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Myers, T. S., Mateus O., Polcyn M. J., Vineyard D., & Jacobs L. L. (2016).  A new chelonioid turtle from the Paleocene of Cabinda, Angola. Journal of Vertebrate Paleontology, Program and Abstracts, 2016, p. 194.. 194. Abstract

We report a new chelonioid turtle on the basis of a nearly complete skull collected in lower Paleocene, shallow marine deposits, equivalent to the offshore Landana Formation, near the town of Landana in Cabinda Province, Angola. Chelonioid material previously reported from this locality is likely referable to this new taxon. The well-preserved skull is missing the left quadrate, squamosal, and prootic, both opisthotics, and the mandible. The skull possesses a rod-like basisphenoid rostrum, which is a synapomorphy of Chelonioidea, but it differs from other chelonioid skulls in that the contact between the parietal and squamosal is absent, and the posterior palatine foramen is present. Phylogenetic analysis recovers the new taxon as a basal chelonioid. The Paleocene– Eocene strata near Landana have produced a wealth of turtle fossils, including the holotype of the pleurodire Taphrosphys congolensis. A turtle humerus collected from the Landana locality differs morphologically from the humeri of chelonioids and Taphrosphys, indicating the presence of a third taxon. Chelonioid fossil material in the Landana assemblage is rare compared to the abundant fragmentary remains of Taphrosphys that are found throughout the stratigraphic section. This disparity in abundance suggests the new chelonioid taxon preferred open marine habitats, whereas Taphrosphys frequented nearshore environments.

Myers, T. S., Tabor N. J., Jacobs L. L., & Mateus O. (2012).  Estimating soil pCO2 using paleosol carbonates: implications for the relationship between primary productivity and faunal richness in ancient terrestrial ecosystems. Paleobiology. 38(4), 585–604. Abstractmyers_et_al_2012_estimating_soil_paleosols_portugal.pdf

In this paper we present a method for estimating soil pCO2 in ancient environments using the measured carbon-isotope values of pedogenic carbonates and plant-derived organic matter. The validity of soil pCO2 estimates proves to be highly dependent on the organic δ13C values used in the calculations. Organic matter should be sourced from the same paleosol profiles as sampled carbonates to yield the most reliable estimates of soil pCO2. In order to demonstrate the potential use of soil pCO2 estimates in paleoecological and paleoenvironmental studies, we compare samples from three Upper Jurassic localities. Soil pCO2 estimates, interpreted as a qualitative indicator of primary paleoproductivity, are used to rank the Late Jurassic terrestrial environments represented by the Morrison Formation in western North America, the informally named Lourinhã formation in Western Europe, and the Stanleyville Group in Central Africa. Because modern terrestrial environments show a positive correlation between primary productivity and faunal richness, a similar relationship is expected in ancient ecosystems. When the relative paleoproductivity levels inferred for each study area are compared with estimates of dinosaur generic richness, a positive correlation emerges. Both the Morrison and Lourinhã formations have high inferred productivity levels and high estimated faunal richness. In contrast, the Stanleyville Group appears to have had low primary productivity and low faunal richness. Paleoclimatic data available for each study area indicate that both productivity and faunal richness are positively linked to water availability, as observed in modern terrestrial ecosystems.

Myers, T. S., Tabor N. J., Jacobs L. L., & Mateus O. (2012).  Palaeoclimate of the Late Jurassic of Portugal: Comparison with the Western United States. Sedimentology. 59(6), 1695–1717., 2012//01/ Abstractmyers_et_al_2012_palaeoclimate_of_the_late_jurassic_of_portugal_comparison_with_the_western.pdfWebsite

Investigation of the palaeoclimatic conditions associated with Upper Jurassic strata in Portugal and comparison with published palaeoclimate reconstructions of the Upper Jurassic Morrison Formation in western North America provide important insights into the conditions that allowed two of the richest terrestrial faunas of this period to flourish. Geochemical analyses and observations of palaeosol morphology in the informally named Upper Jurassic Lourinhã formation of western Portugal indicate warm and wet palaeoclimatic conditions with strongly seasonal precipitation patterns. Palaeosol profiles are dominated by carbonate accumulations and abundant shrink-swell (vertic) features that are both indicative of seasonal variation in moisture availability. The δ18OSMOW and δDSMOW values of phyllosilicates sampled from palaeosol profiles range from +22·4‰ to +22·7‰ and −53·0‰ to −37·3‰, respectively. These isotope values correspond to temperatures of formation between 32°C and 39°C ± 3°, with an average of 36°C, which suggest surface temperatures between 27°C and 34°C (average 31°C). On average, these surface temperature estimates are 1°C higher than the highest summer temperatures modelled for Late Jurassic Iberia using general circulation models. Elemental analysis of matrix material from palaeosol B-horizons provides proxy (chemical index of alteration minus potassium) estimates of mean annual precipitation ranging from 766 to 1394 mm/year, with an average of approximately 1100 mm/year. Palaeoclimatic conditions during deposition of the Lourinhã formation are broadly similar to those inferred for the Morrison Formation, except somewhat wetter. Seasonal variation in moisture availability does not seem to have negatively impacted the ability of these environments to support rich and relatively abundant faunas. The similar climate between these two Late Jurassic terrestrial ecosystems is probably one of the factors which explains the similarity of their vertebrate faunas.

Myers, T. S., Mateus O., Polcyn M. J., Vineyard D., & Jacobs L. L. (2016).  A new chelonioid turtle from the Paleocene of Cabinda, Angola. Annual Meeting of the Society of Vertebrate Paleontology. 194.: Journal of Vertebrate Paleontology, Program and Abstracts, 2016 Abstractmyers_etal_2016_chelonoid_angola_turtle_svp_abstract.pdf

We report a new chelonioid turtle on the basis of a nearly complete skull collected in lower Paleocene, shallow marine deposits, equivalent to the offshore Landana Formation, near the town of Landana in Cabinda Province, Angola. Chelonioid material previously reported from this locality is likely referable to this new taxon. The well-preserved skull is missing the left quadrate, squamosal, and prootic, both opisthotics, and the mandible. The skull possesses a rod-like basisphenoid rostrum, which is a synapomorphy of Chelonioidea, but it differs from other chelonioid skulls in that the contact between the parietal and squamosal is absent, and the posterior palatine foramen is present. Phylogenetic analysis recovers the new taxon as a basal chelonioid. The Paleocene– Eocene strata near Landana have produced a wealth of turtle fossils, including the holotype of the pleurodire Taphrosphys congolensis. A turtle humerus collected from the Landana locality differs morphologically from the humeri of chelonioids and Taphrosphys, indicating the presence of a third taxon. Chelonioid fossil material in the Landana assemblage is rare compared to the abundant fragmentary remains of Taphrosphys that are found throughout the stratigraphic section. This disparity in abundance suggests the new chelonioid taxon preferred open marine habitats, whereas Taphrosphys frequented nearshore environments.

Myers, T. S., Mateus O., Polcyn M. J., Vineyard D., & Jacobs L. L. (2016).  A new chelonioid turtle from the Paleocene of Cabinda, Angola. Journal of Vertebrate Paleontology, Program and Abstracts, 2016, p. 194.. 194. Abstract
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Myers, T. S., Tabor N. J., Jacobs L. L., & Mateus O. (2012).  Palaeoclimate of the late jurassic of portugal: Comparison with the western united states. Sedimentology. 59, 1695-1717., Number 6 Abstract
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Myers, T. S., Polcyn M. J., Mateus O., Vineyard D. P., Gonçalves A. O., & Jacobs L. L. (2017).  A new durophagous stem cheloniid turtle from the lower Paleocene of Cabinda, Angola. Papers in Palaeontology. 2017, 1-16. Abstractnew_durophagous_stem_cheloniid_turtle_from_the_lower_paleocene_of_cabinda_angola.pdfWebsite

A new stem cheloniid turtle, Cabindachelys landanensis, gen. et sp. nov., is represented by a nearly complete skull and partial hyoid collected in lower Paleocene shallow marine deposits, equivalent to the offshore Landana Formation, near the town of Landana in Cabinda, Angola. A partial chelonioid carapace previously reported from this locality is referred here to C. landanensis. Cabindachelys landanensis possesses clear synapomorphies of Pan-Cheloniidae, including a rod-like rostrum basisphenoidale, V-shaped basisphenoid crest, and secondary palate, but also retains a reduced foramen palatinum posterius, unlike most other pan-cheloniids. Phylogenetic analysis suggests that C. landanensis forms a weakly-supported clade with Erquelinnesia gosseleti, Euclastes acutirostris, Euclastes wielandi and Terlinguachelys fischbecki, although a close relationship between the protostegid T. fischbecki and these durophagous pan-cheloniids is unlikely. The Paleocene–Eocene strata near Landana have produced a number of turtle fossils, including the holotype specimen of the pleurodire Taphrosphys congolensis. A turtle humerus collected c. 1 m above the holotype skull of C. landanensis differs from humeri of chelonioids and Taphrosphys, indicating that a third turtle taxon is present at Landana. Cheloniid fossil material is rare in the Landana assemblage, in comparison with the abundant remains of Taphrosphys congolensis found throughout the stratigraphic section. This disparity implies that C. landanensis preferred open marine habitats, whereas Taphrosphys congolensis spent more time in nearshore environments. The appearance of new durophagous species such as C. landanensis in the early Paleocene reflects the rapid radiation of pan-cheloniids as they diversified into open niches following the K–Pg extinction.

Myers, T. S., Polcyn M. J., Mateus O., Vineyard D. P., Gon?alves A. O., & Jacobs L. L. (2018).  A new durophagous stem cheloniid turtle from the lower Paleocene of Cabinda, Angola. Papers in Palaeontology. 4, 161-176., Number 2 Abstract
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Myers, T. S., Mateus O., Polcyn {M. J. }, Vineyard D., & Jacobs L. L. (2016).  A new chelonioid turtle from the Paleocene of Cabinda, Angola. Abstract

We report a new chelonioid turtle on the basis of a nearly complete skull collected in lower Paleocene, shallow marine deposits, equivalent to the offshore Landana Formation, near the town of Landana in Cabinda Province, Angola. Chelonioid material previously reported from this locality is likely referable to this new taxon. The well-preserved skull is missing the left quadrate, squamosal, and prootic, both opisthotics, and the mandible. The skull possesses a rod-like basisphenoid rostrum, which is a synapomorphy of Chelonioidea, but it differs from other chelonioid skulls in that the contact between the parietal and squamosal is absent, and the posterior palatine foramen is present. Phylogenetic analysis recovers the new taxon as a basal chelonioid. The Paleocenetextendash Eocene strata near Landana have produced a wealth of turtle fossils, including the holotype of the pleurodire Taphrosphys congolensis. A turtle humerus collected from the Landana locality differs morphologically from the humeri of chelonioids and Taphrosphys, indicating the presence of a third taxon. Chelonioid fossil material in the Landana assemblage is rare compared to the abundant fragmentary remains of Taphrosphys that are found throughout the stratigraphic section. This disparity in abundance suggests the new chelonioid taxon preferred open marine habitats, whereas Taphrosphys frequented nearshore environments.

Muchagata, J., & Mateus O. (2016).  Sexual display and rostral variation in extinct beaked whale, Globicetus hiberus. XIV EAVP Meeting. 136., Haarlem, The Netherlands Abstract
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Muchagata, J., Mateus O., & Prieto R. (2018).  Treasures from the depths: first record of Tusciziphius (fossil beaked whale) from the Azores islands deep waters and the importance of local communities to science. XVI Annual Meeting of the European Association of Vertebrate Palaeontologists. 132., Caparica, Portugal June 26th-July 1st, 2018 Abstractmuchagata_et_al_2018_eavp_abstract.pdf

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Muchagata, J., & Mateus O. (2016).  Sexual display and rostral variation in extinct beaked whale, Globicetus hiberus. XIV EAVP Meeting. 136., Haarlem, The Netherlands: XIV EAVP Meeting, Programme and Abstract Book Abstractmuchagata_et_al_2016_eavp_abstractbook_finalversionjjl_26062016-1.pdf

Iberian extinct ziphiid, Globicetus hiberus, bears a peculiar large bony sphere in the rostrum, the Mesorostral Process of the Premaxillae or MPP. The MPP varies in size and shape of growth in the six specimens studied and seems to have an allometrically growth in one subgroup, but not in the other, suggesting subgroups correspond to males and females (sexual dimorphism). Even more, some rostral structures, such as the medial pad of the premaxillae seem to be associated with the specimens with lower and leaner MPP’s and ossification of the mesorostral canal by the vomer can also be of value in differentiating sex. Beaked whales are deep-diving, echolocation-user odontocetes and able to perceive bones as distinctive echoic images with their sonar; therefore the MPP may work as a secondary sexual organ (“antlers inside” hypothesis by Gol´din, 2014), a mute display structure acting as an “acoustic flag” to be perceived through echolocation by other individuals, giving information about the shape and size of the MPP.

Muchagata, J., & Mateus O. (2016).  Sexual display and rostral variation in extinct beaked whale, Globicetus hiberus. : XIV EAVP Meeting, Programme and Abstract Book Abstract
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Muchagata, J., & Mateus O. (2016).  Sexual display and rostral variation in extinct beaked whale, Globicetus hiberus. XIV EAVP Meeting. 136., Haarlem, The Netherlands: XIV EAVP Meeting, Programme and Abstract Book Abstract
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Moreno-Azanza, M., Gasca J. M., Canudo J. I., Lázaro B. B., & Mateus O. (2016).  The evolution of the ornithischian eggshell: State of the art and perspectives. Abstracts with Programs, the 2016 Annual Meeting, the Palaeontological Society of Japan. 6., Fukui, Japanmoreno_azanza_et_al_2016_the_evolution_of_the_ornithischian_eggshell.pdf
Moreno-Azanza, M., Bauluz B., Canudo J. I., & Mateus O. (2017).  The conservative structure of the ornithopod eggshell: electron backscatter diffraction characterization of Guegoolithus turolensis from the Early Cretaceous of Spain. Journal of Iberian Geology. 43, 235–243., jun, Number 2: Springer Nature AbstractWebsite
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Moreno-Azanza, M., Gasca J. M., Canudo J. I., L\áżaro B. B., & Mateus O. (2016).  The evolution of the ornithischian eggshell: State of the art and perspectives. Abstracts with Programs, the 2016 Annual Meeting, the Palaeontological Society of Japan. 6., Fukui, Japan Abstract
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Moreno-Azanza, M., Gasca J. M., Canudo J. I., Lázaro B. B., & Mateus O. (2016).  The evolution of the ornithischian eggshell: State of the art and perspectives. : Palaeontological Society of Japan Abstract
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Moreno-Azanza, M., Gasca J. M., Canudo J. I., Lázaro B. B., & Mateus O. (2016).  The evolution of the ornithischian eggshell: State of the art and perspectives. Abstracts with Programs, the 2016 Annual Meeting, the Palaeontological Society of Japan. 6., Fukui, Japan Abstract
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Moreno-Azanza, M., Bauluz B., Canudo J. I., & Mateus O. (2017).  The conservative structure of the ornithopod eggshell: electron backscatter diffraction characterization of Guegoolithus turolensis from the Early Cretaceous of Spain. Journal of Iberian Geology. 1-9., 2017 Abstractthe_conservative_structure_of_the_ornithopod_eggshell.pdfWebsite

The Spheroolithidae oospecies Guegoolithus turolensis, putatively attributed to non-hadrosauroid styracosterns was first described in the Barremian of the Iberian Basin, and later reported in the Valanginian–Hauterivian of the Cameros Basin, with both occurrences separated by a few hundred kilometres but by over 10 million years.

Mil\à\}n, J., Mateus O., Marzola M., & Clemmensen L. B. (2016).  Plesiosaur remains from the Lower Jurassic part of the Kap Steward Formation, Jameson Land, East Greenland \–\} evidence of the earliest marine incursion. 60th Annual Meeting Palaeontological Association. 91-92., Lyon, France Abstract
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Milàn, J., Mateus O., Marzola M., & Clemmensen L. B. (2016).  Plesiosaur remains from the Lower Jurassic part of the Kap Steward Formation, Jameson Land, East Greenland – evidence of the earliest marine incursion. 60th Annual Meeting Palaeontological Association. 91-92., Lyon, France: Palaeontological Association Abstract
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Milan, J., Christiansen P., & Mateus O. (2005).  A three-dimensionally preserved sauropod manus impression from the Upper Jurassic of Portugal: Implications for sauropod manus shape and locomotor mechanics. Kaupia. 14, 47-52. Abstractmilan_et_al_mateus_2005_-_3d_sauropod_manus_track_l_j_portugal.pdfWebsite

Sauropods were the largest animals ever to walk the earth, and evolved several specializations in their limbs in order to support their body mass. Their legs became columnar and their manual digits became reduced and encapsulated in tissue to form a single weight-bearing unit in the derived sauropods. A new three-dimensionally preserved cast of a sauropod manus, found in the Upper Jurassic Lourinhã Formation, Portugal, demonstrates not only the shape, but also the actual movements of the sauropod manus during the stride. The manus cast is 32 cm deep, and show the manus to be hoof-shaped and lacking any impressions of individual digits, except for digit I, the pollex. Well preserved striations from skin on the sides of the cast show that the manus was covered in rough, tubercular skin. The width of the manus cast is consistent from top to bottom, demonstrating that the manus was brought down and lifted vertically before any parasaggital movement of the upper limb took place.

Milan, J., & Mateus O. (2005).   Jagten på Europas største dinosaur. Naturens Verden. 88(10), 2-13. Abstract
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Milàn, J., Mateus O., Mau M., Rudra A., Sanei H., & Clemmensen L. B. (2021).  A possible phytosaurian (Archosauria, Pseudosuchia) coprolite from the Late Triassic Fleming Fjord Group of Jameson Land, central East Greenland. Bulletin of the Geological Society of Denmark. 69, 71-80. Abstractmilan_et_al_2021_coprolites_greenland_bull69-71-80.pdfWebsite

A large, well-preserved vertebrate coprolite was found in a lacustrine sediment in the Malmros Klint Formation of the Late Triassic Fleming Fjord Group in the Jameson Land Basin, central East Greenland. The size and internal and external morphology of the coprolite is consistent with that of crocodilian coprolites and one end of the coprolite exhibits evidence of post-egestion trampling. As the associated vertebrate fauna of the Fleming Fjord Group contains abundant remains of pseudosuchian phytosaurs, the coprolite is interpreted as being from a large phytosaur.

Milàn, J., Clemmensen L. B., Adolfssen J. S., Estrup E. J., Frobøse N., Klein N., Mateus O., & Wings O. (2012).  A preliminary report on coprolites from the Late Triassic part of the Kap Stewart Formation, Jameson Land, East Greenland. New Mexico Museum of Natural History and Science, Bulletin. 57, 203-205. Abstractmilan_et_al_2012_greenland_coprolites_triassic.pdf

The basal part of the Triassic-Jurassic (Rhaetian-Sinemurian) Kap Stewart Formation, exposed at Jameson Land, East Greenland, yields an extensive coprolite collection from black, parallel-laminated mudstone (“paper shale”), representing an open lacustrine system. Preliminary investigations show three different types of coprolites: elongated cylindrical masses, composed of irregularly wrapped layers; elongated cylindrical masses with constriction marks; and spirally-coiled specimens.

Milan, J., & Mateus O. (2005).  Jagten på Europas største dinosaur.. Naturens Verden. 88(10), 2-13.. AbstractWebsite

[In danish. Title translation: The hunt for the biggest dinosaur in Europe]

Milàn, J., Mateus O., Marzola M., & Clemmensen L. B. (2016).  Plesiosaur remains from the Lower Jurassic part of the Kap Steward Formation, Jameson Land, East Greenland – evidence of the earliest marine incursion. 60th Annual Meeting Palaeontological Association. 91-92., Lyon, France: Palaeontological Associationmilan_et_al_2016__-_kap_stewart_fm_plesiosaur_-_palass_2016.pdf
Milàn, J., & Mateus O. (2003).  Fra strandbred til museum p{\aa} syv dage: historien om et gigantisk dinosaur fodspor. Varv. 8–14., Number 2003/3 Abstract
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Mateus, O., & Antunes M. T. (2000).  On the presence of Ceratosaurus sp. (Dinosauria: Theropoda) in the Late Jurassic of Portugal. Abstract volume of the 31st International Geological Congress. , Rio de Janeiro, Brazil Abstract
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Mateus, O., & Milan J. (2008).  Ichnological evidence for giant ornithopod dinosaurs in the Upper Jurassic Lourinhã Formation, Portugal. Oryctos. 8, 47-52. Abstractmateus_and_milan_2008_ichnological_evidence_for_giant_ornithopod_big_ornithopod_track_from_u_j_lourinha_fm_portugal.pdfWebsite

The Upper Jurassic Lourinhã Formation (Lusitanian Basin, Portugal) contains a diverse dinosaur fauna comprising theropods, sauropods, stegosaurs, ankylosaurs and several genera of ornithopods. The sedimentology in the area favours preservation of tracksways, and tracks from most of the dinosaurs are also represented by skeletal remains. During fieldwork in the summer of 2003 a new, large, tridactyl track was found at the beach of Vale Frades, approximately 6 km north of Lourinhã (central west Portugal). The track was found together with a stegosaur track on a clay bed exposed within the intertidal zone. Due to the immediate danger of erosion, the track was collected and is now on display at Museu da Lourinhã. The track is 70 cm long and 69 cm wide, the toes are short and broad, with indications of short blunt claws, and there is a high angle of divarication between the outer digits. The shape and dimensions of the track identifies it as deriving from an ornithopod dinosaur with an estimated hip height around three metres. Although very large ornithopods are known from the Cretaceous, the largest known Jurassic ornithopod is Camptosaurus from North America, and the largest known from Portugal is the camptosaurid Draconyx loureiroi. Neither of these reached the body size suggested by the new track. So far the track described herein is the only evidence for a Jurassic ornithopod of that size.

Mateus, I., Mateus H., Antunes M. T., Mateus O., Taquet P., Ribeiro V., & Manuppella G. (1997).  Couvée, œufs et embryons d'un Dinosaure Théropode du Jurassique supérieur de Lourinhã (Portugal). Comptes Rendus de l'Academie de Sciences - Serie IIa: Sciences de la Terre et des Planetes. 325, 71–78., Number 1 Abstract
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Mateus, O., Walen A., & Antunes M. T. (2006).  The large theropod fauna of the Lourinhã Formation (Portugal) and its similarity to the Morrison Formation, with a description of a new species of Allosaurus. New Mexico Museum of Natural History and Science Bulletin. 36, 123-129. Abstractmateus_walen_antunes_-_2006_-_the_large_theropod_fauna_of_the_lourinha_formation__portugal__and_its_similarity_to_the_morrison_formation__with_a_description_of_a_new_species_of_allosaurus.pdf

Late Jurassic theropod dinosaurs have been known in Portugal since 1863 but only now are they being fully understood, with the recognition of genera such as Allosaurus, Aviatyrannis, Ceratosaurus, Lourinhanosaurus, and Torvosaurus from the Lourinhã and Alcobaça Formations (Kimmeridgian/Tithonian). Ceratosaurus dentisulcatus can now be reported from Portugal. It represents the only occurrence of this species outside the Morrison Formation.
New cranial elements confirm the presence of Torvosaurus tanneri, in Portugal. Torvosaurus was the largest Late Jurassic land carnivore. New postcranial and cranial elements allow the erection of a new species from Portugal, Allosaurus europaeus n.sp. The theropod assemblage of Portugal is similar to that of the Morrison Formation.

Mateus, O. (2014).  Eggs and eggshells of crocodylomorpha from the Late Jurassic of Portugal. Journal of Vertebrate Paleontology. Program and Abstracts, 2014, 218., Number NA: Taylor & Francis Abstract
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Mateus, O. (2013).  Crocodylomorphs from the Mesozoic of Portugal and a new skull of eusuchian from the Late Cretaceous. 2013 Hwaseong International Dinosaurs Expedition Symposium, pp.66-67.. , Hwaseong, South Korea Abstractmateus_2013_crocodylomorphs_portugal_new_skull.pdf

The diversity of fossil crocodylomorphs in Portugal is high, with occurrence as old as Mystriosaurus (=Steneosaurus) bollensis from the Lower Jurassic. The Late Jurassic forms are the better documented, and include the following taxa: Machimosaurus hugii, Lisboasaurus estesi Seiffert, 1973, Lusitanisuchus mitrocostatus Seiffert, 1975; Schwarz & Fechner 2004, Theriosuchus guimarotae Schwarz and Salisbury 2005, Cf. Alligatorium, Goniopholis baryglyphaeus, and a crocodylomorph-like eggs in dinosaur nests (Mateus et al., 1998; Ricqlès et al., 2001). From the Lower Cretaceous were reported a few dinosaurs but its record is strangely scarce in crocodylomorphs (Mateus et al., 2011). The Upper Cretaceous crocodiles show a large diversity, but it is mostly based in fragmentary material that require revision, such as “Crocodylus” blavieri? Grey from the Upper Campanian - Maastrichtian of Viso, near Aveiro (initially reported by Sauvage 1897-98), Goniopholis cf. crassidens Owen 1841 and Oweniasuchus pulchelus Jonet 1981. Moreover there is a fascinating, but poorly understood, crocodylomorph diversity in the Cenomanian of Portugal, documented by fragmentary specimens that have been doubtfully assigned to Thoracosaurus Leidy 1852 of the Middle Cenomanian of Cacém, to the nomen dubium Oweniasuchus lusitanicus Sauvage 1897-98 (interpreted as a mesosuchian goniopholid) based in a fragmentary mandible from the Campanian-Maastrichtian, and also from the Middle Cenomanian of Portugal, Buffetaut and Lauverjat (1978) report an fragmentary unidentified possible dyrosaurid from Nazaré. All this specimens are too incomplete to be compared with the specimen here described. In contrast, Cenozoic crocodiles of Portugal are often known after complete skulls and several individuals. The taxa list include Iberosuchus macrodon (Lower to Middle Eocene), Tomistoma calaritanus (Early Miocene) and T. lusitanica (Burdigalian-Helvetian), and Diplocynodon sp. (Antunes, 1961, 1987, 1994).
At least, two different morphotypes of crocodylomorph eggs from the Late Jurassic of Lourinhã Formation are also known.
A new specimen here reported of crocodile based in a partial skull and mandible (ML1818) from the Uppermost Middle Cenomanian platform carbonates of Baixo Mondego, west central Portugal (Tentúgal Fm., Callapez, 2004). The taxon is phylogenetically positioned as a basal Eusuchia, due to the choanae enclosed by the pterygoid, and closely related with stem Crocodylia and Borealosuchus. This specimen represents the only well documented and valid eusuchian species in the Cenomanian of Europe and is the oldest representative of an eusuchian crocodylomorph, with the exception for the Barremian Hylaeochampsa vectiana.

Mateus, O. (2013).  Cathetosaurus as a valid sauropod genus and comparisons with Camarasaurus. Journal of Vertebrate Paleontology. 173., 1 Abstract
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Mateus, O. (2009).  Colecções paleontológicas do Museu da Lourinhã (Portugal) / Paleontological collections of the Museum of Lourinhã (Portugal). (Unknown Unknown, Ed.).Journal of Paleontological Techniques. 18–19.., 1 Abstract
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Mateus, O. (2011).  New fossil whales from Angola. Abstracts of the 71st Annual Meeting of the Society of Vertebrate Paleontology. 119., 1 Abstract
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Mateus, O., Maidment S. C. R., & Christiansen N. A. (2008).  A new specimen aff. Dacentrurus armatus (Dinosauria: Stegosauridae) from the Late Jurassic of Portugal. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. 157–157., Neuquén, Argentina Abstract
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Mateus, O., & Campos H. (2018).  Loulé há mais de 220 Milhões de anos: os vertebrados fósseis do Algarve triásico. Loulé: Territórios. Memórias. Identidades. 651-659.: Museu Nacional de Arqueologia | Imprensa Nacionalmateus_campos2018_algarve_triasico.pdf
Mateus, O., Dyke G., Motchurova-Dekova N., Ivanov P., & Kamenov G. D. (2008).  The Bulgarian dinosaur: did it exist? European late Cretaceous ornithomimosaurs. 56th Symposium of Vertebrate Palaeontology and Comparative Anatomy. 47., Dublin Abstract
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Mateus, O., & Milan J. (2010).  A diverse Upper Jurassic dinosaur ichnofauna from central-west Portugal. Lethaia. 43, 245–257., Jan Abstractmateus__milan_2010_-_diverse_l_j_ichnofauna_from_lourinha_fm_portugal.pdfWebsite

A newly discovered dinosaur track-assemblage from the Upper Jurassic Lourinha˜ Formation (Lusitanian Basin, central-west Portugal), comprises medium- to large-sized sauropod tracks with well-preserved impressions of soft tissue anatomy, stegosaur tracks and tracks from medium- to large-sized theropods. The 400-m-thick Lourinha˜ Formation consists of mostly aluvial sediments, deposited during the early rifting of the Atlantic Ocean in the Kimmeridgian and Tithonian. The stratigraphic succession shows several shifts between flood-plain mud and fluvial sands that favour preservation and fossilization of tracks. The studied track-assemblage is found preserved as natural casts on the underside of a thin bivalve-rich carbonate bed near the Tithonian–Kimmeridgian boundary. The diversity of the tracks from the new track assemblage is compared with similar faunas from the Upper Jurassic of Asturias, Spain and the Middle Jurassic Yorkshire Coast of England. The Portuguese record of Upper Jurassic dinosaur body fossils show close similarity to the track fauna from the Lourinha˜ Formation.

Mateus, O., Laven T., & Knotschke N. (2004).  A dwarf between giants?: A new late Jurassic sauropod from Germany. Journal of Vertebrate Paleontology. 23, 90., Number suppl. to 3 Abstract
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Mateus, O., & Jacinto J. J. (2002).  Contribuição para o estudo de Hemidactylus turcicus (Reptilia, Gekkonidae): ritmos de actividade e microhabitat em Évora, Portugal. VII Congresso Luso-Espanhol de Herpetologia. 136., Évora Abstract

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