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Mateus, O., Neto de Carvalho C., & Klompmaker A. A. (2013).  Decapod crustacean body and ichnofossils from the Mesozoic of Portugal. 5th Symposium on Mesozoic and Cenozoic Decapod Crustaceans. , 25–27 June 2013, Warszawa: Polish Geological Institute − National Research Institute & AGH University of Science and Technologymateus_et_al_2013_crustacea_mesozoic_portugal_5th_decapod_crustaceans_meeting_2013.pdf
Mateus, O. (2013).  Decapod crustacean body and ichnofossils from the Mesozoic of Portugal. NA, , 1 Abstract

Book of abstracts of the 5th Symposium on Mesozoic and Decapod Crustaceans

Leal, A. S., Mateus O., Tomás C., & Dionísio A. (2014).  Decay and conservation trial of Late Jurassic sandstone with dinosaur tracks in a museum environment (Museum of Lourinhã, Portugal). Buletini i Shkencave Gjeologjike. 1(2014), 410. Abstractleal_et_al_2014_cbgassav1-_abstract_dinosaur_footprints__page_410.pdf

Abstract
Late Jurassic dinosaur footprints were found on a coastline cliff in Lourinhã, Porto das Barcas, Lagido do Forno (coordinate 39°14.178’N, 9°20.397’W, Portugal) in June 2001. The locality is characterized by steep cliffs with high slopes that are composed of gray and red sandstones/ siltstones. The location belongs to the successions of Lusitanian Basin representing the Porto Novo Member of the Lourinhã Formation. Three natural infills of tridactyl tracks, possibly ascribed to ornithopod, a bipedal herbivore were found, representing a left foot movement, a right and a left one, respectively. Footprints are 300- 400mm wide and have a height of 330-360mm. The footprints are characterized by round fingers, which are elongated due to some degradation/ erosion. The footprints were collected from the field in 2001 and subsequently cleaned, consolidated and glued in the laboratory of the Museum of Lourinhã before being exhibited in a museum display. Stone matrix was removed and a consolidation product was applied, probably a polyvinyl acetate. The footprint with broken central digit was glued with an epoxy resin, Araldite. Both applied products were confirmed by analysis of μ- FTIR and both presented colour change and detachment surface problems. The footprints have been exposed in the palaeontology hall of the Museum of Lourinhã, Portugal from 2004 without climate controlling. These trace fossils form an important part of the palaeontological collection of Late Jurassic vertebrate fossils from Lourinhã Formation. Presently, it is considered a unique heritage in danger of disappearing due to high decay level of disaggregation of its geological structure. The footprints display several pathologies, such as “Blistering”, “Powdering”, “Exfoliation”’ as well as “Dirt”, “Fracture”’, “Inscriptions”, “Consolidants” and “Adhesives” and are now in very poor conditions. Laboratorial analysed were made to evaluate the presence of salts. Moreover a microclimatic study was conducted inside the museum to evaluate the influence of thermo-hygrometric parameters on the decay processes. The future interventions will depend on the results of consolidation trials that are currently under progress by using stone samples taken from the same layer and location from Porto das Barcas applying different commercial consolidation products.

Jacobs, L. L., Polcyn M. J., Mateus O., & Schulp A. S. (2023).  Deep time conservation paleobiology of the Atlantic jigsaw puzzle and the future of the southwestern Angolan coast. Bulletin of the Florida Museum of Natural History. 60(2), 90.: In: Abstracts of the 2nd Conservation Paleobiology Symposium. https://doi … Abstractjacobs_et_al_2023_jigsaw.pdf

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Hayashi, S., Carpenter K., Watabe M., Mateus O., & Barsbold R. (2008).  Defensive weapons of thyreophoran dinosaurs: histological comparisons and structural differences in spikes and clubs of ankylosaurs and stegosaurs. Journal of Vertebrate Paleontology. 28(3, Supplement), 89A-90A., Number Suppl. to 3 Abstracthayashi_et_al_2008_histology_stegosaurs_defensive_weapons_of_thyreophoran_dinosaurs-_histological_comparisons_and_structural_differences_in_spikes_and_clubs_of_ankylosaurs_and_stegosaurs.pdfWebsite

Thyreophoran dinosaurs have spike- and club-shaped osteoderms probably used for defensive weapons. The structural and histological variations have been little known. Here, we provide the comparisons of the internal structures in defensive weapons of ankylosaurs and stegosaurs, using spikes of a polacanthid (Gastonia) and a nodosaurid (Edmontonia), clubs of ankylosaurids (Saichania and Ankylosauridae indet. from Canada), and spikes of stegosaurids (Stegosaurus and Dacentrurus), which sheds light on understandings of evolutionary history and functional implications of defensive weapons in thyreophorans. In ankylosaurs, the structural and histological features of spikes and clubs are similar with those of small osteoderms in having thin compact bones, thick cancellous bones with large vascular canals, and abundant collagen fibers. A previous study demonstrated that each of three groups of ankylosaurs (polacanthid, nodosaurid, and ankylosaurid) has distinct characteristic arrangements of collagen fibers in small osteoderms. This study shows that spikes and clubs of ankylosaurs maintain the same characteristic features for each group despite of the differences in shapes and sizes. Conversely, the spike-shaped osteoderms in primitive (Dacentrurus) and derived (Stegosaurus) stegosaurids have similar structure to each other and are significantly different from the other types of stegosaur osteoderms (throat bony ossicles and plates) in having thick compact bones with a medullary cavity. These lack abundant collagen fibers unlike ankylosaur osteoderms. The spikes of ankylosaurs and stegosaurs are similar in shape, but their structural and histological features are different in having unique structures of collagen fibers for ankylosaurs and thick compact bones for stegosaurs, providing enough strength to have large spikes and to use them as defensive weapons. Although the shapes of ankylosaur clubs are different from spikes, the internal structures are similar, suggesting that ankylosaurs maintain similar structures despite of different shapes in osteoderms. These results indicate that ankylosaurs and stegosaurs used different strategies independently to evolve defensive weapons.

Hayashi, S., Carpenter K., Watabe M., Mateus O., & Barsbold R. (2008).  Defensive weapons of thyreophoran dinosaurs: histological comparisons and structural differences in spikes and clubs of ankylosaurs and stegosaurs. 28 (3, Supplement), 89A-90A. Journal of Vertebrate Paleontology. 28, 89-90., Number Suppl. to 3 Abstract
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Hayashi, S., Carpenter K., Watabe M., Mateus O., & Barsbold R. (2008).  Defensive weapons of thyreophoran dinosaurs: histological comparisons and structural differences in spikes and clubs of ankylosaurs and stegosaurs. 28 (3, Supplement), 89A-90A. Journal of Vertebrate Paleontology. 28, 89–90., Number Suppl. to Abstract
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Leal, S., Mateus O., Tomás C., & Dionisio A. (2014).  Degradation processes and consolidation of Late Jurassic sandstone dinosaur tracks in museum environment (Museum of Lourinhã, Portugal). EGU General Assembly 2014 - Geophysical Research Abstracts. Vol. 16, EGU2014-9026-1, 2014.leal_et_al_2014_tracks_lab_egu2014-9026-1.pdf
Mateus, O. (2014).  Degradation processes and consolidation of Late Jurassic sandstone dinosaur tracks in museum environment (Museum of Lourinhã, Portugal). Geophysical Research Abstracts. Geophysical Research Abstracts, EGU2014–9026–1, 2014., 1 Abstract
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Hendrickx, C., Mateus O., & Araújo R. (2015).  The dentition of megalosaurid theropods. Acta Palaeontologica Polonica. 60(3), 627–642. Abstracthendrickx_et_al_2015_theropod_teeth_app.pdfWebsite

Theropod teeth are particularly abundant in the fossil record and frequently reported in the literature. Yet, the dentition of many theropods has not been described comprehensively, omitting details on the denticle shape, crown ornamentation and enamel texture. This paucity of information has been particularly striking in basal clades, thus making identification of isolated teeth difficult, and taxonomic assignments uncertain. We here provide a detailed description of the dentition of Megalosauridae, and a comparison to and distinction from superficially similar teeth of all major theropod clades. Megalosaurid dinosaurs are characterized by a mesial carina facing mesiolabially in most mesial teeth, centrally positioned carinae on both most mesial and lateral crowns, a mesial carina terminating above the cervix, and short to well-developed interdenticular sulci between distal denticles. A discriminant analysis performed on a dataset of numerical data collected on the teeth of 62 theropod taxa reveals that megalosaurid teeth are hardly distinguishable from other theropod clades with ziphodont dentition. This study highlights the importance of detailing anatomical descriptions and providing additional morphometric data on teeth with the purpose of helping to identify isolated theropod teeth in the future.

Hendrickx, C., Mateus O., & Araújo R. (2014).  The dentition of megalosaurid theropods, with a proposed terminology on theropod teeth. XII EAVP Meeting XII Annual Meeting of the European Association of Vertebrate Palaeontologists – Abstract Book. p. 75., Torino 24-28 June 2014hendrickx_et_al_2014_megalosaurid_teeth_eavp.pdf
Hendrickx, C., Mateus O., & Araújo R. (2014).  The dentition of Megalosauridae (Theropoda: Dinosauria). {APP}. : Polska Akademia Nauk Instytut Paleobiologii (Institute of Paleobiology, Polish Academy of Sciences) AbstractWebsite
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Mateus, O. (2009).  Dinolourinhã – a integração dos jovens na paleontologia: o caso-estudo do Museu da Lourinhã.. Journal of Paleontological Techniques. 28–29., 1 Abstract
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Tomas, C., Mateus O., & Abreu C. (2009).  Dinolourinhã – a integração dos jovens na paleontologia: o caso-estudo do Museu da Lourinhã.. Journal of Paleontological Techniques 5: 28-29.. Abstract
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Tomas, C., Mateus O., & Abreu C. (2009).  Dinolourinhã; a integração dos jovens na paleontologia: o caso-estudo do Museu da Lourinhã. Journal of Paleontological Techniques 5: 28-29.. 28-29., Jan Abstracttomas_et_al_2009_dinolourinha_abstracts_jpt.pdf

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Martins, R. M. S., Beckmann F., Castanhinha R., Mateus O., & Pranzas P. K. (2011).  Dinosaur and crocodile fossils from the mesozoic of Portugal: Neutron tomography and synchrotron-radiation based micro-computed tomography. Materials Research Society Symposium Proceedings. 1319, 319-332. Abstract
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Mateus, O., Jacobs L. L., Polcyn M. J., Schulp A. S., Neto A. B., & Antunes M. T. (2008).  Dinosaur and turtles from the Turonian of Iembe, Angola. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontologia de Vertebrados. 156., Neuquén, Argentina Abstractmateus_et_al_2008_dinosaur_and_turtles_from_the_turonian_of_iembe_angola.pdf

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Mateus, O., Jacobs L. L., Polcyn M. J., Schulp A. S., Neto A. B., & Antunes M. T. (2008).  Dinosaur and turtles from the Turonian of Iembe, Angola. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 156., Neuquén, Argentina Abstract
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Mateus, O., Jacobs L. L., Polcyn M. J., Schulp A. S., Neto A. B., & Antunes M. T. (2008).  Dinosaur and turtles from the Turonian of Iembe, Angola. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 156–156., Neuquén, Argentina Abstract
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Castanhinha, R., Araújo R., & Mateus O. (2009).  Dinosaur eggshell and embryo localities in Lourinhã Formation, Late Jurassic, Portugal. Journal of Vertebrate Paleontology, 29(3): . 76A. Abstractcastanhinhaetal2009dinosaureggshellp.pdf

Four different localities from the Late Jurassic of Lourinhã formation with eggshells and embryos were studied: Paimogo (lower Amoreira-Porto Novo member), Peralta (Praia Azul member), Porto das barcas (Bombarral member) and Casal da Rôla (Amoreira-Porto Novo member). All but Casal da Rôla have embryonic material. Preliminary results show that eggshells from Paimogo correspond to obliquiprismatic morphotype (0.92mm thick), similar to those from Morrison Formation. Within Paimogo site a different type of eggshell was discovered, having a radial section of 153 μm with a mammilary layer measuring 65 μm. Porto das Barcas eggshells represent a discretispherulitic morphotype (1,23 mm thick).
This locality presents a nest 60-cm diameter containing many eggshells but an indeterminate number of eggs. Some embryonic bones were discovered between the eggshells including teeth and skull bones showing that the eggs belong to a saurischian, tentatively a sauropod dinosaur. Peralta nest eggshells are preliminary ascribed to obliquiprismatic morphotype (column: 0,56mm and mammilla: 0,21mm) probably related to Paimogo’s nest taxon (Lourinhanosaurus). Peralta site bears embryonic bones namely small theropod teeth associated with bone fragments, and unidentifiable dinosaur vertebra. Only eggshells have been collected at Casal da Rôla (ML1194). The eggshells (0,78mm thick) are prismatic morphotype and it was impossible to determine the pore system, the outer surface is smooth with no ornamentation.
Lourinhã formation has the oldest sauropod and theropod nest with embryos known so far.

Mateus, O. (2009).  DINOSAUR EGGSHELL AND EMBRYO LOCALITIES IN LOURINHA FORMATION, LATE JURASSIC, PORTUGAL. Journal of Vertebrate Paleontology. 29, 76A–76A., 1 Abstract
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Castanhinha, R., Araujo R., & Mateus O. (2009).  Dinosaur eggshell and embryo localities in Lourinhã Formation, Late Jurassic, Portugal. Journal of Vertebrate Paleontology. 29, 76., Number 3 Abstract
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Castanhinha, R., Araujo R., & Mateus O. (2009).  Dinosaur eggshell and embryo localities in Lourinhã Formation, Late Jurassic, Portugal. Journal of Vertebrate Paleontology. 29, 76–76., Number 3 Abstract
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Mateus, O., Overbeeke M., & Rita F. (2008).  Dinosaur Frauds, Hoaxes and "Frankensteins": How to distinguish fake and genuine vertebrate fossils. Journal of Paleontological Techniques. 2, 1-5.. Abstractmateus_et_al_2008_dinosaur_frauds_hoaxes_and_frankensteins-_how_to_distinguish_fake_and_genuine_vertebrate_fossils._journal_of_paleontological_techniques.pdfWebsite

Dinosaurs and other fossils have been artificially enhanced, or totally forged, to increase their commercial value. The most problematic forgeries to detect are based on original fossils that are artificially assembled. Several techniques are suggested for detecting hoaxes: detailed visual examination, chemical analysis, Xray or CT-scan, and ultraviolet light. It is recommended that museums and paleontological researchers do not purchase and/or trade fossils lacking clear provenience information. Exceptions to that general rule should be closely examined using techniques described herein.

Mateus, O., Overbeeke M., & Rita F. (2008).  Dinosaur Frauds, Hoaxes and "Frankensteins": How to distinguish fake and genuine vertebrate fossils. Journal of Paleontological Techniques. 2, 1-5. Abstract
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Mateus, O., Antunes M. T., & Taquet P. (2001).  Dinosaur ontogeny : the case of Lourinhanosaurus (Late Jurassic, Portugal). J. Vertebr. Paleontol. 21, Abstract
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Mateus, O., & Marzola M. (2014).  Dinosaur taphonomy in the Lourinhã Formation (Late Jurassic, Portugal). 7th International Meeting on Taphonomy and Fossilization, Taphos 2014. 60-61., Ferrara, Italymateus__marzola_2014_lourinha_taphonomy_ferrara_taphonomy_meeting_2014.pdf
Mateus, O. (2014).  Dinosaur taphonomy in the Lourinhã Formation (Late Jurassic, Portugal). International Meeting on Taphonomy and Fossilization. 60–61. Abstract
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Marques, M. I. F., & Mateus O. (2021).  Dinosaur tracksites from Portugal, focused on the carbonated platform of North and Central Lusitanian Basin. 3rd Palaeontological Virtual Congress. 210.: ISBN 978-84-09-36657-6 Abstractmarques_mateus_2021_pvc3_tracks.pdf

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Milàn, J., & Mateus O. (2024).  Dinosaurfund fra Jylland. Magasinet Naturen. 2024(1), 52-55.milan__mateus_2024_-_jydske_dinosaurer.pdfWebsite
Mateus, O., & Andersen E. (1998).  Dinosaurrede i Gedser- portugisisk specialitet udstilles i Gedser. GeologiskNyt. 3/98, 7. Abstract
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Antunes, M. T., & Mateus O. (2003).  Dinosaurs of Portugal. Comptes Rendus Palevol. 2, 77-95., Number 1 Abstractantunes_mateus_2003_dinosaurs_of_portugal.pdfWebsite

A synthesis on the state of art on dinosaur knowledge in Portugal is presented. The following genera have been recognized: Ceratosaurus, Torvosaurus, Lourinhanosaurus, Allosaurus, cf. Compsognathus, Stokesosaurus, cf. Richardoestesia, cf. Archaeopteryx, Euronychodon, cf. Paronychodon, Dinheirosaurus, Lourinhasaurus, Lusotitan, cf. Pleurocoelus, Lusitanosaurus, Dacentrurus, Dracopelta, Phyllodon, Hypsilophodon, Alocodon, Trimucrodon, Draconyx, Iguanodon, and Taveirosaurus. Most are from Late Jurassic localities at the Lourinhã area and Guimarota. A new genus, Lusotitan, is here raised to include the Late Jurassic ‘Brachiosaurus’ atalaiensis. Lower Cretaceous until Cenomanian material is scarce, except for dinosaur footprints. An interesting Late-Cretaceous, mostly small dinosaur association has been collected between Aveiro and Taveiro.

Antunes, M. T., & Mateus O. (2003).  Dinosaurs of Portugal. Comptes Rendus Palevol. 2, 77–95., jan, Number 1: Elsevier {BV} AbstractWebsite
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Mateus, O. (1998).  Dinossauros de Portugal e um novo terópode do Jurássico Superior da Lourinhã. , Évora: Universidade de Évora Abstract
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Mateus, O. (2005).  Dinossauros do Jurássico Superior de Portugal, com destaque para os saurísquios. Universidade Nova de Lisboa. , Lisboa
Mateus, O. (1998).  Dinossauros Portugueses. Caderno de resumos do I Congresso de Estudantes de Biologia. 13., Évora Abstractmateus_1998_dinossauros_portugueses_i_congresso_estudantes_biologia.pdf.pdf

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Mateus, O. (1998).  Dinossauros Portugueses. Caderno de resumos do I Congresso de Estudantes de Biologia. 13., Évora Abstract
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Mateus, O. (1998).  Dinossauros Portugueses. Caderno de resumos do I Congresso de Estudantes de Biologia. 13–13., Évora Abstract
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Tschopp, E., & Mateus O. (2016).  Diplodocus Marsh, 1878 (Dinosauria, Sauropoda): proposed designation of D. carnegii Hatcher, 1901 as the type species. Bulletin of Zoological Nomenclature. 73(1), 17-24. Abstracttschopp_mateus_2016_-_case_3700_-_diplodocus_type.pdf

The purpose of this application, under Articles 78.1 and 81.1 of the Code, is to replace Diplodocus longus Marsh, 1878 as the type species of the sauropod dinosaur genus Diplodocus by the much better represented D. carnegii Hatcher, 1901, due to the undiagnosable state of the holotype of D. longus (YPM 1920, a partial tail and a chevron). The holotype of D. carnegii, CM 84, is a well-preserved and mostly articulated specimen. Casts of it are on display in various museums around the world, and the species has generally been used as the main reference for studies of comparative anatomy or phylogeny of the genus. Both species are known from the Upper Jurassic Morrison Formation of the western United States. The genus Diplodocus is the basis for the family-level taxa diplodocinae Marsh, 1884, diplodocidae Marsh, 1884, diplodocimorpha Marsh, 1884 (Calvo & Salgado, 1995) and diplodocoidea Marsh, 1884 (Upchurch, 1995). It is also a specifier of at least 10 phylogenetic clades. With the replacement of D. longus by D. carnegii as type species, Diplodocus could be preserved as a taxonomic name with generally accepted content. Taxonomic stability of the entire clade diplodocoidea, and the proposed definitions of several clades within Sauropoda, could be maintained.

Tschopp, E., & Mateus O. (2016).  Diplodocus Marsh, 1878 (Dinosauria, Sauropoda): proposed designation of D. carnegii Hatcher, 1901 as the type species. Bulletin of Zoological Nomenclature. 73, 17-24. Abstract
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Hendrickx, C., Mateus O., Araújo R., & Choiniere J. (2019).  The distribution of dental features in non-avian theropod dinosaurs: Taxonomic potential, degree of homoplasy, and major evolutionary trends. Palaeontologia Electronica. 22(3), 1-110. Abstractthe_distribution_of_dental_features_in_non-avian_t.pdfWebsite

Isolated theropod teeth are some of the most common fossils in the dinosaur fossil record and are continually reported in the literature. Recently developed quantitative methods have improved our ability to test the affinities of isolated teeth in a repeatable framework. But in most studies, teeth are diagnosed on qualitative characters. This can be problematic because the distribution of theropod dental characters is still poorly documented, and often restricted to one lineage. To help in the identification of isolated theropod teeth, and to more rigorously evaluate their taxonomic and phylogenetic potential, we evaluated dental features in two ways. We first analyzed the distribution of 34 qualitative dental characters in a broad sample of taxa. Functional properties for each dental feature were included to assess how functional similarity generates homoplasy. We then compiled a quantitative data matrix of 145 dental characters for 97 saurischian taxa. The latter was used to assess the degree of homoplasy of qualitative dental characters, address longstanding questions on the taxonomic and biostratigraphic value of theropod teeth, and explore the major evolutionary trends in the theropod dentition.

In smaller phylogenetic datasets for Theropoda, dental characters exhibit higher levels of homoplasy than non-dental characters, yet they still provide useful grouping information and optimize as local synapomorphies of smaller clades. In broader phylogenetic datasets, the degree of homoplasy displayed by dental and non-dental characters is not significantly different. Dental features on crown ornamentations, enamel texture and tooth microstructure have significantly less homoplasy than other dental features and can be used to identify many theropod taxa to ‘family’ or ‘sub-family’ level, and some taxa to genus or species. These features should, therefore, be a priority for investigations seeking to classify isolated teeth.

Our observations improve the taxonomic utility of theropod teeth and in some cases can help make isolated teeth useful as biostratigraphic markers. This proposed list of dental features in theropods should, therefore, facilitate future studies on the systematic paleontology of isolated teeth.

Hendrickx, C., Mateus O., Araújo R., & Choiniere J. (2019).  The distribution of dental features in non-avian theropod dinosaurs: Taxonomic potential, degree of homoplasy, and major evolutionary trends. Palaeontologia Electronica. 22, , Number 3 Abstract
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