Este capítulo apresenta uma visão geral da paleobiodiversidade alfa de Angola com base no registo fóssil disponível, o qual se limita às rochas sedimentares, a sua idade variando entre o Pré‑Câmbrico e o pre‑
sente. O período geológico com a maior paleobiodiversidade no registo fóssil angolano é o Cretácico, com mais de 80% do total dos táxones fósseis conhecidos, especialmente moluscos marinhos, sendo estes na sua maioria
amonites. Os vertebrados representam cerca de 15% da fauna conhecida e cerca de um décimo destes são espécies descritas pela primeira vez com base em espécimes de Angola.

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Dinosaurs and other fossils have been artificially enhanced, or totally forged, to increase their commercial value. The most problematic forgeries to detect are based on original fossils that are artificially assembled. Several techniques are suggested for detecting hoaxes: detailed visual examination, chemical analysis, Xray or CT-scan, and ultraviolet light. It is recommended that museums and paleontological researchers do not purchase and/or trade fossils lacking clear provenience information. Exceptions to that general rule should be closely examined using techniques described herein.

Although nonavian theropod have received considerable interest in the last years, their ontogeny still remains poorly understood, especially the ontogenetical changes affecting their skull (Rauhutand Fechner, 2005). The quadrate, for instance, is preserved in several embryos and juvenile specimens belonging to many clades of theropods such as the Tyrannosauridae (Carr, 1999), Compsognathidae (Dal Sasso and Maganuco, 2011), Therizinosauroidea (Kúndrat et al., 2007), Oviraptoridae (Norell et al., 1994; Norell et al., 2001; Weishampel et al., 2008) and Troodontidae (Varrichio et al., 2002) but very little is usually said about the anatomy of this bone and no one has ever investigated ontogenetical variation in the nonavian theropod quadrate. The discovery of two quadrates belonging to embryos of the sinraptorid Lourinhanosaurus antunesi from Portugal and five isolated quadrates pertaining to juvenile, subadult and adult specimens of Spinosauridae from Morocco fills this gap and allows some ontogenetic information to be drawn for this bone in these two specific clades of Theropoda.

ABSTRACT: Type specimens (holotypes, neotypes, syntypes, etc.) are of crucial importance because they are the only tangible evidence of the nomenclatural act that supports the understanding of paleobiodiversity. The list of the vertebrate species whose type specimen is based on fossils from Portugal is presented here. We counted 206 species, of which there are 45 bony fishes, 39 Late Jurassic Mammaliamorpha, 33 Cenozoic mammals, 28 non-avian dinosaurs, 25 non-dinosaur reptiles, 11 Cenozoic Aves, 22 Chondrichthyes and three other vertebrates. Except for the Quaternary fossil record, the type specimens can be used as a shortcut for measuring the fossil record and paleobiodiversity through geological time and rock units because they correlate in 95% with the fossil record by epoch in the case-study of fossil vertebrates of Portugal.

Abstract We report here new and the first mammaliamorph tracks from the Early Cretaceous of Africa. The tracksite, that also bears crocodylomorph and sauropod dinosaurian tracks, is in the Catoca diamond mine, Lunda Sul Province, Angola. The mammaliamorph tracks have a unique morphology, attributed to Catocapes angolanus ichnogen. et ichnosp. nov. and present an anterolateral projection of digit I and V. The tracks with an average length of 2.7 cm and width of 3.2 cm are the largest mammaliamorph tracks known from the Early Cretaceous unmatched in size in the skeletal fossil record. The crocodylomorph trackways and tracks are attributed to Angolaichnus adamanticus ichnogen. et ichnosp. nov. (‘ichnofamily’ Batrachopodidae) and present a functionally pentadactyl pes, an extremely outwardly rotated handprint, and an unusual tetradactyl and plantigrade manus. One medium-sized sauropod dinosaur trackway preserved skin impressions of a trackmaker with stride length of 1.6 m; a second is that of a small-sized sauropod trackmaker with a pace length of 75 cm.

Stegosaurian dinosaurs have a quadrupedal stance, short forelimbs, short necks, and are generally considered to be low browsers. A new stegosaur, Miragaia longicollum gen. et sp. nov., from the Late Jurassic of Portugal, has a neck comprising at least 17 cervical vertebrae. This is eight additional cervical vertebrae when compared with the ancestral condition seen in basal ornithischians such as Scutellosaurus.
Miragaia has a higher cervical count than most of the iconically long-necked sauropod dinosaurs. Long neck length has been achieved by ‘cervicalization’ of anterior dorsal vertebrae and probable lengthening of centra. All these anatomical features are evolutionarily convergent with those exhibited in the necks of
sauropod dinosaurs. Miragaia longicollum is based upon a partial articulated skeleton, and includes the only known cranial remains from any European stegosaur. A well-resolved phylogeny supports a new clade that unites Miragaia and Dacentrurus as the sister group to Stegosaurus; this new topology challenges the common view of Dacentrurus as a basal stegosaur.

dinosaur genera Diceratops Lull, 1905 and Microceratops Bohlin, 1953 are preoccupied by the Hymenoptera insects, Diceratops Foerster, 1868 and Microceratops Seyrig, 1952, respectively. Therefore, the name of the ceratopsian dinosaur Diceratops Lull, 1905 from the Late Cretaceous of United States is a junior homonym of the hymenoptera Diceratops Foerster, 1868. Diceratus n. gen. (Greek di ‘‘two,’’ Greek ceratos ‘‘horned’’) is proposed as the replacement name of Diceratops Lull, 1905. Some workers have considered Diceratops synonymous with Triceratops (e.g., Dodson and Currie, 1990) but it was reinstated by Forster (1996) after analysis of the characteristics of all existing ceratopsid skulls, and recent reviews (e.g., Dodson et al., 2004) have considered Diceratops a valid genus.
Due to preoccupation, the name of the ceratopsian dinosaur Microceratops Bohlin, 1953 from the Cretaceous of the Gobi is
a junior homonym of the insect Microceratops Seyrig, 1952. Microceratus n. gen. (Greek micro ‘‘small,’’ Greek ceratos ‘‘horned’’) is proposed as the replacing name of Microceratops Bohlin, 1953.
Sereno (2000:489) has declared Microceratops a nomen dubium since the holotype material lacks any diagnostic features, a
convention followed by You and Dodson (2004:480). However, the name is still used by Le Loeuff et al. (2002), Lucas (2006),
Alifanov (2003) and Xu et al. (2002), and such practice justifies the renaming of the genus.
In order to preserve some stability, the names chosen here deliberately preserve the same prefixes.

The precursor of the North Atlantic existed between the North American and Iberian blocks from the earliest Jurassic Hettangian and has been ever expanding since. By the Kimmeridgian and Tithonian, when much of the Morrison Fm rocks were deposited, the proto-Atlantic was more than 300 km wide at 27° paleolatitude between North America and Iberia. Macrovertebrate paleontology reveals a unique story to the isolation of Iberia and instead suggest a paleogeographic land connection between North American and Iberia. Torvosaurus, Allosaurus, Ceratosaurus, Stegosaurus, Supersaurus and others have a distribution restricted to Morrison Formation in North America and Lourinhã Formation in Portugal. A novel paleogeographic model is here suggested: (1) around the Middle–Late Jurassic transition there is a major palaeoceanographic and palaeoclimatic reorganization, coincidental to a major eustatic sea-level drop and uplift associated with the Callovian– Oxfordian Atlantic Regressive Event; (2) creating an ephemeral land bridge presenting a temporary opportunity for terrestrial gateways likely across the Flemish Cap and Galician Bank land masses, allowing large dinosaurian taxa to cross the northern proto-Atlantic in both directions; (3) finally, a Callovian–Oxfordian faunal exchange around the 163 Ma, through latest Kimmeridgian at 152 Ma (the age of equivalent genera in both Morrison and Portugal), is was an interval that allowed speciation, but retaining generic similarity of vertebrates. This model is consistent with the chronology and taxonomy required for speciation of the Iberian and American forms, exemplified by the coeval sister-taxa pairs Torvosaurus tanneri and T. gurneyi, Allosaurus fragilis and A. europaeus, or Supersaurus vivianae and S. lourinhanensis. While some of the smaller animals in the fauna show Morrison/Portugal affinities, most from Iberia have European or even Asian affinities. The larger-bodied fauna are more closely related to Morrison than to mainland Europe (except for dacentrurine stegosaurs). The body size differences and affinities of taxa across paleogeography is comparable to what is observed today across the Wallace Line. Migration may have also occurred in both directions. The closest relative of Torvosaurus is likely the European Bathonian Megalosaurus, thus the presence of the genus in North America represents a European migration. On other hand, Allosaurus and Supersaurus origins are consistent with a North American origin, representing an westto-east migration.

A forelimb of a new sauropod dinosaur (Angolatitan adamastor n. gen. et sp.) from the Late Turonian of Iembe (Bengo Province) represents the first dinosaur discovery in Angola, and is one of the few occurrences of sauropod dinosaurs in sub-Saharan Africa collected with good chronological controls. The marginal marine sediments yielding the specimen are reported to be late Turonian in age and, thus it represents a non-titanosaurian sauropod in sub-Saharan Africa at a time taken to be dominated by titanosaurian forms. Moreover, Angolatitan adamastor is the only basal Somphospondyli known in the Late Cretaceous which implies in the existence of relict forms in Africa.

A survey of Hemidactylus turcicus L. (Reptilia, Gekkonidae) was carried out from March to October
of 1996, in temporally limited transects, using quadrats of 160x160 meters, in the World Heritage site
of Évora. Occasional observations were also made in 1995 and 1996. Hemidactylus turcicus is
common in the sampled area, occurring in 56% of the quadrats, and prefers quiet streets. The species
Tarentola mauritanica was not observed in Évora but it appears 10 Km to the Southwest of this town.
Diurnal activity and winter activity were not observed.