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2008
Araujo, R., Castanhinha R., & Mateus O. (2008).  Major trends in the evolution of teeth and mandibles in ornithopod dinosaurs. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 18., Neuquén, Argentina Abstract
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Araujo, R., Castanhinha R., & Mateus O. (2008).  Major trends in the evolution of teeth and mandibles in ornithopod dinosaurs. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 18–18., Neuquén, Argentina Abstract
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Schulp, A. S., Polcyn M. J., Mateus O., Jacobs L. L., & Morais M. L. (2008).  A new species of Prognathodon (Squamata, Mosasauridae) from the Maastrichtian of Angola, and the affinities of the mosasaur genus Liodon. Proceedings of the Second Mosasaur Meeting, Fort Hays Studies Special Issue 3, Fort Hays State University, Hays, Kansas. 1-12. Abstract
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Mateus, O., Maidment S. C. R., & Christiansen N. A. (2008).  A new specimen aff. Dacentrurus armatus (Dinosauria: Stegosauridae) from the Late Jurassic of Portugal. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 157., Neuquén, Argentina Abstract
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Mateus, O., Maidment S. C. R., & Christiansen N. A. (2008).  A new specimen aff. Dacentrurus armatus (Dinosauria: Stegosauridae) from the Late Jurassic of Portugal. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 157–157., Neuquén, Argentina Abstract
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Mateus, O., Natario C., Araujo R., & Castanhinha R. (2008).  A new specimen of spinosaurid dinosaur aff. Baryonyx from the Early Cretaceous of Portugal. (Universidade de, Coimbra, Ed.).Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 51., Coimbra Abstract
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Mateus, O., Natario C., Araujo R., & Castanhinha R. (2008).  A new specimen of spinosaurid dinosaur aff. Baryonyx from the Early Cretaceous of Portugal. Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 51–51., Coimbra Abstract
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Mateus, O., & Castanhinha R. (2008).  PaleoAngola- Predadores de um oceano primitivo. National Geographic Portugal. 8, 26-33., Number 91 Abstract
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Mateus, O., & Castanhinha R. (2008).  PaleoAngola- Predadores de um oceano primitivo. National Geographic Portugal. 8, 26–33., Number 91 Abstract
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Castanhinha, R., Araujo R., & Mateus O. (2008).  Reptile Egg Sites From Lourinhã Formation, Late Jurassic, Portugal. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. , Neuquén, Argentina Abstract
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Castanhinha, R., Araujo R., & Mateus O. (2008).  Reptile Egg Sites From Lourinhã Formation, Late Jurassic, Portugal. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. , Neuquén, Argentina Abstract
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Mateus, O., & Milan J. (2008).  Sauropod forelimb flexibility deduced from deep manus tracks. (University of, Glasgow, Ed.).52th Paleontological Association Annual Meeting. 18th-21st December 2008. 67-68. Abstract
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Rita, F., Mateus O., & Overbeeke M. (2008).  Tomografia Computorizada na Deteccão de Fraudes em Fósseis. Acta Radiológica Portuguesa. 80, 83-84., Number 20 Abstract
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Rita, F., Mateus O., & Overbeeke M. (2008).  Tomografia Computorizada na Deteccão de Fraudes em Fósseis. Acta Radiológica Portuguesa. 80, 83–84., Number 20 Abstract
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Mateus, O. (2008).  Two ornithischian dinosaurs renamed: Microceratops Bohlin 1953 and Diceratops Lull 1905. Journal of Paleontology. 82, , Number 2 Abstract
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Mateus, O. (2008).   Fósseis de transição, elos perdidos, fósseis vivos e espécies estáveis . Evolução: História e Argumentos. 77-96., Lisboa Abstract
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Mateus, O. (2008).  Checklist for Late Jurassic reptiles and amphibians from Portugal. Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 55., Coimbra Abstractmateus_2008_lista_de_repteis_e_anfibios_do_jurassico_superior_de_portugal__list_congressoherpetolog.pdf

The richness of Late Jurassic vertebrates in Portugal is known since the 19th century by Paul Choffat, Henri Sauvage and other. The Kimmeridgian Guimarota fauna assemblage is the best known, followed by the fauna of Lourinhã formation. Here is presented an attempt to provide a checklist of the reptiles and amphibians of the Late Jurassic. Amphibia: Lissamphibia (Celtedens, cf. Marmorerpeton, Discoglossidae indet.). Chelonia: Eucryptodira (Pleurosternidae indet., Platychelyidae indet., Plesiochelys cf. etalloni, Plesiochelys choffati, Anosteirinae indet.). Squamata: Scincomorpha (Becklesius hoffstetteri; Paramacellodus sp., Saurillodon proraformis, S. henkeli, S. cf. obtusus). Squamata: Anguimorpha (Dorsetisaurus pollicidens, Parviraptor estesi). Crown Lepidosauromorpha (Marmoretta sp.). Choristodera: Cteniogenidae (Ctenogenys reedi). Sauropterygia: Plesiosauria: Cryptoclidoidea: Cryptoclididae indet. Crocodylomorpha (Lisboasaurus estesi, L. mitrocostatus). Crocodyliformes: Neosuchia (Machimosaurus hugii, Goniopholis cf. simus, Goniopholis baryglyphaeus, cf. Bernissartia, Atoposauridae, Theriosuchus guimarotae, cf. Alligatorium, Metriorhynchus sp.). Pterosauria (Rhamphorhynchus sp., Pterodactylus sp.). Dinosauria: Theropoda (Ceratosaurus sp. , Torvosaurus sp., Lourinhanosaurus antunesi, Allosaurus europaeus, Cf. Compsognathus sp., cf. Richardoestesia sp., Dromaeosaurinae indeter., Velociraptorinae indeter., cf. Archaeopteryx sp., aff. Paronychodon). Dinosauria: Sauropoda: Eusauropoda (Dinheirosaurus lourinhanensis, Lourinhasaurus alenquerensis, Lusotitan atalaiensis, Apatosaurus sp.). Dinosauria: Ornithischia: Thyreophora (Dacentrurus armatus, Stegosaurus sp., Dracopelta zbyszewskii). Dinosauria: Ornithischia: Ornithopoda (Phyllodon henkeli, Dryosaurus sp., Hypsilophodon sp., Alocodon kuehnei, Trimucrodon cuneatus, Draconyx loureiroi).

Hayashi, S., Carpenter K., Watabe M., Mateus O., & Barsbold R. (2008).  Defensive weapons of thyreophoran dinosaurs: histological comparisons and structural differences in spikes and clubs of ankylosaurs and stegosaurs. Journal of Vertebrate Paleontology. 28(3, Supplement), 89A-90A., Number Suppl. to 3 Abstracthayashi_et_al_2008_histology_stegosaurs_defensive_weapons_of_thyreophoran_dinosaurs-_histological_comparisons_and_structural_differences_in_spikes_and_clubs_of_ankylosaurs_and_stegosaurs.pdfWebsite

Thyreophoran dinosaurs have spike- and club-shaped osteoderms probably used for defensive weapons. The structural and histological variations have been little known. Here, we provide the comparisons of the internal structures in defensive weapons of ankylosaurs and stegosaurs, using spikes of a polacanthid (Gastonia) and a nodosaurid (Edmontonia), clubs of ankylosaurids (Saichania and Ankylosauridae indet. from Canada), and spikes of stegosaurids (Stegosaurus and Dacentrurus), which sheds light on understandings of evolutionary history and functional implications of defensive weapons in thyreophorans. In ankylosaurs, the structural and histological features of spikes and clubs are similar with those of small osteoderms in having thin compact bones, thick cancellous bones with large vascular canals, and abundant collagen fibers. A previous study demonstrated that each of three groups of ankylosaurs (polacanthid, nodosaurid, and ankylosaurid) has distinct characteristic arrangements of collagen fibers in small osteoderms. This study shows that spikes and clubs of ankylosaurs maintain the same characteristic features for each group despite of the differences in shapes and sizes. Conversely, the spike-shaped osteoderms in primitive (Dacentrurus) and derived (Stegosaurus) stegosaurids have similar structure to each other and are significantly different from the other types of stegosaur osteoderms (throat bony ossicles and plates) in having thick compact bones with a medullary cavity. These lack abundant collagen fibers unlike ankylosaur osteoderms. The spikes of ankylosaurs and stegosaurs are similar in shape, but their structural and histological features are different in having unique structures of collagen fibers for ankylosaurs and thick compact bones for stegosaurs, providing enough strength to have large spikes and to use them as defensive weapons. Although the shapes of ankylosaur clubs are different from spikes, the internal structures are similar, suggesting that ankylosaurs maintain similar structures despite of different shapes in osteoderms. These results indicate that ankylosaurs and stegosaurs used different strategies independently to evolve defensive weapons.

Mateus, O., Jacobs L. L., Polcyn M. J., Schulp A. S., Neto A. B., & Antunes M. T. (2008).  Dinosaur and turtles from the Turonian of Iembe, Angola. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontologia de Vertebrados. 156., Neuquén, Argentina Abstractmateus_et_al_2008_dinosaur_and_turtles_from_the_turonian_of_iembe_angola.pdf

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Mateus, O., Overbeeke M., & Rita F. (2008).  Dinosaur Frauds, Hoaxes and "Frankensteins": How to distinguish fake and genuine vertebrate fossils. Journal of Paleontological Techniques. 2, 1-5.. Abstractmateus_et_al_2008_dinosaur_frauds_hoaxes_and_frankensteins-_how_to_distinguish_fake_and_genuine_vertebrate_fossils._journal_of_paleontological_techniques.pdfWebsite

Dinosaurs and other fossils have been artificially enhanced, or totally forged, to increase their commercial value. The most problematic forgeries to detect are based on original fossils that are artificially assembled. Several techniques are suggested for detecting hoaxes: detailed visual examination, chemical analysis, Xray or CT-scan, and ultraviolet light. It is recommended that museums and paleontological researchers do not purchase and/or trade fossils lacking clear provenience information. Exceptions to that general rule should be closely examined using techniques described herein.

Mateus, O. (2008).  Fósseis de transição, elos perdidos, fósseis vivos e espécies estáveis. (Levy, et al, Ed.).Evolução: História e Argumentos. 77-96., Lisboa: Esfera do Caosmateus_2008_evolucao_fosseis_de_transicao.pdf
Mateus, O., & Jacinto J. (2008).  Hemidactylus turcicus. Atlas dos Anfíbios e Répteis de Portugal. 130-131., Lisboa: A. Loureiro, N. Ferrand de Almeida, M.A. Carretero, O.S. Paulo. Instituto da Conservação da Natureza e da Biodiversidademateus__jacinto_2008_hemidactylus_turcicus_in_atlas_dos_anfibios_e_repteis_de_portugal.pdf
Mateus, O., & Milan J. (2008).  Ichnological evidence for giant ornithopod dinosaurs in the Upper Jurassic Lourinhã Formation, Portugal. Oryctos. 8, 47-52. Abstractmateus_and_milan_2008_ichnological_evidence_for_giant_ornithopod_big_ornithopod_track_from_u_j_lourinha_fm_portugal.pdfWebsite

The Upper Jurassic Lourinhã Formation (Lusitanian Basin, Portugal) contains a diverse dinosaur fauna comprising theropods, sauropods, stegosaurs, ankylosaurs and several genera of ornithopods. The sedimentology in the area favours preservation of tracksways, and tracks from most of the dinosaurs are also represented by skeletal remains. During fieldwork in the summer of 2003 a new, large, tridactyl track was found at the beach of Vale Frades, approximately 6 km north of Lourinhã (central west Portugal). The track was found together with a stegosaur track on a clay bed exposed within the intertidal zone. Due to the immediate danger of erosion, the track was collected and is now on display at Museu da Lourinhã. The track is 70 cm long and 69 cm wide, the toes are short and broad, with indications of short blunt claws, and there is a high angle of divarication between the outer digits. The shape and dimensions of the track identifies it as deriving from an ornithopod dinosaur with an estimated hip height around three metres. Although very large ornithopods are known from the Cretaceous, the largest known Jurassic ornithopod is Camptosaurus from North America, and the largest known from Portugal is the camptosaurid Draconyx loureiroi. Neither of these reached the body size suggested by the new track. So far the track described herein is the only evidence for a Jurassic ornithopod of that size.

Araujo, R., Castanhinha R., & Mateus O. (2008).  Major trends in the evolution of teeth and mandibles in ornithopod dinosaurs. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 18., Neuquén, Argentina Abstractaraujo_et_al._2008._major_trends_in_the_evolution_of_teeth_and_mandibles_in_ornithopod_dinosaurs.pdf

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Schulp, A. S., Polcyn M. J., Mateus O., Jacobs L. L., & Morais M. L. (2008).  A new species of Prognathodon (Squamata, Mosasauridae) from the Maastrichtian of Angola, and the affinities of the mosasaur genus Liodon. Proceedings of the Second Mosasaur Meeting, Fort Hays Studies Special Issue 3, Fort Hays State University, Hays, Kansas. 1-12. Abstractschulp_et_al_2008_prognathodon_kianda.pdf

Here we describe a new species of the mosasaurine genus Prognathodon from the Maastrichtian of Namibe, Angola, on the basis of five specimens which represent most of the cranial skeleton including the diagnostic quadrate. Phylogenetic analysis shows this new taxon, P. kianda nov. sp., to be the sister taxon to all other species of Prognathodon. It is unique amongst Prognathodon in possessing a high marginal tooth count and relatively small pterygoid teeth. The tooth morphology in the new taxon is reminiscent of some species of the genus Liodon, allowing association of Liodon-like dentition with otherwise Prognathodon-like crania, and thus resolves the long-standing question of the phylogenetic affinities of Liodon.

Mateus, O., Maidment S. C. R., & Christiansen N. A. (2008).  A new specimen aff. Dacentrurus armatus (Dinosauria: Stegosauridae) from the Late Jurassic of Portugal. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontologia de Vertebrados. 157., Neuquén, Argentina Abstractmateus_et_al_2008_a_new_specimen_aff._dacentrurus_armatus_dinosauria_stegosauridae_from_the_late_jurassic_of_portugal.pdf

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Castanhinha, R., Araujo R., & Mateus O. (2008).  Reptile Egg Sites From Lourinhã Formation, Late Jurassic, Portugal. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontologia de Vertebrados. , Neuquén, Argentina Abstractcastanhinha_et_al_2008_reptile_egg_sites_from_lourinha_formation_late_jurassic_portugal.pdf

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Mateus, O., & Milan J. (2008).  Sauropod forelimb flexibility deduced from deep manus tracks. 52th Paleontological Association Annual Meeting. 18th-21st December 2008,. 67-68.: University of Glasgow Abstractmateus__milan_2008_palass_sauropod_forelimb_flexibility_deduced_from_deep_manus_tracks.pdf

Sauropods are often considered to have very limited mobility and reduced limb flexibility, mainly due to their giant size and consequent weight. In the Upper Jurassic Lourinhã Formation, central-west Portugal, deep vertical natural casts of sauropod manus tracks are often preserved as the infills of the original tracks. These manus tracks are vertical-walled, with marks of the striations of the skin scales, showing that the movement of the sauropod manus impacting and exiting the mud was totally vertical with no horizontal component of the stride. Some tracks are up to 66 cm deep, which is equivalent to the height of whole sauropod manus. This means that sauropods could lift their anterior feet in a complete vertical manner. Such movement is only possible if there is mobility at elbow and shoulder articulations in a higher degree than previously thought for sauropods. Our vision of sauropod limbs as inflexible columns has to be updated to a more dynamic model for limbs and body.

Mateus, O., & Araújo R. (2008).  Stone-splitters and expansive demolition agents: breaking big rocks with little effort on vertebrate paleontological excavations. Journal of Paleontological Techniques. 3, 1-6. Abstractmateus__araujo_2008_stone-splitters_and_expansive_demolition_agents_jpt_n003_oct.pdfWebsite

Two techniques (stone-splitters and demolition agents) are revealed to be efficient methods for breaking large stone blocks in extreme paleontological excavation. In certain conditions – where security factors, permission issues, terrain conditions, rock properties are problematic – the traditional methods for breaking large rock blocks cannot be applied (e.g. crane trucks or explosives). Using an expansive demolition agent or stone-splitters after drilling equidistant holes not only allows a cheap, quick and safe solution but also permits precise removal of up to 9 ton blocks. Stone-splitters are a three-part tool that when inserted linearly and equidistantly along a brittle rock mass cause a precise fracture.

Rita, F., Mateus O., & Overbeeke M. (2008).  Tomografia Computorizada na Deteccão de Fraudes em Fósseis. Acta Radiológica Portuguesa. 80, 83-84., Number 20 Abstractrita_et_al_2008_tomografia_computorizada_na_deteccao_de_fraudes_em_fosseis.pdfWebsite

The material in analysis is the skull of an Ornitiquous Psittacosaurus of the China Cretaceous suspicious of being a fraud. The fossil described here appeared to be in very good condition and conservation. The skull is almost complete but because it is filled by sediment, this prevents an analyse of the intra-skull anatomy.
With the intuition of confirming or not the existence of fossiled bone elements in the interior of the sedimented mass that filled the Psittacossaurus skull, it was submitted to a Computorized Tomography.
The Psittacosaurus skull showed an unexpected absorption of the x-rays, because of the outstanding differences of density between the bone and the matrix due to the fact that the interior of the skull was composed by an amalgam of materials, where a less compact and relatively homogeneous material (soil and wax) was found and that material united and mounted the whole skull and the normal bone structures were non-existing.
The capacity of the Computorized Axial Tomography of differentiating materials with different densities of absorption of x-rays, permitted an easy and reliable investigation and explained beyond doubt the quality of the fossil specimen studied, concluding with no doubt that in spite of the realistic aspect, we had come before a fraud.

Mateus, O. (2008).  Two ornithischian dinosaurs renamed: Microceratops Bohlin 1953 and Diceratops Lull 1905. Journal of Paleontology. 82, 423., Number 2 Abstractmateus_2008_two_ornithischians_renamed__microceratops_bohlin_1953_and_diceratops_lull_.pdfWebsite

dinosaur genera Diceratops Lull, 1905 and Microceratops Bohlin, 1953 are preoccupied by the Hymenoptera insects, Diceratops Foerster, 1868 and Microceratops Seyrig, 1952, respectively. Therefore, the name of the ceratopsian dinosaur Diceratops Lull, 1905 from the Late Cretaceous of United States is a junior homonym of the hymenoptera Diceratops Foerster, 1868. Diceratus n. gen. (Greek di ‘‘two,’’ Greek ceratos ‘‘horned’’) is proposed as the replacement name of Diceratops Lull, 1905. Some workers have considered Diceratops synonymous with Triceratops (e.g., Dodson and Currie, 1990) but it was reinstated by Forster (1996) after analysis of the characteristics of all existing ceratopsid skulls, and recent reviews (e.g., Dodson et al., 2004) have considered Diceratops a valid genus.
Due to preoccupation, the name of the ceratopsian dinosaur Microceratops Bohlin, 1953 from the Cretaceous of the Gobi is
a junior homonym of the insect Microceratops Seyrig, 1952. Microceratus n. gen. (Greek micro ‘‘small,’’ Greek ceratos ‘‘horned’’) is proposed as the replacing name of Microceratops Bohlin, 1953.
Sereno (2000:489) has declared Microceratops a nomen dubium since the holotype material lacks any diagnostic features, a
convention followed by You and Dodson (2004:480). However, the name is still used by Le Loeuff et al. (2002), Lucas (2006),
Alifanov (2003) and Xu et al. (2002), and such practice justifies the renaming of the genus.
In order to preserve some stability, the names chosen here deliberately preserve the same prefixes.

Mateus, O., Natário C., Araújo R., & Castanhinha R. (2008).  A new specimen of spinosaurid dinosaur aff. Baryonyx from the Early Cretaceous of Portugal. Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 51., Jan Abstract

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2009
Jacobs, L. L., Mateus O., Polcyn M. J., Schulp A. S., Scotese C. R., Goswami A., Ferguson K. M., Robbins J. A., Vineyard D. P., & Neto A. B. (2009).  Cretaceous paleogeography, paleoclimatology, and amniote biogeography of the low and mid-latitude South Atlantic Ocean. Bulletin de la Societe Geologique de France. 180, 333-341., Number 4 Abstract
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Jacobs, L., Polcyn M., Mateus O., Schulp, & Neto A. (2009).  The Cretaceous Skeleton Coast of Angola. Journal of Vertebrate Paleontology. 29, 121., Number 3 Abstract
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Jacobs, L. L., Polcyn M. J., Mateus O., Schulp A. S., & Neto A. (2009).  The Cretaceous Skeleton Coast of Angola. Journal of Vertebrate Paleontology. 29, 121–121., Number 3 Abstract
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Tomas, C., Mateus O., & Abreu C. (2009).  Dinolourinhã – a integração dos jovens na paleontologia: o caso-estudo do Museu da Lourinhã.. Journal of Paleontological Techniques 5: 28-29.. Abstract
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Castanhinha, R., Araujo R., & Mateus O. (2009).  Dinosaur eggshell and embryo localities in Lourinhã Formation, Late Jurassic, Portugal. Journal of Vertebrate Paleontology. 29, 76., Number 3 Abstract
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Castanhinha, R., Araujo R., & Mateus O. (2009).  Dinosaur eggshell and embryo localities in Lourinhã Formation, Late Jurassic, Portugal. Journal of Vertebrate Paleontology. 29, 76–76., Number 3 Abstract
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Brandao, J., Callapez, P., O. Mateus, Castro, P (Eds.). (2009).  International Conference on the Geological Collections and Museums: mission and management. Abstract
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Mateus, O., Maidment S. C. R., & Christiansen N. A. (2009).  A new long-necked {'}sauropod-mimic{'} stegosaur and the evolution of the plated dinosaurs. Proceedings of the Royal Society B: Biological Sciences. 276, 1815-1821., Number 1663 Abstract
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Polcyn, M., Jacobs L., Mateus O., & Schulp A. (2009).  New specimens of Angolasaurus bocagei and comments on the early radiations of plioplatecarpine mosasaurs. Journal of Vertebrate Paleontology. Journal of Vertebrate Paleontology. 29, 165., Number 3 Abstract
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Polcyn, M. J., Jacobs L. L., Mateus O., & Schulp A. S. (2009).  New specimens of Angolasaurus bocagei and comments on the early radiations of plioplatecarpine mosasaurs. Journal of Vertebrate Paleontology. Journal of Vertebrate Paleontology. 29, 165–165., Number 3 Abstract
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Mateus, O., Jacobs L., Polcyn M., Schulp A. S., Vineyard D., Buta Neto A., & Telles Antunes M. (2009).  The oldest African eucryptodiran turtle from the Cretaceous of Angola. Acta Palaeontologica Polonica. 54, 581-588., Number 4 Abstract
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Araujo, R., Mateus O., Walen A., & Christiansen N. (2009).  Preparation techniques applied to a stegosaurian Dinosaur from Portugal: excavation, laboratory mechanical and chemical preparation, moulding and casting, 3D scanning. Journal of Paleontological Techniques 5: 22.. 22. Abstract
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Mateus, O. (2009).  The sauropod Turiasaurus riodevensis in the the Late Jurassic of Portugal. Journal of Vertebrate Paleontology. 29, 144., Number 3 Abstract
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Mateus, O. (2009).  The sauropod Turiasaurus riodevensis in the the Late Jurassic of Portugal. Journal of Vertebrate Paleontology. 29, 144–144., Number 3 Abstract
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Castanhinha, R., Araújo R., & Mateus O. (2009).  Dinosaur eggshell and embryo localities in Lourinhã Formation, Late Jurassic, Portugal. Journal of Vertebrate Paleontology, 29(3): . 76A. Abstractcastanhinhaetal2009dinosaureggshellp.pdf

Four different localities from the Late Jurassic of Lourinhã formation with eggshells and embryos were studied: Paimogo (lower Amoreira-Porto Novo member), Peralta (Praia Azul member), Porto das barcas (Bombarral member) and Casal da Rôla (Amoreira-Porto Novo member). All but Casal da Rôla have embryonic material. Preliminary results show that eggshells from Paimogo correspond to obliquiprismatic morphotype (0.92mm thick), similar to those from Morrison Formation. Within Paimogo site a different type of eggshell was discovered, having a radial section of 153 μm with a mammilary layer measuring 65 μm. Porto das Barcas eggshells represent a discretispherulitic morphotype (1,23 mm thick).
This locality presents a nest 60-cm diameter containing many eggshells but an indeterminate number of eggs. Some embryonic bones were discovered between the eggshells including teeth and skull bones showing that the eggs belong to a saurischian, tentatively a sauropod dinosaur. Peralta nest eggshells are preliminary ascribed to obliquiprismatic morphotype (column: 0,56mm and mammilla: 0,21mm) probably related to Paimogo’s nest taxon (Lourinhanosaurus). Peralta site bears embryonic bones namely small theropod teeth associated with bone fragments, and unidentifiable dinosaur vertebra. Only eggshells have been collected at Casal da Rôla (ML1194). The eggshells (0,78mm thick) are prismatic morphotype and it was impossible to determine the pore system, the outer surface is smooth with no ornamentation.
Lourinhã formation has the oldest sauropod and theropod nest with embryos known so far.