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Guillaume, A. R. D., Natário C., Moreno-Azanza M., & Mateus O. (2021).  New occurences of Celtedens (Lissamphibia: Albanerpetontidae) from the Lourinhã Formation. XVIII Conference of the European Association of Vertebrate Palaeontologists. 87. Abstractguillaume_et_al_2021_celtedens.pdf

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Adams, T. L., Polcyn M. J., Mateus O., Winkler D. A., & Jacobs L. L. (2010).  New occurrence of the long-snouted crocodyliform,Terminonaris cf.T. robusta, from Woodine Formation (Cenomanian) ot Texas. 52A., Jan Abstract

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Rotatori, F., Moreno-Azanza M., & Mateus O. (2019).  New ornithopod dinosaur remains from the Late Jurassic Lourinhã Formation. 17th Conference of the EAVP. 100., Bruxelles: European Association of Vertebrate Paleontologistsrotatori_et_al_2019_ornithopod_portugal_eavp_2019_abstract.pdf
López-Rojas, V., Mateus O., Milàn J., Wings O., Klein N., & Clemmensen L. B. (2021).  A new phytosaur from the Late Triassic of Jameson Land, Greenland. 3rd Palaeontological Virtual Congress. 207.: ISBN 978-84-09-36657-6 Abstractlopez_rojas_2021_pvc3_greenland.pdf

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López-Rojas, V., Clemmensen L. B., Milàn J., Wings O., Klein N., & Mateus O. (2023).  A new phytosaur species (Archosauriformes) from the Upper Triassic of Jameson Land, central East Greenland. Journal of Vertebrate PaleontologyJournal of Vertebrate Paleontology. e2181086., 2023: Taylor & Francis Abstracta_new_phytosaur_species_archosauriformes_from_the_upper_triassic_of_jameson_land_central_east_greenland.pdfWebsite

ABSTRACTHerein we describe phytosaurs from thin fluvial overbank sandstones of the Upper Triassic Malmros Klint Formation of the Fleming Fjord Group (central East Greenland). The new sample includes more than 150 disarticulated bones and teeth from small to large specimens belonging to at least four individuals. The fossils mostly consist of teeth and postcranial elements and permit the recognition of a new species of Mystriosuchus, M. alleroq, diagnosed by an L-shaped quadratojugal whose anterior suture trends anterodorsally and a tripartite degree of heterodonty. Humeral diaphyseal histology of one specimen reveals a fairly compact cortex that surrounds a cancellous medullary region followed by a remodeling zone containing scattered secondary osteons. Primary bone tissue is parallel-fibred with a moderate to low vascular density. The cortex is cyclically interrupted by distinct growth marks indicating a seasonal environment. A change in growth rate from moderate to low is documented within the outer cortex, indicating that at least this individual was close to somatic maturity. Mystriosuchus has formerly been known as an exclusively European taxon. The new findings support the European faunal influence in East Greenland during the Late Triassic inferred from other taxa such as temnospondyls and archosaurs. The mid-late Norian age of European Mystriosuchus suggests an additional age constraint for the vertebrate-bearing portion of the Malmros Klint Formation.

Puértolas-Pascual, E., Marx M., Mateus O., Saleiro A., Fernandes A. E., Marinheiro J., Tomás C., & Mateus S. (2021).  A new plesiosaur from the Lower Jurassic of Portugal and the early radiation of Plesiosauroidea. Acta Palaeontologica Polonica. 66(2), 369-388. Abstracta_new_plesiosaur_from_the_lower_jurassic_of_portugal_and_the_early_radiation_of_plesiosauroidea.pdfWebsite

A new plesiosaur partial skeleton, comprising most of the trunk and including axial, limb, and girdle bones, was collected in the lower Sinemurian (Coimbra Formation) of Praia da Concha, near São Pedro de Moel in central west Portugal. The specimen represents a new genus and species, Plesiopharos moelensis gen. et sp. nov. Phylogenetic analysis places this taxon at the base of Plesiosauroidea. Its position is based on this exclusive combination of characters: presence of a straight preaxial margin of the radius; transverse processes of mid-dorsal vertebrae horizontally oriented; ilium with sub-circular cross section of the shaft and subequal anteroposterior expansion of the dorsal blade; straight proximal end of the humerus; and ventral surface of the humerus with an anteroposteriorly long shallow groove between the epipodial facets. In addition, the new taxon has the following autapomorphies: iliac blade with less expanded, rounded and convex anterior flank; highly developed ischial facet of the ilium; apex of the neural spine of the first pectoral vertebra inclined posterodorsally with a small rounded tip. This taxon represents the most complete and the oldest plesiosaur species in the Iberian Peninsula. It is also the most complete, best preserved, and oldest marine vertebrate in the region and testifies to the incursion of marine reptiles in the newly formed proto-Atlantic sea, prior to the Atlantic Ocean floor spreading in the Early Cretaceous.

Smith, A. S., Araújo R., & Mateus O. (2012).  A new plesiosauroid from the Toarcian (Lower Jurassic) of Alhadas, Portugal. Acta Palaeontologica Polonica. 57(2), 257–266., Jan Abstractsmith__araujo__mateus_2012_a_new_plesiosauroid_from_the_toarcian_lower_jurassic_of_alhadas_portugal.pdf

A partial plesiosauroid skull from the São Gião Formation (Toarcian, Lower Jurassic) of Alhadas, Portugal is re−evaluated and described as a new taxon, Lusonectes sauvagei gen. et sp. nov. It has a single autapomorphy, a broad triangular parasphenoid cultriform process that is as long as the posterior interpterygoid vacuities, and also a unique character combination, including a jugal that contacts the orbital margin, a distinct parasphenoid–basisphenoid suture exposed between the posterior interpterygoid vacuities, lack of an anterior interpterygoid vacuity, and striations on the ventral surface of
the pterygoids. Phylogenetic analysis of Jurassic plesiosauroids places Lusonectes as outgroup to “microcleidid elasmosaurs”, equivalent to the clade Plesiosauridae. Lusonectes sauvagei is the only diagnostic plesiosaur from Portugal, and the westernmost occurrence of any plesiosaurian in Europe.

Smith, A. S., Araújo R., & Mateus O. (2012).  A new plesiosauroid from the Toarcian (Lower Jurassic) of Alhadas, Portugal. Acta Palaeontologica Polonica. 57, 257-266., Number 2 Abstract
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Schulp, A., Mateus O., Polcyn M., & Jacobs L. (2006).  A new Prognathodon (Squamata : Mosasauridae) from the Cretaceous of Angola. JOURNAL OF VERTEBRATE PALEONTOLOGY. 26, 122A-122A., Jan Abstractschulp_et_al_2006.pdf

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Marinheiro, J., Mateus O., Alaoui A., Amani F., Nami M., & Ribeiro C. (2014).  New Quaternary fossil sites from the Middle Atlas of Morocco. Comunicações Geológicas. 101, Especial I, 485-488. Abstractmarinheiro_et_al_2014_new_quaternary_fossil_sites_from_the_middle_atlas_of_morocco.pdf

The paleontological richness of Morocco has been scientifically known since at least the early 20th century. The region of the Middle Atlas, more specifically the Boulemane area, has been however only sparsely studied since the 1960s when it provided vertebrate fossils from the Middle Jurassic. In September 2013, a Moroccan-Portuguese expedition to the village of Taghrout, Boulemane, made excavations in a Pleistocene fossil site that once was a small high-altitude sedimentary basin, uncharted in previous geological maps. The excavations yielded bone material from large mammals, the most common findings are elephants ascribed to Elephas, but artiodactyls, turtles, and in-situ Acheulean tools were also collected. This represents a new and important paleontological and archeological site. In addition to the discoveries of Taghrout, the expedition also retrieved Quaternary vertebrate material from a nearby cave and found new Jurassic localities, with arcosaur bones and dinosaur footprints, in El Mers.

Marinheiro, J., Mateus O., Alaoui A., Amani F., Nami M., & Ribeiro C. (2014).  New Quaternary fossil sites from the Middle Atlas of Morocco. Comunica\\c cões Geológicas. 101, Especial I, 485-488. Abstract

The paleontological richness of Morocco has been scientifically known since at least the early 20th century. The region of the Middle Atlas, more specifically the Boulemane area, has been however only sparsely studied since the 1960s when it provided vertebrate fossils from the Middle Jurassic. In September 2013, a Moroccan-Portuguese expedition to the village of Taghrout, Boulemane, made excavations in a Pleistocene fossil site that once was a small high-altitude sedimentary basin, uncharted in previous geological maps. The excavations yielded bone material from large mammals, the most common findings are elephants ascribed to Elephas, but artiodactyls, turtles, and in-situ Acheulean tools were also collected. This represents a new and important paleontological and archeological site. In addition to the discoveries of Taghrout, the expedition also retrieved Quaternary vertebrate material from a nearby cave and found new Jurassic localities, with arcosaur bones and dinosaur footprints, in El Mers.

Marinheiro, J., Mateus O., Alaoui A., Amani F., Nami M., & Ribeiro C. (2014).  New Quaternary fossil sites from the Middle Atlas of Morocco. Comunicações Geológicas. 101, 485–488., Number Especial I: Laboratório Nacional de Energia e Geologia Abstract
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Marinheiro, J., Mateus O., Alaoui A., Amani F., Nami M., & Ribeiro C. (2014).  New quaternary fossil sites from the middle atlas of morocco,Novas jazidas fossilíferas do Quaternário do Médio-Atlas de Marrocos. Comunicacoes Geologicas. 101, 485-488. Abstract
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Guillaume, A. R. D., Moreno-Azanza M., & Mateus O. (2022).  New species of Albanerpetontidae (Lissamphibia) from the Lourinhã Formation. XXXVII Jornadas SEP y V Congreso Ibérico de Paleontología. 94., Cuenca: ISBN 13: 978-84-09-44478-6 Abstractguillaume_et_al_2022_albanerpetontidae_jornadas_sep_cuenca_abstract_2022.pdf

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Brusatte, S. L., Butler R. J., Mateus O., & Steyer S. J. (2015).  A new species of Metoposaurus from the Late Triassic of Portugal and comments on the systematics and biogeography of metoposaurid temnospondyls. Journal of Vertebrate PaleontologyJournal of Vertebrate Paleontology. e912988., 2015: Taylor & Francis Abstractbrusatte_et_al2015metoposaurusportugal.pdfWebsite

ABSTRACTMetoposaurids are a group of temnospondyl amphibians that filled crocodile-like predatory niches in fluvial and lacustrine environments during the Late Triassic. Metoposaurids are common in the Upper Triassic sediments of North Africa, Europe, India, and North America, but many questions about their systematics and phylogeny remain unresolved. We here erect Metoposaurus algarvensis, sp. nov., the first Metoposaurus species from the Iberian Peninsula, based on several new specimens from a Late Triassic bonebed in Algarve, southern Portugal. We describe the cranial and pectoral anatomy of M. algarvensis and compare it with other metoposaurids (particularly other specimens of Metoposaurus from Germany and Poland). We provide a revised diagnosis and species-level taxonomy for the genus Metoposaurus, which is currently represented with certainty by three European species (M. diagnosticus, M. krasiejowensis, M. algarvensis). We also identify cranial characters that differentiate these three species, and may have phylogenetic significance. These include features of the braincase and mandible, which indicate that metoposaurid skulls are more variable than previously thought. The new Portuguese bonebed provides further evidence that metoposaurids congregated in fluvial and lacustrine settings across their geographic range and often succumbed to mass death events. We provide an updated paleogeographic map depicting all known metoposaurid occurrences, which shows that these temnospondyls were globally distributed in low latitudes during the Late Triassic and had a similar, but not identical, paleogeographic range as phytosaurs.http://zoobank.org/urn:lsid:zoobank.org:pub:083C80C6-0AB6-49E1-A636-6A8BDBC06A47ABSTRACTMetoposaurids are a group of temnospondyl amphibians that filled crocodile-like predatory niches in fluvial and lacustrine environments during the Late Triassic. Metoposaurids are common in the Upper Triassic sediments of North Africa, Europe, India, and North America, but many questions about their systematics and phylogeny remain unresolved. We here erect Metoposaurus algarvensis, sp. nov., the first Metoposaurus species from the Iberian Peninsula, based on several new specimens from a Late Triassic bonebed in Algarve, southern Portugal. We describe the cranial and pectoral anatomy of M. algarvensis and compare it with other metoposaurids (particularly other specimens of Metoposaurus from Germany and Poland). We provide a revised diagnosis and species-level taxonomy for the genus Metoposaurus, which is currently represented with certainty by three European species (M. diagnosticus, M. krasiejowensis, M. algarvensis). We also identify cranial characters that differentiate these three species, and may have phylogenetic significance. These include features of the braincase and mandible, which indicate that metoposaurid skulls are more variable than previously thought. The new Portuguese bonebed provides further evidence that metoposaurids congregated in fluvial and lacustrine settings across their geographic range and often succumbed to mass death events. We provide an updated paleogeographic map depicting all known metoposaurid occurrences, which shows that these temnospondyls were globally distributed in low latitudes during the Late Triassic and had a similar, but not identical, paleogeographic range as phytosaurs.http://zoobank.org/urn:lsid:zoobank.org:pub:083C80C6-0AB6-49E1-A636-6A8BDBC06A47

Brusatte, S. L., Butler R. J., Mateus O., & Steyer J. S. (2015).  A new species of Metoposaurus from the Late Triassic of Portugal and comments on the systematics and biogeography of metoposaurid temnospondyls. Journal of Vertebrate Paleontology. 35, , Number 3 Abstract
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Schulp, A. S., Polcyn M. J., Mateus O., Jacobs L. L., & Morais M. L. (2008).  A new species of Prognathodon (Squamata, Mosasauridae) from the Maastrichtian of Angola, and the affinities of the mosasaur genus Liodon. Proceedings of the Second Mosasaur Meeting, Fort Hays Studies Special Issue 3, Fort Hays State University, Hays, Kansas. 1-12. Abstractschulp_et_al_2008_prognathodon_kianda.pdf

Here we describe a new species of the mosasaurine genus Prognathodon from the Maastrichtian of Namibe, Angola, on the basis of five specimens which represent most of the cranial skeleton including the diagnostic quadrate. Phylogenetic analysis shows this new taxon, P. kianda nov. sp., to be the sister taxon to all other species of Prognathodon. It is unique amongst Prognathodon in possessing a high marginal tooth count and relatively small pterygoid teeth. The tooth morphology in the new taxon is reminiscent of some species of the genus Liodon, allowing association of Liodon-like dentition with otherwise Prognathodon-like crania, and thus resolves the long-standing question of the phylogenetic affinities of Liodon.

Schulp, A. S., Polcyn M. J., Mateus O., Jacobs L. L., & Morais M. L. (2008).  A new species of Prognathodon (Squamata, Mosasauridae) from the Maastrichtian of Angola, and the affinities of the mosasaur genus Liodon. Proceedings of the Second Mosasaur Meeting, Fort Hays Studies Special Issue 3, Fort Hays State University, Hays, Kansas. 1-12. Abstract
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Mateus, O., Maidment S. C. R., & Christiansen N. A. (2008).  A new specimen aff. Dacentrurus armatus (Dinosauria: Stegosauridae) from the Late Jurassic of Portugal. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontologia de Vertebrados. 157., Neuquén, Argentina Abstractmateus_et_al_2008_a_new_specimen_aff._dacentrurus_armatus_dinosauria_stegosauridae_from_the_late_jurassic_of_portugal.pdf

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Mateus, O., Maidment S. C. R., & Christiansen N. A. (2008).  A new specimen aff. Dacentrurus armatus (Dinosauria: Stegosauridae) from the Late Jurassic of Portugal. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 157., Neuquén, Argentina Abstract
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Mateus, O., Maidment S. C. R., & Christiansen N. A. (2008).  A new specimen aff. Dacentrurus armatus (Dinosauria: Stegosauridae) from the Late Jurassic of Portugal. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 157–157., Neuquén, Argentina Abstract
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Ferrari, L., Rotatori F. M., Camilo B., Moreno-Azanza M., & Mateus O. (2022).  New specimen of dryomorphan (Ornithischia, Iguanodontia) remains from the Upper Jurassic of Portugal. Abstract book of the XIX Annual Conference of the European Association of Vertebrate Palaeontologists, Benevento/Pietraroja, Italy, 27th June-2nd July 2022.. 61-62.: PalaeoVertebrata, Special Volume 1- 2022, 224. Doi: 10.18563/pv.eavp Abstractferrarietal_2022.pdf

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Mateus, O., Natário C., Araújo R., & Castanhinha R. (2008).  A new specimen of spinosaurid dinosaur aff. Baryonyx from the Early Cretaceous of Portugal. Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 51., Jan Abstract

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Mateus, O., Natario C., Araujo R., & Castanhinha R. (2008).  A new specimen of spinosaurid dinosaur aff. Baryonyx from the Early Cretaceous of Portugal. (Universidade de, Coimbra, Ed.).Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 51., Coimbra Abstract
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Mateus, O., Natario C., Araujo R., & Castanhinha R. (2008).  A new specimen of spinosaurid dinosaur aff. Baryonyx from the Early Cretaceous of Portugal. Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 51–51., Coimbra Abstract
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Mateus, O., Araújo R., Natário C., & Castanhinha R. (2011).  A new specimen of the theropod dinosaur Baryonyx from the early Cretaceous of Portugal and taxonomic validity of Suchosaurus. Zootaxa. 2827, 54–68., Jan Abstractmateus_et_al_2011_a_new_specimen_of_the_theropod_dinosaur_baryonyx_from_the_early_cretaceous_of_portugal_and_taxonomic_validity_of_suchosaurus.pdf

Although the Late Jurassic of Portugal has provided abundant dinosaur fossils, material from the Early Cretaceous is scarce. This paper reports new cranial and postcranial material of the theropod dinosaur Baryonyx walkeri found in the Barremian (Papo Seco Formation) of Portugal. This specimen, found at Praia das Aguncheiras, Cabo Espichel, consists of a partial dentary, isolated teeth, pedal ungual, two calcanea, presacral and caudal vertebrae, fragmentary pubis, scapula, and rib fragments. It represents the most complete spinosaurid yet discovered in the Iberian Peninsula and the most complete dinosaur from the Early Cretaceous of Portugal. This specimen is confidently identified as a member of Baryonychinae due to the presence of conical teeth with flutes and denticles in a dentary rosette. The specimen ML1190 shares the following characteristics with Baryonyx walkeri: enamel surface with small (nearly vertical) wrinkles, variable denticle size along the carinae, 6–7 denticles per mm, wrinkles forming a 45 degree angle near the carinae, and tooth root longer than crown. In addition, dubious taxa based on teeth morphology such as Suchosaurus cultridens (Owen, 1840–1845), and Suchosaurus girardi (Sauvage 1897–98; Antunes & Mateus 2003) are discussed, based on comparisons with well-known material such as Baryonyx walkeri Charig & Milner, 1986. Suchosaurus cultridens and S. girardi are considered as nomina dubia due to the lack of diagnostic apomorphies, but both specimens are referred to Baryonychinae incertae sedis.

Mateus, O., Araújo R., Natário C., & Castanhinha R. (2011).  A new specimen of the theropod dinosaur Baryonyx from the early Cretaceous of Portugal and taxonomic validity of Suchosaurus. Zootaxa. 54-68., Number 2827 Abstract
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Mateus, O. (2009).  New specimens of Angolasaurus bocagei and comments on the early radiations of plioplatecarpine mosasaurs. Journal of Vertebrate Paleontology. Journal of Vertebrate Paleontology. 29, 165A., 1, Number 3: Taylor & Francis Abstract

NEW SPECIMENS OF ANGOLASAURUS BOCAGEI AND COMMENTS ON THE EARLY RADIATIONS OF PLIOPLATECARPINE MOSASAURS POLCYN, Michael, SMU, Dallas, TX, USA; JACOBS, Louis, SMU, Dallas, TX, USA; MATEUS, Octávio, Museu da Lourinhã, Lourinhã, Portugal; SCHULP, Anne, Natuurhistorisch Museum Maastricht, Maastricht, Netherlands New, well preserved material of the Turonian mosasaur Angolasaurus bocagei from the Tadi Beds of the Itombe Formation in northern Angola, allows detailed redescription of its morphology and reassessment of its phylogenetic relationships. Angolasaurus had been previously referred to the genus Platecarpus; however, phylogenetic analysis confirms the valid taxonomic status of A. bocagei, and reconstructs that taxon within a clade that also includes the genera Selmasaurus and Ectenosaurus. These forms are united by an elaborated infrastapedial process of the quadrate and a unique ridge-like descending process of the parietal forming the supraoccipital articulation, but also retain a relatively plesiomorphic configuration of the braincase. That clade is united with all other plioplatecarpines by a number of derived characters including the presence of a novel basicranial circulation pattern. In Africa, North and South America, early plioplatecarpines are known by the Middle Turonian and Angolasaurus and closely related forms appear by the Upper Turonian. Selmasaurus and Ectenosaurus are a rare faunal component of the Santonian and Campanian of North America. Platecarpus planifrons appears in the Coniacian of North America and represents the plesiomorphic condition of the clade containing the remaining species of Platecarpus and Plioplatecarpus, that appears in the Santonian and persist until the end of the Cretaceous, reaching global distribution. The temporal and geographic distribution of these radiations suggest influence of paleogeography and eustatic sea levels.

Polcyn, M., Jacobs L., Mateus O., & Schulp A. (2009).  New specimens of Angolasaurus bocagei and comments on the early radiations of plioplatecarpine mosasaurs. Journal of Vertebrate Paleontology. Journal of Vertebrate Paleontology. 29, 165., Number 3 Abstract
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Polcyn, M. J., Jacobs L. L., Mateus O., & Schulp A. S. (2009).  New specimens of Angolasaurus bocagei and comments on the early radiations of plioplatecarpine mosasaurs. Journal of Vertebrate Paleontology. Journal of Vertebrate Paleontology. 29, 165–165., Number 3 Abstract
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Mateus, O., & Estraviz-López D. (2022).  A new theropod dinosaur from the early cretaceous (Barremian) of Cabo Espichel, Portugal: Implications for spinosaurid evolution. PLOS ONE. 17, 1-48., 02, Number 2: Public Library of Science Abstractjournal.pone_.0262614.pdfWebsite

Spinosaurids are some of the most enigmatic Mesozoic theropod dinosaurs due to their unique adaptations to aquatic environments and their relative scarcity. Their taxonomy has proven to be especially problematic. Recent discoveries from Western Europe in general, specifically Iberia, provide some of the best specimens for the understanding of their phylogeny, leading to the description of the spinosaurid Vallibonavenatrix cani and the recognition of the Iberian dinosaur Camarillasaurus cirugedae as one of them. Portuguese associated spinosaurid remains (ML1190) from the Papo Seco Formation (early Barremian) were previously assigned to Baryonyx walkeri but new material recovered in 2020 along with new phylogenetic analyses suggests a different phylogenetic placement, making their revision necessary. Here we show that these remains are not attributable to Baryonyx walkeri, but to a new genus and species, Iberospinus natarioi, gen. et sp. nov. The new taxon is characterized by the presence of a single Meckelian foramen in the Meckelian sulcus, a straight profile of the ventral surface of the dentary and a distal thickening of the acromion process of the pubis between other characters. Iberospinus natarioi is recovered as a sister taxon of the clade formed by Baryonyx and Suchomimus, and outside Spinosaurinae when Vallibonaventrix cani is excluded from the analysis. The description of this taxon reinforces Iberia as a hotspot for spinosaur biodiversity, with several endemic taxa for the region. As expected for the clade, the dentary displays a highly vascularized neurovascular network. The morphometric analysis of parts of the skeleton (pedal phalanx and caudal vertebrae, among others) shows an intermediate condition between basal tetanurans and spinosaurines.

Clemmensen, L. B., Lindström S., Mateus O., Mau M., Milàn J., & Kent D. V. (2021).  A new vertebrate fossil-bearing layer in the Rhætelv Formation (Kap Stewart Group) of central East Greenland: evidence of a Hettangian marine incursion into the continental Jameson Land Basin. Lethaia. n/a, , Number n/a Abstractlet.12449.pdfWebsite

The Kap Stewart Group (Rhaetian-Sinemurian, Triassic–Early Jurassic) of the Jameson Land Basin in central East Greenland has traditionally been regarded as a strictly continental unit with delta and perennial lake sediments. New finds of plesiosaur bone remain in a thin storm deposited sandstone bed in the middle part of the Rhætelv Formation of the Kap Stewart Group, however, indicates a likely period of marine influence. At the study area at the eastern margin of the basin, the Rhætelv Formation is 300-m thick and overlies unconformably the Norian Fleming Fjord Group. The bone-bearing sandstone occurs 190 m above the base of the group and is closely associated with black laminated mudstones; palynological investigation of three samples from these mudstones indicates that they are of a younger Hettangian age. The Hettangian was a relatively short stage (201.3–199.5 Ma) and elsewhere characterized by two episodes of sea-level highstands. Assuming that the marine incursion in the Jameson land Basin evidenced by the plesiosaur fossil remains took place during the youngest of these sea-level highstands, the bone-bearing bed of the Rhætelv Formation can be dated to 200 Ma and thereby gives the first numerical age constraint of this hitherto poorly dated succession.

Hendrickx, C., Aráujo R., & Mateus O. (2015).  The non-avian theropod quadrate I: Standardized terminology with an overview of the anatomy and function. PeerJ. 2015, , Number 9 Abstract
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Hendrickx, C., Araújo R., & Mateus O. (2014).  The nonavian theropod quadrate II: systematic usefulness, major trends and cladistic and phylogenetic morphometrics analyses. PeerJ PrePrints. 2, e380v2., 2014 AbstractWebsite

The skull-bone quadrate in nonavian theropods is very diverse morphologically alongside the disparity of the group as a whole. However this disparity has been underestimated for taxonomic purposes. In order to evaluate the phylogenetic potential and investigate the evolutionary transformations of the quadrate, we conducted a Catalano-Goloboff phylogenetic morphometric analysis as well as a cladistic analysis using 98 discrete quadrate related characters. The cladistic analysis provides a fully resolved tree mirroring to some degree the classification of nonavian theropods. The quadrate morphology by its own provides a wealth of data with strong phylogenetic signal and allows inference of major trends in the evolution of this bone. Important synapomorphies include: for Abelisauroidea, a lateral ramus extending to the ectocondyle; for Tetanurae, the absence of the lateral process; for Spinosauridae, a medial curvature of the ventral part of the pterygoid ramus occurring just above the mandibular articulation; for Avetheropoda, an anterior margin of the pterygoid flange formed by a roughly parabolic margin; and for Tyrannosauroidea, a semi-oval pterygoid flange shape in medial view. The Catalano-Goloboff phylogenetic morphometric analysis reveals two main morphotypes of the mandibular articulation of the quadrate linked to function. The first morphotype, characterized by an anteroposteriorly broad mandibular articulation with two ovoid/subcircular condyles roughly subequal in size, is found in Ceratosauria, Tyrannosauroidea and Oviraptorosauria. This morphotype allows a very weak displacement of the mandible laterally. The second morphotype is characterized by an elongate and anteroposteriorly narrow mandibular articulation and a long and parabolic/sigmoid ectocondyle. Present in Megalosauroidea, Carcharodontosauridae and Dromaeosauridae, this morphotype permits the lower jaw rami to be displaced laterally when the mouth opened.

Hendrickx, C., Araújo R., & Mateus O. (2012).  The nonavian theropod quadrate: systematics usefulness, major trends and phylogenetic morphometrics analysis. : Journal of Vertebrate Paleontology, Program and Abstracts, 2012, p.110. ISSN 1937-2809 Abstracthendrickx_araujo__mateus_2012_quadrate_theropods_svp_2012_abstract_book_meeting_abstracts.pdf.pdf

The quadrate in nonavian theropods is incredibly diverse morphologically; however this morphological disparity has been underestimated for taxonomic purposes. The quadrate topological homologies and anatomy, as well as the terminology, among nonavian theropod clades are reviewed. In order to evaluate the phylogenetic potential and investigate the evolutionary transformations of the quadrate, we conducted a Catalano-Goloboff phylogenetic morphometric analysis using 3 morphometric characters, a total of 28 landmarks coded for 23 taxa, as well as a cladistic analysis using 115 discrete quadrate-related characters coded for 43 taxa. The cladistic analysis provides a fully resolved tree mirroring the current classification of nonavian theropods. The quadrate morphology by its own provides a wealth of data with strong phylogenetic signal. Several unambiguous synapomorphies support nonavian theropod relationships and the resulting consensus tree allows inference of major trends in the evolution of this bone. Important synapomorphies include: for Abelisauridae, a lateral ramus extending to the ectocondyle; for Tetanurae, the absence of the lateral process; for Spinosauridae, a medial curvature of the ventral part of the pterygoid ramus occurring just above the mandibular articulation; for Neotetanurae, an anterior margin of the pterygoid flange formed by a roughly parabolic margin; and for Tyrannosauroidea, a semi-oval pterygoid flange shape in medial view. The Catalano-Goloboff phylogenetic morphometric analysis reveals two main morphotypes of the mandibular articulation of the quadrate linked to function. The first morphotype, characterized by an anteroposteriorly broad mandibular articulation with two ovoid/subcircular condyles roughly subequal in size, is found in Ceratosauria, Tyrannosauroidea and Oviraptorosauria. This morphotype allows a very weak displacement of the mandible laterally. The second morphotype is characterized by an elongate and anteroposteriorly narrow mandibular articulation and a long and parabolic/sigmoid ectocondyle. Present in Megalosauroidea, Allosauroidea and Dromaeosauridae, this morphotype permits the lower jaw rami to be displaced laterally when the mouth opened.

Polcyn, M. J., Jacobs L. L., Schulp A. S., & Mateus O. (2010).  The North African Mosasaur Globidens phosphaticus from the Maastrichtian of Angola. Historical Biology. 22, 175-185., Number 1 Abstract
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Mateus, O. (2007).  Notes and review of the ornithischian dinosaurs of Portugal. Journal of Vertebrate Paleontology. 27, 114A-114A., Jan: Society of Vertebrate Paleontology Abstractmateus_2007_notes_and_review_of_ornithichians_of_portugall.pdf

The record of ornithischian dinosaurs from Portugal is substantial but incomplete in terms of our understanding of taxonomic composition and details of the anatomy of many forms. New data and reinterpretation of these forms are provided. The basal thyreophoran from the Lower Jurassic (the nomen dubium “Lusitanosaurus liasicus”) is the most primitive dinosaur from Iberia. Concerning the Late Jurassic, new material from the Lourinhã Formation (Kimmeridgian/Tithonian) referable to the stegosaur Dacentrurus provides additional information on the systematic position and osteology.
The new material shows two rows of paired triangular plates, with a notch in the anterior rim. A maxillary of an ankylosaur from the Vale Frades (Lourinhã Fm.) is here reported, but cannot be referable to Dracopelta, the only ankylosaur genus currently known from Portugal. The ornithopod Alocodon kuehnei reported as Middle Jurassic (Callovian) is probably Oxfordian in age. A right dentary (ML768 from Zimbral) from the Lourinhã Formation, Kimmeridgian/Tithonian, and shares affinities with Dryosaurus but possesses more denticulation and no secondary ridges, suggesting the occurrence of a new or unreported species for the Late Jurassic of Portugal, which is here tentatively ascribed to aff. Dryosaurus sp. In summary, the Late Jurassic ornithischians species/genera from Portugal include Dacentrurus armatus, Stegosaurus cf. ungulatus, Dracopelta zbyszewskii, Phyllodon henkeli, Hypsilophodon sp., Alocodon kuehnei, Trimucrodon cuneatus, aff. Dryosaurus and Draconyx loureiroi. The Lower Cretaceous (Hauterivian / Barremian) of Praia das Aguncheiras, in Cabo Espichel, provided a partial left maxillary (CPGP.1.99.7) of a basal iguanodontian that shows affinities with Camptosaurus, and is tentatively assigned to this genus. The maxillary teeth denticles differ from Iguanodon or other Iguanodontoidea because not show mammillations. More material is necessary to validate but, to be true, that would confirm the presence of this genus in the lower Cretaceous. The Iguanodon has been well reported in the Lower Cretaceous of Cabo Espichel.

Mateus, O. (2007).  Notes and review of the ornithischian dinosaurs of Portugal. Journal of Vertebrate Paleontology. 27(suppl. to 3), 114. Abstract
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Burigo, A., & Mateus O. (2024).  Notes on the type specimens of Allosaurus atrox and A. Ferox (Theropoda, Allosauridae). Paleolusitana. 2, 18-20. Abstractaburigo__omateus_v2.pdf

Allosaurus Marsh, 1877 is one of the most iconic theropods of the Late Jurassic of USA and Portugal. However,
since the first description the genus was used as taxonomical wastebasket to which several species were
described. We revisit three type specimens, Allosaurus fragilis YPM 1930, A. atrox YPM 1890 and A. ferox USNM
2315. We interpret the vertebral material of A. atrox and support it as a junior synonym of A. fragilis, and A. ferox
as Allosaurus indet. rather than A. fragilis.

Mateus, O., & Antunes M. T. (2002).  Novos vestígios de dinossauros saurópodes do Jurássico Superior da Lourinhã (Portugal). Congresso Ibérico de Herpetologia. , Évora Abstract
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Mateus, O., & Antunes M. T. (2002).  Novos vestígios de dinossauros saurópodes do Jurássico Superior da Lourinhã (Portugal). Congresso Ibérico de Herpetologia. , Évora Abstract
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Mateus, O., & Antunes M. T. (2002).  Novos vestígios de dinossauros saurópodes do Jurássico Superior da Lourinhã (Portugal).. Congresso Ibérico de Herpetologia. , Évora, Portugal Abstract

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