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Mateus, O. (2009).  The Cretaceous Skeleton Coast of Angola. 29, , 1 Abstract
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Mateus, O. (2013).  Cathetosaurus as a valid sauropod genus and comparisons with Camarasaurus. Journal of Vertebrate Paleontology. 173., 1 Abstract
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Mateus, O. (2016).  Exemplos bizarros de evolução em dinossauros e alguns casos portugueses. Do Big Bang ao Homem. 81-95., Porto: U.Porto Ediçõesmateus_2016_capitulo_livro_dinosaurs.pdf
Mateus, O., Jacobs L. L., Polcyn M. J., Schulp A. S., Neto A. B., & Antunes M. T. (2008).  Dinosaur and turtles from the Turonian of Iembe, Angola. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 156–156., Neuquén, Argentina Abstract
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Mateus, O. (2011).  New fossil whales from Angola. Abstracts of the 71st Annual Meeting of the Society of Vertebrate Paleontology. 119., 1 Abstract
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Mateus, O., Morais M. L., Schulp A. S., Jacobs L. L., & Polcyn M. J. (2006).  The Cretaceous of Angola. Journal of Vertebrate Paleontology. 26, 96-97., Number (Suppl. To 3) Abstract
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Mateus, O., Callapez P. M., Polcyn M. J., Schulp A. S., Gonçalves A. O., & Jacobs L. L. (2019).  The Fossil Record of Biodiversity in Angola Through Time: A Paleontological Perspective. (Huntley, Brian J., Russo, Vladimir, Lages, Fernanda, Ferrand, Nuno, Ed.).Biodiversity of Angola: Science {&} Conservation: A Modern Synthesis. 53–76., Cham: Springer International Publishing Abstract

This chapter provides an overview of the alpha paleobiodiversity of Angola based on the available fossil record that is limited to the sedimentary rocks, ranging in age from Precambrian to the present. The geological period with the highest paleobiodiversity in the Angolan fossil record is the Cretaceous, with more than 80{%} of the total known fossil taxa, especially marine molluscs, including ammonites as a majority among them. The vertebrates represent about 15{%} of the known fauna and about one tenth of them are species firstly described based on specimens from Angola.

Mateus, O., & Antunes T. M. (2001).  Draconyx loureiroi, a new camptosauridae (Dinosauria, Ornithopoda) from the Late Jurassic of Lourinhã, Portugal. Annales de Paleontologie. 87, 61-73. Abstractmateus_antunes_2001_draconyx_loureiroi_a_new_camptosauridae_dinosauria_ornithopoda_from_the_late_jurassic_of_lourinha_portugal.pdfWebsite

A new ornithopod dinosaur is described here under the name of Draconyx loureiroi n. gen., n. sp. on teeth, caudal vertebrae, forelimb, hindlimb, and foot material that were found in association in the Late Jurassic-Tithonian of Lourinhã, Portugal. Draconyx is a Camptosauridae related to Camptosaurus.

Mateus, O. (1998).  Serão as aves dinossauros?. CiênciaJ. 6, 5. Abstract
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Mateus, O. (1996).  Situação populacional de Hemidactylus turcicus em Évora-Portugal. Actas do IV Congresso Luso-Espanhol de Herpetologia. 45–45., Porto Abstract
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Mateus, O., & Araújo R. (2008).  Stone-splitters and expansive demolition agents: breaking big rocks with little effort on vertebrate paleontological excavations. Journal of Paleontological Techniques. 3, 1-6. Abstractmateus__araujo_2008_stone-splitters_and_expansive_demolition_agents_jpt_n003_oct.pdfWebsite

Two techniques (stone-splitters and demolition agents) are revealed to be efficient methods for breaking large stone blocks in extreme paleontological excavation. In certain conditions – where security factors, permission issues, terrain conditions, rock properties are problematic – the traditional methods for breaking large rock blocks cannot be applied (e.g. crane trucks or explosives). Using an expansive demolition agent or stone-splitters after drilling equidistant holes not only allows a cheap, quick and safe solution but also permits precise removal of up to 9 ton blocks. Stone-splitters are a three-part tool that when inserted linearly and equidistantly along a brittle rock mass cause a precise fracture.

Mateus, O. (2014).  Elephas and other vertebrate fossils near Taghrout, Morocco. Journal of Vertebrate Paleontology. Program and Abstracts, 2014, 178., 1 Abstract
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Mateus, O. (2009).  Preparation techniques applied to a stegosaurian Dinosaur from Portugal. Journal of Paleontological Techniques. 5, 1–24., 1, Number NA Abstract
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Mateus, O., Mannion P. D., & Upchurch P. (2014).  Zby atlanticus, a new turiasaurian sauropod (Dinosauria, Eusauropoda) from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology. 34, 618–634., Number 3 Abstract
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Mateus, O. (2008).  Checklist for Late Jurassic reptiles and amphibians from Portugal. Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 55–55., Coimbra Abstract
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Mateus, O. (2007).  Notes and review of the ornithischian dinosaurs of Portugal. Journal of Vertebrate Paleontology. 27(suppl. to 3), 114. Abstract
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Mateus, O. (2010).  Paleontological collections of the Museum of Lourinhã (Portugal). (JM Brandão, Callapez, PM, O. Mateus, Castro, P, Ed.).Colecções e museus de Geologia: missão e gestão. 121–126., 1: Ed. Universidade de Coimbra e Centro de Estudos de História e Filosofia da Ciência Abstract
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Mateus, O., Callapez P. M., & Puértolas-Pascual E. (2017).  The oldest Crocodylia? a new eusuchian from the Late Cretaceous (Cenomanian) of Portugal. Journal of Vertebrate Paleontology, Program and Abstracts. 2017, 160. Abstract
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Mateus, O., & Antunes M. T. (2000).  Torvosaurus sp.(Dinosauria : Theropoda) in the Late Jurassic of Portugal. Livro de Resumos do I Congresso Ibérico de Paleontologia. 115-117. Abstract
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Mateus, O., & Jacinto J. J. (1997).  Ritmos de Actividade e habitat de Hemidactylus turcicus (Reptilia: Gekkonidae) em Évora, Portugal. Cuadernos de INICE. 74-75, 207-214. Abstract
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Mateus, O. (2008).  Checklist for Late Jurassic reptiles and amphibians from Portugal. Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 55., Coimbra Abstractmateus_2008_lista_de_repteis_e_anfibios_do_jurassico_superior_de_portugal__list_congressoherpetolog.pdf

The richness of Late Jurassic vertebrates in Portugal is known since the 19th century by Paul Choffat, Henri Sauvage and other. The Kimmeridgian Guimarota fauna assemblage is the best known, followed by the fauna of Lourinhã formation. Here is presented an attempt to provide a checklist of the reptiles and amphibians of the Late Jurassic. Amphibia: Lissamphibia (Celtedens, cf. Marmorerpeton, Discoglossidae indet.). Chelonia: Eucryptodira (Pleurosternidae indet., Platychelyidae indet., Plesiochelys cf. etalloni, Plesiochelys choffati, Anosteirinae indet.). Squamata: Scincomorpha (Becklesius hoffstetteri; Paramacellodus sp., Saurillodon proraformis, S. henkeli, S. cf. obtusus). Squamata: Anguimorpha (Dorsetisaurus pollicidens, Parviraptor estesi). Crown Lepidosauromorpha (Marmoretta sp.). Choristodera: Cteniogenidae (Ctenogenys reedi). Sauropterygia: Plesiosauria: Cryptoclidoidea: Cryptoclididae indet. Crocodylomorpha (Lisboasaurus estesi, L. mitrocostatus). Crocodyliformes: Neosuchia (Machimosaurus hugii, Goniopholis cf. simus, Goniopholis baryglyphaeus, cf. Bernissartia, Atoposauridae, Theriosuchus guimarotae, cf. Alligatorium, Metriorhynchus sp.). Pterosauria (Rhamphorhynchus sp., Pterodactylus sp.). Dinosauria: Theropoda (Ceratosaurus sp. , Torvosaurus sp., Lourinhanosaurus antunesi, Allosaurus europaeus, Cf. Compsognathus sp., cf. Richardoestesia sp., Dromaeosaurinae indeter., Velociraptorinae indeter., cf. Archaeopteryx sp., aff. Paronychodon). Dinosauria: Sauropoda: Eusauropoda (Dinheirosaurus lourinhanensis, Lourinhasaurus alenquerensis, Lusotitan atalaiensis, Apatosaurus sp.). Dinosauria: Ornithischia: Thyreophora (Dacentrurus armatus, Stegosaurus sp., Dracopelta zbyszewskii). Dinosauria: Ornithischia: Ornithopoda (Phyllodon henkeli, Dryosaurus sp., Hypsilophodon sp., Alocodon kuehnei, Trimucrodon cuneatus, Draconyx loureiroi).

Mateus, O. (2014).  Cracking dinosaur endothermy: paleophysiology unscrambled. NA, , 1 Abstract

The amniote eggshell functions as a respiratory structure adapted for the optimal transmission of respiratory gasses to and from the embryo according to its physiological requirements. Therefore amniotes with higher oxygen requirements, such as those that sustain higher metabolic rates, can be expected to have eggshells that can maintain a greater gas flux to and from the egg. Studies of extant amniotes have found that eggshells of reduced porosity impose a limit on the metabolic rate of the offspring. Here we show a highly significant relationship between metabolic rates and eggshell porosity in extant amniotes that predicts highly endothermic metabolic rates in dinosaurs. This study finds the eggshell porosity of extant endotherms to be significantly higher than that of extant ectotherms. Eggshell porosity values of dinosaurs are found to be significantly higherthan that of extant ectotherms, but not extant endotherms. Dinosaur eggshells are commonly preserved in the fossil record, and porosity may be readily identified and measured. This provides a simple tool to identify metabolic rates in extinct egg-laying tetrapods whose eggs possessed a mineralized shell

Mateus, O., Estraviz López D., & Madeira Mateus S. G. (2022).  Type specimens alone have a strong correlation with taxa record by geological epoch: the case study of the fossil vertebrates named from {Portuguese} types. Comunicações Geológicas. 109, 57–64. Abstractmateus-et-al_2022_type_specimens.pdfWebsite

ABSTRACT: Type specimens (holotypes, neotypes, syntypes, etc.) are of crucial importance because they are the only tangible evidence of the nomenclatural act that supports the understanding of paleobiodiversity. The list of the vertebrate species whose type specimen is based on fossils from Portugal is presented here. We counted 206 species, of which there are 45 bony fishes, 39 Late Jurassic Mammaliamorpha, 33 Cenozoic mammals, 28 non-avian dinosaurs, 25 non-dinosaur reptiles, 11 Cenozoic Aves, 22 Chondrichthyes and three other vertebrates. Except for the Quaternary fossil record, the type specimens can be used as a shortcut for measuring the fossil record and paleobiodiversity through geological time and rock units because they correlate in 95% with the fossil record by epoch in the case-study of fossil vertebrates of Portugal.

Mateus, O. (2012).  New dinosaur and pterosaur tracksites from the Late Jurassic of Portugal. , Chongqing, China: 2012 Abstract Book of Qijiang International Dinosaur Tracks Symposium Abstractmateus_2012_dinosaur_tracks_portugal__abstract_book_qijiang_int_dinosaur_tracks_symposium.pdf

Portugal is rich on dinosaur remains (bones, eggs, and tracks) from Early Jurassic to Late
Cretaceous ages, but mainly from the Late Jurassic, in which dozen of tracksites have been reported.
Here are reported new or poorly known track localities:
1) Five tracksites share the preservation substrate (marine carbonated limestone), age (late Jurassic), geographic area (Leiria district of Portugal), kind of preservation (true tracks), and completeness (trackways of multiple individuals):
i) Praia dos Salgados includes eight trackways, mostly ornithopods and theropods, and one wide gauge sauropod, made in very soft sediment; some preserve the hallux impression.
ii) Serra de Mangues is mostly covered with vegetation but seems to include dozens of tracks comprising theropods, thyreophorans, ornithopods and sauropods.
iii) Sobral da Lagoa (Pedreira do Rio Real) include six trackways but poorly preserved;
and
iv) Serra de Bouro that preserves four sauropod trackways in one single layer.
v) Pedrógão, preserved, at least, one theropod trackway and several isolated tracks of
theropods and ornithopods were found in different layers in the Early Oxfordian.
2) The locality in Praia de Porto das Barcas yielded natural casts of stegosaur tracks
(including pes print with skin impression) and a very large sauropod pes print with about
1.2 m long pes.
3) A new pterosaur tracksite was found in the Late Jurassic of Peralta, Lourinhã (Sobral Member, Lourinhã Fm.; Late Kimmeridgian/Early Tithonian). More than 220 manus and pes tracks have been collected in about five square meters, all ascribed to pterosaurs. The tracks were produced in a thin mud layer that has been covered by sand which preserved them as sandstone mould infill (natural casts). The manus of the largest specimens is 13 cm wide and 5.5 cm long and the pes measures 14.5 cm in length and 9 cm in width. This shows the occurrence of very large pterosaurs in the Late Jurassic. Other pterosaur tracksites in the Late Jurassic of Portugal are: Porto das Barcas (Lourinhã Municipality), South of Consolação (Peniche Municipality), and Zambujal de Baixo (Sesimbra Municipality).

Mateus, O. (2009).  DINOSAUR EGGSHELL AND EMBRYO LOCALITIES IN LOURINHA FORMATION, LATE JURASSIC, PORTUGAL. Journal of Vertebrate Paleontology. 29, 76A–76A., 1 Abstract
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Mateus, O., Butler R. J., Brusatte S. L., Whiteside J. H., & Steyer J. S. (2014).  The first phytosaur (Diapsida, Archosauriformes) from the Late Triassic of the Iberian Peninsula. Journal of Vertebrate Paleontology. 34, 970–975., Number 4 Abstract
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Mateus, O., Natario C., Araujo R., & Castanhinha R. (2008).  A new specimen of spinosaurid dinosaur aff. Baryonyx from the Early Cretaceous of Portugal. (Universidade de, Coimbra, Ed.).Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 51., Coimbra Abstract
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Mateus, O., Dyke G., Motchurova-Dekova N., Ivanov P., & Kamenov G. D. (2008).  The Bulgarian dinosaur: did it exist? European late Cretaceous ornithomimosaurs. 56th Symposium of Vertebrate Palaeontology and Comparative Anatomy. 47–47., Dublin Abstract
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Mateus, O. (2010).  First records of crocodyle and pterosaur tracks in the Upper Jurassic of Portugal.. New Mexico Museum of Natural History and Science Bulletin. 51, 83–87., 1, Number NA Abstract

The Upper Jurassic of Portugal has a rich vertebrate fauna well documented from both body and trace fossils. Although the occurrence of crocodyles and pterosaurs is well documented from body fossils, trace fossils from both groups were unknown until now. Here we describe an isolated crocodyle-like track from Praia da Peralta and pterosaur tracks from the Kimmeridgian of Pedreira do Avelino, Sesimbra (Azóia Fm.) and Porto das Barcas, Lourinhã (Lourinhã Fm.). An enigmatic track suggests the possible presence of a small, tail-dragging tetrapod. Possible track-makers are suggested based on the known Late Jurassic vertebrate fauna of Portugal.

Mateus, O. (2010).  Paleontological collections of the Museum of Lourinhã (Portugal). (Brandao, JM, Callapez, PM, O. Mateus, Castro, P, Ed.).Colecções e museus de Geologia: missão e gestão. 121-126., Jan: Ed. Universidade de Coimbra e Centro de Estudos e Filosofia da História da Ciência Coimbra Abstractmateus_2010_paleontological_collections_of_the_museum_of_lourinha__geocoleccoes_omateus.pdf

Abstract: The paleontological collections of the Museum of Lourinhã, in Portugal, has a rich paleontological collection, particularly of Late Jurassic dinosaurs of the Lourinhã Formation (Kimmeridgian-Tithonian). Most salient highlights comprehend the following dinosaur holotype specimens: stegosaur Miragaia longicollum, theropod Lourinhanosaurus antunesi, sauropod Dinheirosaurus lourinhanensis, ornithopod Draconyx loureiroi, theropod Allosaurus europaeus, and, a mammal, Kuehneodon hahni. Other dinosaur specimens are referred including the nest and eggs and embryos of Lourinhanosaurus. Portugal is very productive in Late Jurassic vertebrates, being the seventh country bearing more dinosaur taxa.

Mateus, O., Callapez P. M., Polcyn M. J., Schulp A. S., Gonçalves A. O., & Jacobs L. L. (2019).  The Fossil Record of Biodiversity in Angola Through Time: A Paleontological Perspective. (Huntley, Brian J., Russo, Vladimir, Lages, Fernanda, Ferrand, Nuno, Ed.).Biodiversity of Angola: Science & Conservation: A Modern Synthesis. 53–76.: Springer International Publishing Abstractmateus2019_chapter_thefossilrecordofbiodiversityi.pdf

This chapter provides an overview of the alpha paleobiodiversity of Angola based on the available fossil record that is limited to the sedimentary rocks, ranging in age from Precambrian to the present. The geological period with the highest paleobiodiversity in the Angolan fossil record is the Cretaceous, with more than 80{%} of the total known fossil taxa, especially marine molluscs, including ammonites as a majority among them. The vertebrates represent about 15{%} of the known fauna and about one tenth of them are species firstly described based on specimens from Angola.

Mateus, O., Laven T., & Knotschke N. (2004).  A dwarf between giants?: A new late Jurassic sauropod from Germany. Journal of Vertebrate Paleontology. 23, 90–90., Number suppl. to Abstract
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Mateus, O., Jacobs L., Polcyn M., Schulp A. S., Vineyard D., Neto A. B., & Antunes M. T. (2009).  The oldest African eucryptodiran turtle from the Cretaceous of Angola. Acta Palaeontologica Polonica. 54, 581-588., Jan: Univ Agostinho Neto, Univ Nova Lisboa, Museu Lourinha, Acad Ciencias Lisboa, Nat Hist Museum Maastricht, So Methodist Univ Abstractmateus_et_al_2009_the_oldest_african_angolachelys_angola_turtle.pdfWebsite

A new Late Cretaceous turtle, Angolachelys mbaxi gen. et sp. nov., from the Turonian (90 Mya) of Angola, represents the oldest eucryptodire from Africa. Phylogenetic analysis recovers Angolachelys mbaxi as the sister taxon of Sandownia harrisi from the Aptian of Isle of Wight, England. An unnamed turtle from the Albian Glen Rose Formation of Texas (USA) and the Kimmeridgian turtle Solnhofia parsonsi (Germany), are successively more distant sister taxa. Bootstrap analysis suggests those four taxa together form a previously unrecognized monophyletic clade of marine turtles, herein named Angolachelonia clade nov., supported by the following synapomorphies: mandibular articulation of quadrate aligned with or posterior to the occiput, and basisphenoid not visible or visibility greatly reduced in ventral view. Basal eucryptodires and angolachelonians originated in the northern hemisphere, thus Angolachelys represents one of the first marine amniote lineages to have invaded the South Atlantic after separation of Africa and South America.

Mateus, O., & Antunes M. T. (2000).  Late Jurassic dinosaurs of Portugal. Abstracts of the 1st Symposium of European Dinosaurs. , Dusseldorf, Germany. Abstract
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Mateus, O. (1998).  Dinossauros de Portugal e um novo terópode do Jurássico Superior da Lourinhã. , Évora: Universidade de Évora Abstract
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Mateus, I., Mateus H., Antunes M. T., Mateus O., Taquet P., Ribeiro V., & Manuppella G. (1997).  Couvée, øe}ufs et embryons d{\textquotesingle}un Dinosaure Théropode du Jurassique supérieur de Lourinha (Portugal). Comptes Rendus de l{\textquotesingle}Académie des Sciences - Series {IIA} - Earth and Planetary Science. 325, 71–78., jul, Number 1: Elsevier {BV} AbstractWebsite
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Mateus, O., & Milàn J. (2010).  A diverse Upper Jurassic dinosaur ichnofauna from central-west Portugal. Lethaia. 43, 245-257., Number 2 Abstract
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Mateus, O. (2009).  The sauropod Turiasaurus riodevensis in the the Late Jurassic of Portugal. Journal of Vertebrate Paleontology. 29, 144–144., Number 3 Abstract
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Mateus, O. (2012).  Evidence for presence of clavicles and interclavicles in sauropod dinosaurs and its implications on the furcula-clavicle homology. Journal of Vertebrate Paleontology. 184–185., 1 Abstract

Clavicles and interclavicles are plesiomorphically present in Reptilia. However, several groups show reduction or even loss of these elements. Crocodylimorpha, e.g., lost the clavicles, whereas dinosaurs are generally interpreted to only preserve the clavicles, the theropod furcula representing an unique case of fused clavicles. In sauropods, reports of clavicles are relatively frequent in non-titanosauriforms. These elements are elongated, curved, and rather stout bones with a spatulate and a bifurcate end. However, they were always found as single bones, and differ from the relatively short and unbifurcated clavicles found articulated with the scapulae of basal sauropodomorphs. Elements from the Howe Quarry (Late Jurassic; Wyoming, USA) shed new light on these interpretations. Besides the elongated, curved bones (herein named morphotype A), also pairs of symmetric, L-shaped bones were recovered (morphotype B), associated with diplodocid dorsal and cervical vertebrae. Elements resembling morphotype B - articulated between the scapulae - have recently been reported from a diplodocid found near Tensleep, Wyoming. Taphonomic evidence, as well as the fact that they were preserved in symmetrical pairs, therefore implies that morphotype B represents the true sauropod clavicles. Contrary to earlier reports, morphotype A elements from the Howe Quarry, as well as of previously reported specimens show a symmetry plane following the long axis of the elements. It is thus possible that the morphotype A elements were single bones from the body midline. The only such element present in the pectoral girdle of tetrapods are the interclavicle and the furcula. Comparison with crocodilian and lacertiform interclavicles indicates that the bifurcate end of the sauropod elements might represent the reduced transverse processes of the anterior end, and the spatulate end would have covered the coracoids or sternal plates ventrally. The presence of both clavicles and interclavicles in the pectoral girdle stiffens the anterior trunk, and enhances considerably its stability. Such an enforcement might have been needed in diplodocids due to the strong lateral forces induced to the fore-limbs by the posteriorly placed center of mass (due to shorter fore- than hind-limbs), as well as lateral movements of the enormously elongated necks and tails. The absence of clavicles and interclavicles in titanosauriforms coincides with the development of wide-gauge locomotion style. The presence of interclavicles in sauropods supports the recently proposed homology of the furcula with the interclavicle, instead of representing fused clavicles. Interclavicles were thus not lost, but may have remained cartilaginous or have yet to be found in basal dinosauriforms.

Mateus, O. (2008).  Fósseis de transição, elos perdidos, fósseis vivos e espécies estáveis. (Levy, et al, Ed.).Evolução: História e Argumentos. 77-96., Lisboa: Esfera do Caosmateus_2008_evolucao_fosseis_de_transicao.pdf
Mateus, O., & Milan J. (2008).  Sauropod forelimb flexibility deduced from deep manus tracks. (University of, Glasgow, Ed.).52th Paleontological Association Annual Meeting. 18th-21st December 2008. 67-68. Abstract
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Mateus, O., Milàn J., Romano M., & Whyte M. A. (2011).  New finds of stegosaur tracks from the Upper Jurassic Lourinhã formation, Portugal. Acta Palaeontologica Polonica. 56, 651-658., Number 3 Abstract
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Mateus, O., & Milan J. (2005).  Ichnological evidence for giant ornithopod dinosaurs in the Late Jurassic Lourinhã Formation, Portugal. Abstract Book of the International Symposium on Dinosaurs and Other Vertebrates Palaeoichnology. 60–60., Fumanya, Barcelona Abstract
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