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Marx, M. P., Mateus O., Polcyn M. J., Schulp A. S., Gonçalves O. A., & Jacobs L. L. (2021).  The cranial anatomy and relationships of Cardiocorax mukulu (Plesiosauria: Elasmosauridae) from Bentiaba, Angola. PLOS ONE. 16(8), e0255773 - ., 2021/08/17: Public Library of Science Abstractmarx_et_al_2021_cardiocorax_angola.pdfWebsite

We report a new specimen of the plesiosaur Cardiocorax mukulu that includes the most complete plesiosaur skull from sub-Saharan Africa. The well-preserved three-dimensional nature of the skull offers rare insight into the cranial anatomy of elasmosaurid plesiosaurians. The new specimen of Cardiocorax mukulu was recovered from Bentiaba, Namibe Province in Angola, approximately three meters above the holotype. The new specimen also includes an atlas-axis complex, seventeen postaxial cervical vertebrae, partial ribs, a femur, and limb elements. It is identified as Cardiocorax mukulu based on an apomorphy shared with the holotype where the cervical neural spine is approximately as long anteroposteriorly as the centrum and exhibits a sinusoidal anterior margin. The new specimen is nearly identical to the holotype and previously referred material in all other aspects. Cardiocorax mukulu is returned in an early-branching or intermediate position in Elasmosauridae in four out of the six of our phylogenetic analyses. Cardiocorax mukulu lacks the elongated cervical vertebrae that is characteristic of the extremely long-necked elasmosaurines, and the broad skull with and a high number of maxillary teeth (28–40) which is characteristic of Aristonectinae. Currently, the most parsimonious explanation concerning elasmosaurid evolutionary relationships, is that Cardiocorax mukulu represents an older lineage of elasmosaurids in the Maastrichtian.

Mateus, O., Polcyn M. J., Jacobs L. L., Araújo R., Schulp A. S., Marinheiro J., Pereira B., & Vineyard D. (2012).  Cretaceous amniotes from Angola: dinosaurs, pterosaurs, mosasaurs, plesiosaurs, and turtles. V Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno. 71-105., Salas de los Infantes, Burgos Abstractmateus_et_al_2012_amniotes_from_angola_cretaceous_amniotes_from_angola_dinosaurs_pterosaurs_mosasaurs.pdf

Although rich in Cretaceous vertebrate fossils, prior to 2005 the amniote fossil record of Angola was poorly known. Two horizons and localities have yielded the majority of the vertebrate fossils collected thus far; the Turonian Itombe Formation of Iembe in Bengo Province and the Maastrichtian Mocuio Formation of Bentiaba in Namibe Province. Amniotes of the Mesozoic of Angola are currently restricted to the Cretaceous and include eucryptodire turtles, plesiosaurs, mosasaurs, pterosaurs, and dinosaurs. Recent collecting efforts have greatly expanded our knowledge of the amniote fauna of Angola and most of the taxa reported here were unknown prior to 2005.

Mateus, O., Morais M., Schulp A., Jacobs L., & Polcyn M. (2006).  The Cretaceous of Angola. JOURNAL OF VERTEBRATE PALEONTOLOGY. 26, 96A-97A., Jan Abstractmateus_et_al_2006_svp_abstracts_cretaceous_fo_angola.pdf

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Mateus, O., Morais M. L., Schulp A. S., Jacobs L. L., & Polcyn M. J. (2006).  The Cretaceous of Angola. Journal of Vertebrate Paleontology. 26, 96-97., Number (Suppl. To 3) Abstract
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Mateus, O., Morais M. L., Schulp A. S., Jacobs L. L., & Polcyn M. J. (2006).  The Cretaceous of Angola. Journal of Vertebrate Paleontology. 26, 96–97., Number (Suppl. T Abstract
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Jacobs, L. L., Mateus O., Polcyn M. J., Schulp A. S., Scotese C. R., Goswami A., Ferguson K. M., Robbins J. A., Vineyard D. P., & Neto A. B. (2009).  Cretaceous paleogeography, paleoclimatology, and amniote biogeography of the low and mid-latitude South Atlantic Ocean. BULLETIN DE LA SOCIETE GEOLOGIQUE DE FRANCE. 180, 333-341., Jan: Univ Agostinho Neto, Univ Nova Lisboa, So Methodist Univ, Univ Texas Arlington, Museu Lourinha, Nat Hist Museum Abstractjacobs_mateus_et_al_2009_cretaceous_paleogeography_paleoclimatology_and_amniote_biogeography_of_the_south_atlantic_ocean_angola_africa_currents.pdf

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Jacobs, L. L., Mateus O., Polcyn M. J., Schulp A. S., Scotese C. R., Goswami A., Ferguson K. M., Robbins J. A., Vineyard D. P., & Neto A. B. (2009).  Cretaceous paleogeography, paleoclimatology, and amniote biogeography of the low and mid-latitude South Atlantic Ocean. Bulletin de la Societe Geologique de France. 180, 333-341., Number 4 Abstract
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Jacobs, L., Polcyn M., Mateus O., Schulp A. S., & Neto A. B. (2009).  The Cretaceous Skeleton Coast of Angola. Journal of Vertebrate Paleontology. 29, 121A., Jan Abstractjacobs_et_al_2009cretaceousskeletoncoas.pdfWebsite

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Mateus, O. (2009).  The Cretaceous Skeleton Coast of Angola. Journal of Vertebrate Paleontology. 29, 121A., 1, Number 3: Taylor & Francis Abstract

THE CRETACEOUS SKELETON COAST OF ANGOLA JACOBS, Louis, SMU, Dallas, TX, USA; POLCYN, Michael, SMU, Dallas, TX, USA; MATEUS, Octávio, Museu da Lourinhã, Lourinhã, Portugal; SCHULP, Anne, Natuurhistorisch Museum Maastricht, Maastricht, Netherlands; NETO, André , Universidade Agostinho Neto, Luanda, Angola Cretaceous coastal sediments of Angola present a rich and diverse fauna of marine amniotes, including turtles, mosasaurs, and plesiosaurs. The abundance of mosasaurs in particular suggests a highly productive coastal area. Angola today lies at the northern limit of the Namibian Desert, the so-called Skeleton Coast, which results from prevailing southeasterly winds of the descending limb of the southern Hadley Cell sweeping across the African coast. The Benguela upwelling and a highly productive sea are found today off the Namibian Desert coast. However, the Benguela upwelling system, based on results of DSDP studies, is said to have originated in the late Neogene and therefore cannot explain the productivity found along the length of the West African coast. The explanation is found in the northward drift of Africa through the arid climate zone, and is demonstrated by the tracing of the paleogeographic position of fossil localities through time.

Mateus, O. (2009).  The Cretaceous Skeleton Coast of Angola. 29, , 1 Abstract
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Jacobs, L., Polcyn M., Mateus O., Schulp, & Neto A. (2009).  The Cretaceous Skeleton Coast of Angola. Journal of Vertebrate Paleontology. 29, 121., Number 3 Abstract
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Jacobs, L. L., Polcyn M. J., Mateus O., Schulp A. S., & Neto A. (2009).  The Cretaceous Skeleton Coast of Angola. Journal of Vertebrate Paleontology. 29, 121–121., Number 3 Abstract
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Gaspar, A., Avelar T., & Mateus O. (2007).  Criacionismo e Sociedade no Séc. XX. (Avelar, T., O. Mateus, Almada, F., Gaspar, A., Ed.).Evolução e Criacionismo: Uma Relação Impossível. 133-160., Lisboa: Quasi ed. gasparavelarmateus2007evoluoecriacio.pdf
Gaspar, A., Avelar T., & Mateus O. (2007).  Criacionismo e Sociedade no Séc. XX.  Evolução e Criacionismo: Uma Relação Impossível. 133-160., Lisboa Abstract
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Russo, J., Mateus O., Balbino A., & Marzola M. (2014).  Crocodylomorph eggs and eggshells from the Lourinhã Fm. (Upper Jurassic), Portugal. Comunicações Geológicas. 101, Especial I, 563-566. Abstractrusso_et_al_2014_crocodylomorph_eggs_and_eggshells_from_the_lourinha_fm_upper_jurassic_portugal.pdf

We here present fossil Crocodylomorpha eggshells from the Upper Jurassic Lourinhã Formation of Portugal, recovered from five sites: one nest from Cambelas with 13 eggs, and three partial eggs and various fragments from, Paimogo N (I), Paimogo S (II), Casal da Rola, and Peralta. All specimens but the nest were found in association with dinosaur egg material. Our research reveals that on a micro- and ultrastructural analysis, all samples present the typical characters consistent with crocodiloid eggshell morphotype, such as the shell unit shape, the organization of the eggshell layers, and the triangular blocky extinction observed with crossed nicols. We assign the material from the Lourinhã Formation to the oofamily Krokolithidae, making it the oldest crocodylomorph eggs known so far, as well as the best record for eggs of non- crocodylian crocodylomorphs. Furthermore, our study indicates that the basic structure of crocodiloid eggshells has remained stable since at least the Upper Jurassic.

Russo, J., Mateus O., Balbino A., & Marzola M. (2014).  Crocodylomorph eggs and eggshells from the Lourinhã Fm. (Upper Jurassic), Portugal. Comunica\\c cões Geológicas. 101, Especial I, 563-566. Abstract
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Russo, J., Mateus O., Balbino A., & Marzola M. (2014).  Crocodylomorph eggs and eggshells from the Lourinhã Fm. (Upper Jurassic), Portugal. Comunica\\c cões Geológicas. 101, Especial I, 563-566. Abstract
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Russo, J., Mateus O., Balbino A., & Marzola M. (2014).  Crocodylomorph eggs and eggshells from the Lourinhã Fm. (Upper Jurassic), Portugal. Comunica\\c cões Geológicas. 101, Especial I, 563-566. Abstract

We here present fossil Crocodylomorpha eggshells from the Upper Jurassic Lourinhã Formation of Portugal, recovered from five sites: one nest from Cambelas with 13 eggs, and three partial eggs and various fragments from, Paimogo N (I), Paimogo S (II), Casal da Rola, and Peralta. All specimens but the nest were found in association with dinosaur egg material. Our research reveals that on a micro- and ultrastructural analysis, all samples present the typical characters consistent with crocodiloid eggshell morphotype, such as the shell unit shape, the organization of the eggshell layers, and the triangular blocky extinction observed with crossed nicols. We assign the material from the Lourinhã Formation to the oofamily Krokolithidae, making it the oldest crocodylomorph eggs known so far, as well as the best record for eggs of non- crocodylian crocodylomorphs. Furthermore, our study indicates that the basic structure of crocodiloid eggshells has remained stable since at least the Upper Jurassic.

Guillaume, A. R. D., Moreno-Azanza M., Puértolas-Pascual E., & Mateus O. (2018).  Crocodylomorph teeth from the Lourinhã Formation, Portugal (Late Jurassic). XVI Annual Meeting of the European Association of Vertebrate Palaeontologists. 80., Caparica, Portugal June 26th-July 1st, 2018 Abstractguillaume_et_al_2018_eavp_abstract.pdf

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Guillaume, A. R. D., Moreno-Azanza M., Puértolas-Pascual E., & Mateus O. (2018).  Crocodylomorph teeth from the Lourinhã Formation, Portugal (Late Jurassic). XVI Annual Meeting of the European Association of Vertebrate Palaeontologists. 80., Caparica, Portugal June 26th-July 1st, 2018 Abstract

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Mateus, O. (2013).  Crocodylomorphs from the Mesozoic of Portugal and a new skull of eusuchian from the Late Cretaceous. 2013 Hwaseong International Dinosaurs Expedition Symposium, pp.66-67.. , Hwaseong, South Korea Abstractmateus_2013_crocodylomorphs_portugal_new_skull.pdf

The diversity of fossil crocodylomorphs in Portugal is high, with occurrence as old as Mystriosaurus (=Steneosaurus) bollensis from the Lower Jurassic. The Late Jurassic forms are the better documented, and include the following taxa: Machimosaurus hugii, Lisboasaurus estesi Seiffert, 1973, Lusitanisuchus mitrocostatus Seiffert, 1975; Schwarz & Fechner 2004, Theriosuchus guimarotae Schwarz and Salisbury 2005, Cf. Alligatorium, Goniopholis baryglyphaeus, and a crocodylomorph-like eggs in dinosaur nests (Mateus et al., 1998; Ricqlès et al., 2001). From the Lower Cretaceous were reported a few dinosaurs but its record is strangely scarce in crocodylomorphs (Mateus et al., 2011). The Upper Cretaceous crocodiles show a large diversity, but it is mostly based in fragmentary material that require revision, such as “Crocodylus” blavieri? Grey from the Upper Campanian - Maastrichtian of Viso, near Aveiro (initially reported by Sauvage 1897-98), Goniopholis cf. crassidens Owen 1841 and Oweniasuchus pulchelus Jonet 1981. Moreover there is a fascinating, but poorly understood, crocodylomorph diversity in the Cenomanian of Portugal, documented by fragmentary specimens that have been doubtfully assigned to Thoracosaurus Leidy 1852 of the Middle Cenomanian of Cacém, to the nomen dubium Oweniasuchus lusitanicus Sauvage 1897-98 (interpreted as a mesosuchian goniopholid) based in a fragmentary mandible from the Campanian-Maastrichtian, and also from the Middle Cenomanian of Portugal, Buffetaut and Lauverjat (1978) report an fragmentary unidentified possible dyrosaurid from Nazaré. All this specimens are too incomplete to be compared with the specimen here described. In contrast, Cenozoic crocodiles of Portugal are often known after complete skulls and several individuals. The taxa list include Iberosuchus macrodon (Lower to Middle Eocene), Tomistoma calaritanus (Early Miocene) and T. lusitanica (Burdigalian-Helvetian), and Diplocynodon sp. (Antunes, 1961, 1987, 1994).
At least, two different morphotypes of crocodylomorph eggs from the Late Jurassic of Lourinhã Formation are also known.
A new specimen here reported of crocodile based in a partial skull and mandible (ML1818) from the Uppermost Middle Cenomanian platform carbonates of Baixo Mondego, west central Portugal (Tentúgal Fm., Callapez, 2004). The taxon is phylogenetically positioned as a basal Eusuchia, due to the choanae enclosed by the pterygoid, and closely related with stem Crocodylia and Borealosuchus. This specimen represents the only well documented and valid eusuchian species in the Cenomanian of Europe and is the oldest representative of an eusuchian crocodylomorph, with the exception for the Barremian Hylaeochampsa vectiana.

Azanza, M. M., Coimbra R., Puértolas-Pascual E., Russo J., Bauluz B., & Mateus O. (2019).  Crystallography of Lourinhanosaurus eggshells (Dinosauria, Theropoda, Allosauroidea). Journal of Vertebrate Paleontology, Program and Abstracts. 156-157.moreno_azanza_et_al_2019_svp_abstract.pdf
Azanza, M. M., Coimbra R., Puértolas-Pascual E., Russo J., Bauluz B., & Mateus O. (2019).  Crystallography of Lourinhanosaurus eggshells (Dinosauria, Theropoda, Allosauroidea). Journal of Vertebrate Paleontology, Program and Abstracts. 156-157. Abstract
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Marzola, M., Mateus O., Shubin N. H., & Clemmensen L. B. (2017).  Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland). Journal of Vertebrate Paleontology. e1303501., 2017: Taylor & Francis Abstractmarzola_et_al_2017_cyclotosaurus_greenland.pdfWebsite

ABSTRACTCyclotosaurus naraserluki, sp. nov., is a new Late Triassic capitosaurid amphibian from lacustrine deposits in the Fleming Fjord Formation of the Jameson Land Basin in Greenland. It is based on a fairly complete and well-preserved skull associated with two vertebral intercentra. Previously reported as Cyclotosaurus cf. posthumus, C. naraserluki is unique among cyclotosaurs for having the postorbitals embaying the supratemporals posteromedially. The anterior palatal vacuity presents an autapomorphic complete subdivision by a wide medial premaxillary-vomerine bony connection. The parasphenoid projects between the pterygoids and the exoccipitals, preventing a suture between the two, a primitive condition shared with Rhinesuchidae, Eryosuchus, and Kupferzellia. Within Cyclotosaurus, the Greenlandic skull has a distinctive combination of circular choanae (shared with C. ebrachensis, C. posthumus, and C. robustus) and a convex posteromedial margin of the tabulars (also present in C. ebrachensis and C. intermedius). A phylogenetic analysis indicates that C. naraserluki is the sister taxon of the middle Norian C. mordax from southern Germany, with which it shares a pair of premaxillary foramina. Cyclotosaurus is one of the most successful and diverse genera of Late Triassic temnospondyls, with at least eight species reported from middle Carnian to late Norian. Cyclotosaurus naraserluki is the largest amphibian ever reported from Greenland and one of the Late Triassic vertebrates with the highest northern paleolatitude currently known.http://zoobank.org/urn:lsid:zoobank.org:pub:43AAA541-031C-4EE1-B819-4846EBBD1BBBSUPPLEMENTAL DATA?Supplemental materials are available for this article for free at www.tandfonline.com/UJVPCitation for this article: Marzola, M., O. Mateus, N. H. Shubin, and L. B. Clemmensen. 2017. Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland). Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2017.1303501.ABSTRACTCyclotosaurus naraserluki, sp. nov., is a new Late Triassic capitosaurid amphibian from lacustrine deposits in the Fleming Fjord Formation of the Jameson Land Basin in Greenland. It is based on a fairly complete and well-preserved skull associated with two vertebral intercentra. Previously reported as Cyclotosaurus cf. posthumus, C. naraserluki is unique among cyclotosaurs for having the postorbitals embaying the supratemporals posteromedially. The anterior palatal vacuity presents an autapomorphic complete subdivision by a wide medial premaxillary-vomerine bony connection. The parasphenoid projects between the pterygoids and the exoccipitals, preventing a suture between the two, a primitive condition shared with Rhinesuchidae, Eryosuchus, and Kupferzellia. Within Cyclotosaurus, the Greenlandic skull has a distinctive combination of circular choanae (shared with C. ebrachensis, C. posthumus, and C. robustus) and a convex posteromedial margin of the tabulars (also present in C. ebrachensis and C. intermedius). A phylogenetic analysis indicates that C. naraserluki is the sister taxon of the middle Norian C. mordax from southern Germany, with which it shares a pair of premaxillary foramina. Cyclotosaurus is one of the most successful and diverse genera of Late Triassic temnospondyls, with at least eight species reported from middle Carnian to late Norian. Cyclotosaurus naraserluki is the largest amphibian ever reported from Greenland and one of the Late Triassic vertebrates with the highest northern paleolatitude currently known.http://zoobank.org/urn:lsid:zoobank.org:pub:43AAA541-031C-4EE1-B819-4846EBBD1BBBSUPPLEMENTAL DATA?Supplemental materials are available for this article for free at www.tandfonline.com/UJVPCitation for this article: Marzola, M., O. Mateus, N. H. Shubin, and L. B. Clemmensen. 2017. Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland). Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2017.1303501.

Marzola, M., Mateus O., Shubin N. H., & Clemmensen L. B. (2017).  Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland). Journal of Vertebrate Paleontology. e1303501., may: Informa {UK} Limited AbstractWebsite
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