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Myers, T. S., Polcyn M. J., Mateus O., Vineyard D. P., Gonçalves A. O., & Jacobs L. L. (2017).  A new durophagous stem cheloniid turtle from the lower Paleocene of Cabinda, Angola. Papers in Palaeontology. 2017, 1-16. Abstractnew_durophagous_stem_cheloniid_turtle_from_the_lower_paleocene_of_cabinda_angola.pdfWebsite

A new stem cheloniid turtle, Cabindachelys landanensis, gen. et sp. nov., is represented by a nearly complete skull and partial hyoid collected in lower Paleocene shallow marine deposits, equivalent to the offshore Landana Formation, near the town of Landana in Cabinda, Angola. A partial chelonioid carapace previously reported from this locality is referred here to C. landanensis. Cabindachelys landanensis possesses clear synapomorphies of Pan-Cheloniidae, including a rod-like rostrum basisphenoidale, V-shaped basisphenoid crest, and secondary palate, but also retains a reduced foramen palatinum posterius, unlike most other pan-cheloniids. Phylogenetic analysis suggests that C. landanensis forms a weakly-supported clade with Erquelinnesia gosseleti, Euclastes acutirostris, Euclastes wielandi and Terlinguachelys fischbecki, although a close relationship between the protostegid T. fischbecki and these durophagous pan-cheloniids is unlikely. The Paleocene–Eocene strata near Landana have produced a number of turtle fossils, including the holotype specimen of the pleurodire Taphrosphys congolensis. A turtle humerus collected c. 1 m above the holotype skull of C. landanensis differs from humeri of chelonioids and Taphrosphys, indicating that a third turtle taxon is present at Landana. Cheloniid fossil material is rare in the Landana assemblage, in comparison with the abundant remains of Taphrosphys congolensis found throughout the stratigraphic section. This disparity implies that C. landanensis preferred open marine habitats, whereas Taphrosphys congolensis spent more time in nearshore environments. The appearance of new durophagous species such as C. landanensis in the early Paleocene reflects the rapid radiation of pan-cheloniids as they diversified into open niches following the K–Pg extinction.

Myers, T. S., Tabor N. J., Jacobs L. L., & Mateus O. (2012).  Palaeoclimate of the late jurassic of portugal: Comparison with the western united states. Sedimentology. 59, 1695-1717., Number 6 Abstract
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Myers, T. S., Mateus O., Polcyn M. J., Vineyard D., & Jacobs L. L. (2016).  A new chelonioid turtle from the Paleocene of Cabinda, Angola. Journal of Vertebrate Paleontology, Program and Abstracts, 2016, p. 194.. 194. Abstract

We report a new chelonioid turtle on the basis of a nearly complete skull collected in lower Paleocene, shallow marine deposits, equivalent to the offshore Landana Formation, near the town of Landana in Cabinda Province, Angola. Chelonioid material previously reported from this locality is likely referable to this new taxon. The well-preserved skull is missing the left quadrate, squamosal, and prootic, both opisthotics, and the mandible. The skull possesses a rod-like basisphenoid rostrum, which is a synapomorphy of Chelonioidea, but it differs from other chelonioid skulls in that the contact between the parietal and squamosal is absent, and the posterior palatine foramen is present. Phylogenetic analysis recovers the new taxon as a basal chelonioid. The Paleocene– Eocene strata near Landana have produced a wealth of turtle fossils, including the holotype of the pleurodire Taphrosphys congolensis. A turtle humerus collected from the Landana locality differs morphologically from the humeri of chelonioids and Taphrosphys, indicating the presence of a third taxon. Chelonioid fossil material in the Landana assemblage is rare compared to the abundant fragmentary remains of Taphrosphys that are found throughout the stratigraphic section. This disparity in abundance suggests the new chelonioid taxon preferred open marine habitats, whereas Taphrosphys frequented nearshore environments.

Myers, T. S., Mateus O., Polcyn M. J., Vineyard D., & Jacobs L. L. (2016).  A new chelonioid turtle from the Paleocene of Cabinda, Angola. Journal of Vertebrate Paleontology, Program and Abstracts, 2016, p. 194.. 194. Abstract
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Myers, T. S., Tabor N. J., Jacobs L. L., & Mateus O. (2012).  Estimating soil pCO2 using paleosol carbonates: implications for the relationship between primary productivity and faunal richness in ancient terrestrial ecosystems. Paleobiology. 38(4), 585–604. Abstractmyers_et_al_2012_estimating_soil_paleosols_portugal.pdf

In this paper we present a method for estimating soil pCO2 in ancient environments using the measured carbon-isotope values of pedogenic carbonates and plant-derived organic matter. The validity of soil pCO2 estimates proves to be highly dependent on the organic δ13C values used in the calculations. Organic matter should be sourced from the same paleosol profiles as sampled carbonates to yield the most reliable estimates of soil pCO2. In order to demonstrate the potential use of soil pCO2 estimates in paleoecological and paleoenvironmental studies, we compare samples from three Upper Jurassic localities. Soil pCO2 estimates, interpreted as a qualitative indicator of primary paleoproductivity, are used to rank the Late Jurassic terrestrial environments represented by the Morrison Formation in western North America, the informally named Lourinhã formation in Western Europe, and the Stanleyville Group in Central Africa. Because modern terrestrial environments show a positive correlation between primary productivity and faunal richness, a similar relationship is expected in ancient ecosystems. When the relative paleoproductivity levels inferred for each study area are compared with estimates of dinosaur generic richness, a positive correlation emerges. Both the Morrison and Lourinhã formations have high inferred productivity levels and high estimated faunal richness. In contrast, the Stanleyville Group appears to have had low primary productivity and low faunal richness. Paleoclimatic data available for each study area indicate that both productivity and faunal richness are positively linked to water availability, as observed in modern terrestrial ecosystems.

Myers, T. S., Mateus O., Polcyn M. J., Vineyard D., & Jacobs L. L. (2016).  A new chelonioid turtle from the Paleocene of Cabinda, Angola. Annual Meeting of the Society of Vertebrate Paleontology. 194.: Journal of Vertebrate Paleontology, Program and Abstracts, 2016 Abstractmyers_etal_2016_chelonoid_angola_turtle_svp_abstract.pdf

We report a new chelonioid turtle on the basis of a nearly complete skull collected in lower Paleocene, shallow marine deposits, equivalent to the offshore Landana Formation, near the town of Landana in Cabinda Province, Angola. Chelonioid material previously reported from this locality is likely referable to this new taxon. The well-preserved skull is missing the left quadrate, squamosal, and prootic, both opisthotics, and the mandible. The skull possesses a rod-like basisphenoid rostrum, which is a synapomorphy of Chelonioidea, but it differs from other chelonioid skulls in that the contact between the parietal and squamosal is absent, and the posterior palatine foramen is present. Phylogenetic analysis recovers the new taxon as a basal chelonioid. The Paleocene– Eocene strata near Landana have produced a wealth of turtle fossils, including the holotype of the pleurodire Taphrosphys congolensis. A turtle humerus collected from the Landana locality differs morphologically from the humeri of chelonioids and Taphrosphys, indicating the presence of a third taxon. Chelonioid fossil material in the Landana assemblage is rare compared to the abundant fragmentary remains of Taphrosphys that are found throughout the stratigraphic section. This disparity in abundance suggests the new chelonioid taxon preferred open marine habitats, whereas Taphrosphys frequented nearshore environments.

Myers, T. S., Mateus O., Polcyn {M. J. }, Vineyard D., & Jacobs L. L. (2016).  A new chelonioid turtle from the Paleocene of Cabinda, Angola. Abstract

We report a new chelonioid turtle on the basis of a nearly complete skull collected in lower Paleocene, shallow marine deposits, equivalent to the offshore Landana Formation, near the town of Landana in Cabinda Province, Angola. Chelonioid material previously reported from this locality is likely referable to this new taxon. The well-preserved skull is missing the left quadrate, squamosal, and prootic, both opisthotics, and the mandible. The skull possesses a rod-like basisphenoid rostrum, which is a synapomorphy of Chelonioidea, but it differs from other chelonioid skulls in that the contact between the parietal and squamosal is absent, and the posterior palatine foramen is present. Phylogenetic analysis recovers the new taxon as a basal chelonioid. The Paleocenetextendash Eocene strata near Landana have produced a wealth of turtle fossils, including the holotype of the pleurodire Taphrosphys congolensis. A turtle humerus collected from the Landana locality differs morphologically from the humeri of chelonioids and Taphrosphys, indicating the presence of a third taxon. Chelonioid fossil material in the Landana assemblage is rare compared to the abundant fragmentary remains of Taphrosphys that are found throughout the stratigraphic section. This disparity in abundance suggests the new chelonioid taxon preferred open marine habitats, whereas Taphrosphys frequented nearshore environments.

Myers, T. S., Tabor N. J., Jacobs L. L., & Mateus O. (2012).  Palaeoclimate of the Late Jurassic of Portugal: Comparison with the Western United States. Sedimentology. 59(6), 1695–1717., 2012//01/ Abstractmyers_et_al_2012_palaeoclimate_of_the_late_jurassic_of_portugal_comparison_with_the_western.pdfWebsite

Investigation of the palaeoclimatic conditions associated with Upper Jurassic strata in Portugal and comparison with published palaeoclimate reconstructions of the Upper Jurassic Morrison Formation in western North America provide important insights into the conditions that allowed two of the richest terrestrial faunas of this period to flourish. Geochemical analyses and observations of palaeosol morphology in the informally named Upper Jurassic Lourinhã formation of western Portugal indicate warm and wet palaeoclimatic conditions with strongly seasonal precipitation patterns. Palaeosol profiles are dominated by carbonate accumulations and abundant shrink-swell (vertic) features that are both indicative of seasonal variation in moisture availability. The δ18OSMOW and δDSMOW values of phyllosilicates sampled from palaeosol profiles range from +22·4‰ to +22·7‰ and −53·0‰ to −37·3‰, respectively. These isotope values correspond to temperatures of formation between 32°C and 39°C ± 3°, with an average of 36°C, which suggest surface temperatures between 27°C and 34°C (average 31°C). On average, these surface temperature estimates are 1°C higher than the highest summer temperatures modelled for Late Jurassic Iberia using general circulation models. Elemental analysis of matrix material from palaeosol B-horizons provides proxy (chemical index of alteration minus potassium) estimates of mean annual precipitation ranging from 766 to 1394 mm/year, with an average of approximately 1100 mm/year. Palaeoclimatic conditions during deposition of the Lourinhã formation are broadly similar to those inferred for the Morrison Formation, except somewhat wetter. Seasonal variation in moisture availability does not seem to have negatively impacted the ability of these environments to support rich and relatively abundant faunas. The similar climate between these two Late Jurassic terrestrial ecosystems is probably one of the factors which explains the similarity of their vertebrate faunas.

Muchagata, J., & Mateus O. (2016).  Sexual display and rostral variation in extinct beaked whale, Globicetus hiberus. XIV EAVP Meeting. 136., Haarlem, The Netherlands Abstract
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Muchagata, J., & Mateus O. (2016).  Sexual display and rostral variation in extinct beaked whale, Globicetus hiberus. : XIV EAVP Meeting, Programme and Abstract Book Abstract
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Muchagata, J., Mateus O., & Prieto R. (2018).  Treasures from the depths: first record of Tusciziphius (fossil beaked whale) from the Azores islands deep waters and the importance of local communities to science. XVI Annual Meeting of the European Association of Vertebrate Palaeontologists. 132., Caparica, Portugal June 26th-July 1st, 2018 Abstractmuchagata_et_al_2018_eavp_abstract.pdf

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Muchagata, J., & Mateus O. (2016).  Sexual display and rostral variation in extinct beaked whale, Globicetus hiberus. XIV EAVP Meeting. 136., Haarlem, The Netherlands: XIV EAVP Meeting, Programme and Abstract Book Abstractmuchagata_et_al_2016_eavp_abstractbook_finalversionjjl_26062016-1.pdf

Iberian extinct ziphiid, Globicetus hiberus, bears a peculiar large bony sphere in the rostrum, the Mesorostral Process of the Premaxillae or MPP. The MPP varies in size and shape of growth in the six specimens studied and seems to have an allometrically growth in one subgroup, but not in the other, suggesting subgroups correspond to males and females (sexual dimorphism). Even more, some rostral structures, such as the medial pad of the premaxillae seem to be associated with the specimens with lower and leaner MPP’s and ossification of the mesorostral canal by the vomer can also be of value in differentiating sex. Beaked whales are deep-diving, echolocation-user odontocetes and able to perceive bones as distinctive echoic images with their sonar; therefore the MPP may work as a secondary sexual organ (“antlers inside” hypothesis by Gol´din, 2014), a mute display structure acting as an “acoustic flag” to be perceived through echolocation by other individuals, giving information about the shape and size of the MPP.

Muchagata, J., & Mateus O. (2016).  Sexual display and rostral variation in extinct beaked whale, Globicetus hiberus. XIV EAVP Meeting. 136., Haarlem, The Netherlands: XIV EAVP Meeting, Programme and Abstract Book Abstract
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Moreno-Azanza, M., Bauluz B., Canudo J. I., & Mateus O. (2017).  The conservative structure of the ornithopod eggshell: electron backscatter diffraction characterization of Guegoolithus turolensis from the Early Cretaceous of Spain. Journal of Iberian Geology. 1-9., 2017 Abstractthe_conservative_structure_of_the_ornithopod_eggshell.pdfWebsite

The Spheroolithidae oospecies Guegoolithus turolensis, putatively attributed to non-hadrosauroid styracosterns was first described in the Barremian of the Iberian Basin, and later reported in the Valanginian–Hauterivian of the Cameros Basin, with both occurrences separated by a few hundred kilometres but by over 10 million years.

Moreno-Azanza, M., Gasca J. M., Canudo J. I., L\áżaro B. B., & Mateus O. (2016).  The evolution of the ornithischian eggshell: State of the art and perspectives. Abstracts with Programs, the 2016 Annual Meeting, the Palaeontological Society of Japan. 6., Fukui, Japan Abstract
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Moreno-Azanza, M., Gasca J. M., Canudo J. I., Lázaro B. B., & Mateus O. (2016).  The evolution of the ornithischian eggshell: State of the art and perspectives. Abstracts with Programs, the 2016 Annual Meeting, the Palaeontological Society of Japan. 6., Fukui, Japanmoreno_azanza_et_al_2016_the_evolution_of_the_ornithischian_eggshell.pdf
Moreno-Azanza, M., Pérez-Pueyo M., Puértolas-Pascual E., Núñez-Lahuerta C., Mateus O., Bauluz B., Bádenas B., & Canudo J. I. (2022).  A new crocodylomorph related ootaxon from the late Maastrichtian of the Southern Pyrenees (Huesca, Spain). Historical Biology. 1-10.: Taylor & Francis Abstracta_new_crocodylomorph_related_ootaxon_from_the_late_maastrichtian_of_the_southern_pyrenees_huesca_spain.pdfWebsite

ABSTRACTCrocodylomorph eggs and eggshells are known as old as the Late Jurassic and are frequent components of most multiootaxic eggshell assemblages. Classified within the oofamily Krokolithidae, thei histo- and ultrastructures are conservative throughout geological time, characterised by inverted-trapezoid-shaped shell units that grow from highly spaced basal knobs and present a diagnostic tabular ultrastructure. Here, we report 327 eggshell fragments from a new fossil site from the Maastrichtian of the Southern Pyrenees, Veracruz 1, and erect a new oogenus and oospecies, Pachykrokolithus excavatum oogen. et oosp. nov. characterised by crocodyloid morphotype and a prominent rugosocavate ornamentation. Eggshells from the slightly older locality of Blasi 2b, previously reported as aff. Krokolithidae, are also assigned to this new ootaxon. Different crocodylomorph taxa coexisted during the Late Cretaceous of the Tremp Basin, hindering the attribution of Pachykrokolithus excavatum oogen. et oosp. nov. to a single clade. Nevertheless, allodaposuchid eusuchians were dominant in this ecosystem, and are the most probable producers of Pachykrokolithus excavatum oogen. et oosp. nov. eggs.

Moreno-Azanza, M., Ezquerro L., Gasca J. M., Díaz-Berenguer E., Nuñez-Lahuerta C., Pérez-Pueyo M., Puértolas-Pascual E., Sellés A. G., Vila B., & Laita E. (2021).  Two latest Cretaceous egg localities in the external sierras (Southern Pyrenees, Huesca Province, NE Spain). Palaeovertebrata - XVIII annual conference of the European Association of Vertebrate Palaeontologists. 44, 120-121. Abstractmoreno-azanza_et_al_2021_eggs_eavp_abstract.pdf

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Moreno-Azanza, M., Gasca J. M., Canudo J. I., Lázaro B. B., & Mateus O. (2016).  The evolution of the ornithischian eggshell: State of the art and perspectives. : Palaeontological Society of Japan Abstract
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Moreno-Azanza, M., Pérez-Pueyo M., Puértolas-Pascual E., Núñez-Lahuerta C., Mateus O., Bauluz B., Bádenas B., & Canudo J. I. (2022).  Cáscaras de huevo de los últimos cocodrilomorfos del Cretácico (Huesca, España). XXXVII Jornadas de Paleontología SEP - V Congreso Ibérico de Paleontología. p. 119. Abstract2022_moreno-azanza_etal_sep.pdf

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Moreno-Azanza, M., Mateus O., Bauluz B., Coimbra R., Ezquerro L., & Núñez-Lahuerta C. (2021).  Hatching in Portugal: a new look to old eggs. XIX Encontro de Jovens Investigadores em Paleontologia. 22. Abstractmoreno-azanza_et_al_2021_eggs_ejip.pdf

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Moreno-Azanza, M., Gasca J. M., Canudo J. I., Lázaro B. B., & Mateus O. (2016).  The evolution of the ornithischian eggshell: State of the art and perspectives. Abstracts with Programs, the 2016 Annual Meeting, the Palaeontological Society of Japan. 6., Fukui, Japan Abstract
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Moreno-Azanza, M., Balauz B., Ezquerro L., Coimbra R., Mateus O., & Nuñez-Lahuerta C. (2022).  Unbroken dinosaur eggs accumulated together do not guarantee autochthony of the fossil assemblage. Journal of Taphonomy. 16(1-4), 129-130. Abstractmoreno-azanza_et_al_2020_unbroken_eggs.pdf

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Moreno-Azanza, M., Bauluz B., Canudo J. I., & Mateus O. (2017).  The conservative structure of the ornithopod eggshell: electron backscatter diffraction characterization of Guegoolithus turolensis from the Early Cretaceous of Spain. Journal of Iberian Geology. 43, 235–243., jun, Number 2: Springer Nature AbstractWebsite
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Mil\à\}n, J., Mateus O., Marzola M., & Clemmensen L. B. (2016).  Plesiosaur remains from the Lower Jurassic part of the Kap Steward Formation, Jameson Land, East Greenland \–\} evidence of the earliest marine incursion. 60th Annual Meeting Palaeontological Association. 91-92., Lyon, France Abstract
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Milàn, J., & Mateus O. (2024).  Dinosaurfund fra Jylland. Magasinet Naturen. 2024(1), 52-55.milan__mateus_2024_-_jydske_dinosaurer.pdfWebsite
Milàn, J., Clemmensen L. B., Adolfssen J. S., Estrup E. J., Frobøse N., Klein N., Mateus O., & Wings O. (2012).  A preliminary report on coprolites from the Late Triassic part of the Kap Stewart Formation, Jameson Land, East Greenland. New Mexico Museum of Natural History and Science, Bulletin. 57, 203-205. Abstractmilan_et_al_2012_greenland_coprolites_triassic.pdf

The basal part of the Triassic-Jurassic (Rhaetian-Sinemurian) Kap Stewart Formation, exposed at Jameson Land, East Greenland, yields an extensive coprolite collection from black, parallel-laminated mudstone (“paper shale”), representing an open lacustrine system. Preliminary investigations show three different types of coprolites: elongated cylindrical masses, composed of irregularly wrapped layers; elongated cylindrical masses with constriction marks; and spirally-coiled specimens.

Milan, J., Christiansen P., & Mateus O. (2005).  A three-dimensionally preserved sauropod manus impression from the Upper Jurassic of Portugal: Implications for sauropod manus shape and locomotor mechanics. Kaupia. 14, 47-52. Abstractmilan_et_al_mateus_2005_-_3d_sauropod_manus_track_l_j_portugal.pdfWebsite

Sauropods were the largest animals ever to walk the earth, and evolved several specializations in their limbs in order to support their body mass. Their legs became columnar and their manual digits became reduced and encapsulated in tissue to form a single weight-bearing unit in the derived sauropods. A new three-dimensionally preserved cast of a sauropod manus, found in the Upper Jurassic Lourinhã Formation, Portugal, demonstrates not only the shape, but also the actual movements of the sauropod manus during the stride. The manus cast is 32 cm deep, and show the manus to be hoof-shaped and lacking any impressions of individual digits, except for digit I, the pollex. Well preserved striations from skin on the sides of the cast show that the manus was covered in rough, tubercular skin. The width of the manus cast is consistent from top to bottom, demonstrating that the manus was brought down and lifted vertically before any parasaggital movement of the upper limb took place.

Milàn, J., & Mateus O. (2023).  To nye dinosaurfund fra Hasle. Natur På Bornholm. 2023, 74-76.milan__mateus_2023_-_to_nye_dinosaurfund_fra_hasle.pdf
Milàn, J., Mateus O., Marzola M., & Clemmensen L. B. (2016).  Plesiosaur remains from the Lower Jurassic part of the Kap Steward Formation, Jameson Land, East Greenland – evidence of the earliest marine incursion. 60th Annual Meeting Palaeontological Association. 91-92., Lyon, France: Palaeontological Associationmilan_et_al_2016__-_kap_stewart_fm_plesiosaur_-_palass_2016.pdf
Milàn, J., Mateus O., Marzola M., & Clemmensen L. B. (2016).  Plesiosaur remains from the Lower Jurassic part of the Kap Steward Formation, Jameson Land, East Greenland – evidence of the earliest marine incursion. 60th Annual Meeting Palaeontological Association. 91-92., Lyon, France: Palaeontological Association Abstract
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Milan, J., & Mateus O. (2005).  Jagten på Europas største dinosaur.. Naturens Verden. 88(10), 2-13.. AbstractWebsite

[In danish. Title translation: The hunt for the biggest dinosaur in Europe]

Milan, J., & Mateus O. (2005).   Jagten på Europas største dinosaur. Naturens Verden. 88(10), 2-13. Abstract
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Milàn, J., & Mateus O. (2024).  A Turiasaurian (Dinosauria, Sauropoda) Tooth from the Pliensbachian Hasle Formation of Bornholm, Denmark, Shows an Early Jurassic Origin of the Turiasauria. Diversity. 16, , Number 1 Abstractmilan__mateus_2023_diversity-16-00012.pdfWebsite

Turiasauria is a clade of basal sauropod dinosaurs hitherto only known from the Middle Jurassic (Bathonian) to the Lower Cretaceous (Valanginian). A new find of a shed tooth crown from the Lower Jurassic (Pliensbachian), Halse Formation of Bornholm, Denmark, is spoon-like, asymmetrical, and heart-shaped, which identifies the tooth as turiasaurian, pushing the origin of the Turiasauria some 17 My back into the Lower Jurassic. This suggests a North Pangean/Laurasian origin of the turiasaurian clade, which then, during the Middle to Late Jurassic, dispersed through Europe, India, and Africa, with their latest representatives found in the Early Cretaceous of England and North America. Furthermore, this is the first record of a sauropod from the Pliensbachian in Europe.

Milàn, J., Mateus O., Mau M., Rudra A., Sanei H., & Clemmensen L. B. (2021).  A possible phytosaurian (Archosauria, Pseudosuchia) coprolite from the Late Triassic Fleming Fjord Group of Jameson Land, central East Greenland. Bulletin of the Geological Society of Denmark. 69, 71-80. Abstractmilan_et_al_2021_coprolites_greenland_bull69-71-80.pdfWebsite

A large, well-preserved vertebrate coprolite was found in a lacustrine sediment in the Malmros Klint Formation of the Late Triassic Fleming Fjord Group in the Jameson Land Basin, central East Greenland. The size and internal and external morphology of the coprolite is consistent with that of crocodilian coprolites and one end of the coprolite exhibits evidence of post-egestion trampling. As the associated vertebrate fauna of the Fleming Fjord Group contains abundant remains of pseudosuchian phytosaurs, the coprolite is interpreted as being from a large phytosaur.

Milàn, J., & Mateus O. (2003).  Fra strandbred til museum p{\aa} syv dage: historien om et gigantisk dinosaur fodspor. Varv. 8–14., Number 2003/3 Abstract
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Mateus, O., & Antunes M. T. (2003).  A New Dinosaur Tracksite in the Lower Cretaceous of Portugal. Ciências da Terra. 15, 253-262. Abstractmateus__antunes_2003-_early_k_dino_tracks_portugal.pdfWebsite

A new Lower Cretaceous (Aptian-Albian) dinosaur tracksite at the Olhos de Água beach is described. It is the first vertebrate fossil finding ever found in the concerned unit, and yielded 128 tracks in 17 trackways within an area of ca. 80 square metres.
Three tridactyl footprint morphotypes have been recognized: - Type 1 (“Iguanodontipus-like”) - trackways D, F, K, J and P; - Type 2 (large theropod), although larger in size, typically from a Grallator-like theropod footprint, i.e. A, B, G, H and O trackways; - Type 3 (medium size theropod); M is the only track of this type. There are other, poorly preserved, unidentified trackways. The theropod, swinging trackway B was produced by an animal that was limping. The theropod track M starts eastwards but drastically changes westwards, speeding up at the same time; this dinosaur decided to turn around and run in the opposite direction.
This site shows three main trackway directions: to the South, to the East, and westwards. Except for the trackway O, large theropods A, B, G and H walked southwards. Perpendicularly to these, ornithopods, small theropods and unidentified trackmakers walked towards East (5) and West (7). The segregation of trackmakers and directions, with large theropod trackways southwards and other dinosaurs’ west or eastwards, may mean that large theropods patrolled a walkway area to an important resource, most probably water, often frequented by ornithopods and smaller theropods. There is no evidence of social behavior or gregarism: footprints’ overposition shows that the large, southwards walking theropods passed on different occasions. Three trackway sequences can be established by chronologic order.

Mateus, O., Maidment S. C. R., & Christiansen N. A. (2009).  A new long-necked {'}sauropod-mimic{'} stegosaur and the evolution of the plated dinosaurs. Proceedings of the Royal Society B: Biological Sciences. 276, 1815-1821., Number 1663 Abstract
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Mateus, O., & Tschopp E. (2013).  Cathetosaurus as a valid sauropod genus and comparisons with Camarasaurus. Journal of Vertebrate Paleontology, Program and Abstracts, 2013. 173.mateus__tschopp_2013_cathetosaurs_camarasaurus__svp_meeting_abstracts_213.pdf
Mateus, O. (2013).  Cabinda revisited: age and environment of new Cenozoic vertebrate fossils from northern Angola. 45, , 1 Abstract

In the early 20thcentury, Belgian naturalists reported Paleocene and Eocene sharks, the bothremydid pleurodiran turtleTaphosphrys(formerlyBantuchelys), and a neosuchian and the dyrosaurid crocodyliformCongosaurusfrom coastal outcrops near Landana in the northern province of Cabinda, Angola. In 1935, rare and fragmentary mammals were reported from strata at Malembo Point, south of Landana, and originally considered to be Miocene in age. Subsequent re-identification of the mammal fossils suggested that the Malembo deposits are Lower Oligocene based on the presence of an arsinoithere and hyracoids. An anthropoid canine, originally identified as a carnivore incisor, was also recognized. In four recent expeditions to Cabinda, Projecto PaleoAngola has traced and measured the stratigraphy between Landana and Malembo Point and sampled these deposits for pollen, stable isotopes, detrital zircons, paleomagnetic stratigraphy, and for U/Pb dating of bones, teeth, and coprolites. Although the faunas from all Cabinda localities are dominated by sharks and rays, new discoveries from Landana include a complete cheloniid cryptodire turtle skull, a small snake vertebra, and a bird bone. Discoveries from the Malembo level include a narrow-snouted crocodyliform similar toCongosaurusandEuthecodon, an arsinoithere anterior tooth, an upper molar similar to that of the ptolemaiidanKelba, an unidentified mammalian incisor, and a large primate-like premolar. Recent biostratigraphic advances in East Africa and the new fossil discoveries in Cabinda suggest similarity to late Oligocene faunas in Ethiopia and Kenya, although the large primate-like premolar is unique. To this day, the fossil localities of Malembo provide the only coastal, low latitude, low elevation record of West African Cenozoic terrestrial mammals. https://gsa.confex.com/gsa/2013AM/webprogram/Paper232115.html

Mateus, O., & Antunes M. T. (2008).  Landmarks in the history of dinosaur paleontology in Portugal, focusing on skeletal remains. Abstract volume, Dinosaurs - A Historical Perspective, 6-7 may 2008. , London Abstract
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Mateus, O., Jacobs L. L., Polcyn M. J., Myers T. S., & Schulp A. S. (2015).  The fossil record of testudines from angola from the turonian to oligocene. Society of Vertebrate Paleontology Annual Meeting. 177., Dallasmateus_et_al_2015_testudines_angola_svp_abstract.pdf
Mateus, O. (2011).  Evolutionary major trends of ornithopod dinosaurs teeth. (Unknown Unknown, Ed.).Dinosaurios y paleontología desde América Latina. 25–31., 1: EDIUNC, Editorial de la Universidad Nacional de Cuyo Abstract
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