tomistoma calaritanus (Early Miocene) and T. lusitanica (Burdigalian-Helvetian), and Diplocynodon sp. (Antunes, 1961, 1987, 1994).
At least, two different morphotypes of crocodylomorph eggs from the Late Jurassic of Lourinhã Formation are also known.
A new specimen here reported of crocodile based in a partial skull and mandible (ML1818) from the Uppermost Middle Cenomanian platform carbonates of Baixo Mondego, west central Portugal (Tentúgal Fm., Callapez, 2004). The taxon is phylogenetically positioned as a basal Eusuchia, due to the choanae enclosed by the pterygoid, and closely related with stem Crocodylia and Borealosuchus. This specimen represents the only well documented and valid eusuchian species in the Cenomanian of Europe and is the oldest representative of an eusuchian crocodylomorph, with the exception for the Barremian Hylaeochampsa vectiana.

Mateus,  O., de da Terra D. C. T. D. C. -, & e GeoBioTec - Geobiociências G. G. (2014).  The Late Triassic of Jameson Land revisited. Abstract
Mateus, O. (1996).  Situação populacional de Hemidactylus turcicus em Évora-Portugal. Actas do IV Congresso Luso-Espanhol de Herpetologia. 45., Porto Abstractmateus_1996_population_situation_of_hemidactyus_turcicus_in_evora_portugal_iv_congresso_lusoespanhol_herpetologia.pdf

A survey of Hemidactylus turcicus L. (Reptilia, Gekkonidae) was carried out from March to October
of 1996, in temporally limited transects, using quadrats of 160x160 meters, in the World Heritage site
of Évora. Occasional observations were also made in 1995 and 1996. Hemidactylus turcicus is
common in the sampled area, occurring in 56% of the quadrats, and prefers quiet streets. The species
Tarentola mauritanica was not observed in Évora but it appears 10 Km to the Southwest of this town.
Diurnal activity and winter activity were not observed.

Mateus, O., & The Gigantic dinosaur E. (2006).  The European Enigmatic Dinosaur Evolution (in Japanese). Abstract
Mateus, O., Taquet P., Antunes M. T., Mateus H., & Ribeiro V. (1998).  Theropod dinosaur nest from Lourinhã, Portugal. Journal of Vertebrate Paleontology. 18, 61A., Number (Suppl. 3) Abstractmateus_et_al_1998_theropod_dinosaur_nest_from_lourinha_portugal_svp.pdfWebsite

More than 100 well preserved dinosaur eggs have been discovered in the Upper Jurassic levels (Tithonian) of Lourinhã, Portugal. The eggshels dispersed in a big area with 11 meters in the highest diameter with high concentration in the middle.

Mateus, O., & Antunes M. T. (2003).  A new dinosaur tracksite in the Lower Cretaceous of Portugal. Ciências da Terra (UNL). 15, 253–262. Abstract
Mateus, O., & Campos H. (2018).  Loulé há mais de 220 Milhões de anos: os vertebrados fósseis do Algarve triásico. Loulé: Territórios. Memórias. Identidades. 651-659.: Museu Nacional de Arqueologia | Imprensa Nacionalmateus_campos2018_algarve_triasico.pdf