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Mateus, O., & Antunes M. T. (2000).  Late Jurassic dinosaurs of Portugal. Abstracts of the 1st Symposium of European Dinosaurs. , Dusseldorf, Germany. Abstract
Mateus, I., Mateus H., Antunes M. T., Mateus O., Taquet P., Ribeiro V., & Manuppella G. (1997).  Couvée, øe}ufs et embryons d{\textquotesingle}un Dinosaure Théropode du Jurassique supérieur de Lourinha (Portugal). Comptes Rendus de l{\textquotesingle}Académie des Sciences - Series {IIA} - Earth and Planetary Science. 325, 71–78., jul, Number 1: Elsevier {BV} AbstractWebsite
Mateus, O., & Milàn J. (2011).  New dinosaur and pterosaur tracksites from the Late Jurassic of Portugal. Dinosaur Tracks 2011 An International Symposium, . , 14-17 April, 201, Obernkirchen, Germany: Universität Göttingenmateus__milan_2012_new_dinosaur_and_pterosaur_tracksites_from_the_late_jurassic.pdf
Mateus, O., & Milàn J. (2010).  A diverse Upper Jurassic dinosaur ichnofauna from central-west Portugal. Lethaia. 43, 245-257., Number 2 Abstract
Mateus, O., Callapez P. M., & Puértolas-Pascual E. (2017).  The oldest Crocodylia? a new eusuchian from the Late Cretaceous (Cenomanian) of Portugal. Journal of Vertebrate Paleontology, Program and Abstracts. 2017, 160. Abstractmateus_et_al_2017_oldest_crocodylia_svp_2017_abstract.pdf


Mateus, O., & Antunes M. T. (2008).  Landmarks in the history of dinosaur paleontology in Portugal, focusing on skeletal remains. Abstract volume, Dinosaurs - A Historical Perspective, 6-7 may 2008. , London Abstract
Mateus, O. (2007).  Notes and review of the ornithischian dinosaurs of Portugal. Journal of Vertebrate Paleontology. 27, 114A-114A., Jan: Society of Vertebrate Paleontology Abstractmateus_2007_notes_and_review_of_ornithichians_of_portugall.pdf

The record of ornithischian dinosaurs from Portugal is substantial but incomplete in terms of our understanding of taxonomic composition and details of the anatomy of many forms. New data and reinterpretation of these forms are provided. The basal thyreophoran from the Lower Jurassic (the nomen dubium “Lusitanosaurus liasicus”) is the most primitive dinosaur from Iberia. Concerning the Late Jurassic, new material from the Lourinhã Formation (Kimmeridgian/Tithonian) referable to the stegosaur Dacentrurus provides additional information on the systematic position and osteology.
The new material shows two rows of paired triangular plates, with a notch in the anterior rim. A maxillary of an ankylosaur from the Vale Frades (Lourinhã Fm.) is here reported, but cannot be referable to Dracopelta, the only ankylosaur genus currently known from Portugal. The ornithopod Alocodon kuehnei reported as Middle Jurassic (Callovian) is probably Oxfordian in age. A right dentary (ML768 from Zimbral) from the Lourinhã Formation, Kimmeridgian/Tithonian, and shares affinities with Dryosaurus but possesses more denticulation and no secondary ridges, suggesting the occurrence of a new or unreported species for the Late Jurassic of Portugal, which is here tentatively ascribed to aff. Dryosaurus sp. In summary, the Late Jurassic ornithischians species/genera from Portugal include Dacentrurus armatus, Stegosaurus cf. ungulatus, Dracopelta zbyszewskii, Phyllodon henkeli, Hypsilophodon sp., Alocodon kuehnei, Trimucrodon cuneatus, aff. Dryosaurus and Draconyx loureiroi. The Lower Cretaceous (Hauterivian / Barremian) of Praia das Aguncheiras, in Cabo Espichel, provided a partial left maxillary (CPGP.1.99.7) of a basal iguanodontian that shows affinities with Camptosaurus, and is tentatively assigned to this genus. The maxillary teeth denticles differ from Iguanodon or other Iguanodontoidea because not show mammillations. More material is necessary to validate but, to be true, that would confirm the presence of this genus in the lower Cretaceous. The Iguanodon has been well reported in the Lower Cretaceous of Cabo Espichel.

Mateus, O., Antunes M. T., & Taquet P. (2001).  Dinosaur ontogeny : the case of Lourinhanosaurus (Late Jurassic, Portugal). J. Vertebr. Paleontol. 21, Abstract
Mateus, O., Taquet P., Antunes M. T., Mateus H., & Ribeiro V. (1998).  Theropod dinosaur nest from Lourinhã, Portugal. Journal of Vertebrate Paleontology. 18, 61A., Number (Suppl. 3) Abstractmateus_et_al_1998_theropod_dinosaur_nest_from_lourinha_portugal_svp.pdfWebsite

More than 100 well preserved dinosaur eggs have been discovered in the Upper Jurassic levels (Tithonian) of Lourinhã, Portugal. The eggshels dispersed in a big area with 11 meters in the highest diameter with high concentration in the middle.

Mateus, O. (1998).  Lourinhanosaurus antunesi, a new Upper Jurassic allosauroid (Dinosauria : Theropoda) from Lourinha, Portugal. Memórias da Academia de Ciências de Lisboa. 37, 111-124. Abstract
Mateus, O., & Antunes M. T. (2008).  Landmarks in the history of dinosaur paleontology in Portugal, focusing on skeletal remains. Abstract volume, Dinosaurs - A Historical Perspective, 6-7 may 2008. , London Abstract

Portugal has been providing dinosaur remains since, at least, 1863. The 18th century tiles depicting the legend of Our Lady in Cabo Espichel are probably the oldest known dinosaur track illustration. To our knowledge, the first remains found in Portugal were theropod teeth collected near Porto das Barcas (Late Jurassic of Lourinhã) in June 20th, 1863 by the geologist Carlos Ribeiro (1813-1882). The first dinosaur paper was written by Henri Sauvage (1842-1917) published in 1896. All remains collected since 19th century were gathered in a work signed by Albert de Lapparent (1905-1975) and Georges Zbyszewski (1909-1999 ) titled Les Dinosauriens du Portugal (1957) that was a significant milestone in the Portuguese dinosaur paleontology and gives the state-of-the-art by the time. Several dinosaurs are named, described, depicted and mapped in that monograph. The first track record is given by Jacinto Pedro Gomes (1844-1916) in 1916. Concerning the non-scientific literature referring to dinosaurs, in 1884 the newspaper Occidente reports the Bernissart findings in Belgium. In the 1959 occurs the first visit to Portugal of Walter Kühne (1911-1991) from the Free University of Berlin. Further visits and work granted the access to the Guimarota Mine and other Late Jurassic deposits in the 1960’s, 70’s and 80’s with a high number of publications. In the 1980’s and early 1990’s starts a progressive era for dinosaur paleontology in Portugal with the works of Peter Galton, Miguel Telles Antunes, the Natural History Museum, the Museum of Lourinhã and the New University of Lisbon, Oliver Rauhut, and others.

Mateus, O. (2014).  Elephas and other vertebrate fossils near Taghrout, Morocco. Journal of Vertebrate Paleontology. Program and Abstracts, 2014, 178., 1 Abstract
Mateus, O., Neto de Carvalho C., & Klompmaker A. A. (2013).  Decapod crustacean body and ichnofossils from the Mesozoic of Portugal. 5th Symposium on Mesozoic and Cenozoic Decapod Crustaceans. , 25–27 June 2013, Warszawa: Polish Geological Institute − National Research Institute & AGH University of Science and Technologymateus_et_al_2013_crustacea_mesozoic_portugal_5th_decapod_crustaceans_meeting_2013.pdf
Mateus, O., Dinis J., & Cunha P. (2014).  Upper Jurassic to Lowermost Cretaceous of the Lusitanian Basin, Portugal-landscapes where dinosaurs walked. Ciências da Terra, special. , Number 8 Abstract
Mateus, O. (2010).  Paleontological collections of the Museum of Lourinhã (Portugal). (JM Brandão, Callapez, PM, O. Mateus, Castro, P, Ed.).Colecções e museus de Geologia: missão e gestão. 121–126., 1: Ed. Universidade de Coimbra e Centro de Estudos de História e Filosofia da Ciência Abstract
Mateus, O. (2009).  The sauropod Turiasaurus riodevensis in the Late Jurassic of Portugal. Journal of Vertebrate Paleontology. 29, 144A., 1, Number 3: Taylor & Francis Abstract

THE SAUROPOD DINOSAUR TURIASAURUS RIODEVENSIS IN THE LATE JURASSIC OF PORTUGAL MATEUS, Octávio, New University of Lisbon (CICEGe-FCT) & Museum of Lourinhã, Lisboa, Portugal A partial sauropod was found in 1996 in Vale Pombas, north of Lourinhã, Central West of Portugal, in the Lourinhã Formation, top of Amoreira Porto Novo member dated as c. 150 M.a. (Early Tithonian, Late Jurassic) and is currently housed at Museum of Lourinhã, in Portugal. The specimen (ML368) comprises a complete tooth with root, anterior chevron and almost complete right forelimb including partial scapula, complete coracoid, humerus, ulna, radius, metacarpals I, III and V, phalanx, and ungual phalanx I. It can be ascribed to Turiasaurus riodevensis, which was previously described from the Villar del Arzobispo Formation at Riodeva (Teruel, Spain). Characters shared with T. riodevensis holotype include: curvature and asymmetry of tooth crown, expansion of crown, outline of humerus, medial deflection of the proximal end of humerus, shape and prominence of deltopectoral crest, vertical ridge in the distal half of the ulna (considered as diagnostic of Turiasauria), configuration of metacarpals, and bone proportions. It differs from T. riodevensis holotype by the smaller size and the more rectangular ungual phalanx in lateral view. The sediments from which the Riodeva specimen was recovered were previsouly thought to be Tithonian to Berriasian in age. The presence of this species in Portugal, in beds confidently dated as Early Tithonian, may allow a more precise date for the Riodeva type locality of early Tithonian in age. The humerus of the Portuguese T. riodevensis is 152 cm long. Although shorter than the Spanish specimen (790 mm), it represents a large individual. All adult sauropods recovered in Portugal thus far are very large individuals: Dinheirosaurus (estimated body length is 20- 25 m), Lusotitan (humerus length estimated to be 205 cm), Lourinhasaurus (femur length: 174 cm), and Turiasaurus here reported. The lack of of small or medium adult body-size sauropods in the Late Jurassic of Portugal, suggests browsing niches thought to be occupied by smaller forms, could be have been available for other dinosaurs, like the long necked stegosaur Miragaia longicollum.

Mateus, O. (2010).  Colecções e museus de Geologia: missão e gestão. , 1: Ed. Universidade de Coimbra e Centro de Estudos de História e Filosofia da Ciência Abstract
Mateus, O., & Jacinto J. J. (2008).  Hemidactylus turcicus. (A Loureiro, N F de Almeida, M.A Carretero, O S Paulo, Ed.).Atlas dos Anfíbio e Répteis de Portugal. 134-135. Abstract
Mateus, O., Callapez P. M., Polcyn M. J., Schulp A. S., Gonçalves A. O., & Jacobs L. L. (2019).  O registo fóssil da biodiversidade em Angola ao longo do tempo: uma perspectiva paleontológica. (Huntley B.J., Russo V., Lages F., Ferrand N., Ed.).Biodiversidade de Angola: Ciência e Conservação - Uma Síntese Moderna. 89-116., Porto: Arte & Ciência Abstractmateus_et_al_2019_paleobiodiversidade_angola.pdf

Este capítulo apresenta uma visão geral da paleobiodiversidade alfa de Angola com base no registo fóssil disponível, o qual se limita às rochas sedimentares, a sua idade variando entre o Pré‑Câmbrico e o pre‑
sente. O período geológico com a maior paleobiodiversidade no registo fóssil angolano é o Cretácico, com mais de 80% do total dos táxones fósseis conhecidos, especialmente moluscos marinhos, sendo estes na sua maioria
amonites. Os vertebrados representam cerca de 15% da fauna conhecida e cerca de um décimo destes são espécies descritas pela primeira vez com base em espécimes de Angola.

Mateus, O., Morais M. L., Schulp A. S., Jacobs L. L., & Polcyn M. J. (2006).  The Cretaceous of Angola. Journal of Vertebrate Paleontology. 26, 96–97., Number (Suppl. T Abstract
Mateus, O., & Antunes M. T. (2000).  Torvosaurus sp. (Dinosauria : Theropoda) in the Late Jurassic of Portugal. Livro de Resumos do I Congresso Ibérico de Paleontologia, pp: 115-117. 115-117. Abstractmateus_antunes_2000_torvosaurus_sp.dinosauria_-_theropoda_in_the_late_jurassic_of_portugal.pdf


Mateus, O., Callapez P. M., & Puértolas-Pascual E. (2017).  The oldest Crocodylia? a new eusuchian from the Late Cretaceous (Cenomanian) of Portugal. Journal of Vertebrate Paleontology, Program and Abstracts. 2017, 160. Abstract
Mateus, O., & Antunes M. T. (2002).  Novos vestígios de dinossauros saurópodes do Jurássico Superior da Lourinhã (Portugal). Congresso Ibérico de Herpetologia. , Évora Abstract
Mateus, O. (1999).  Upper Jurassic dinosaurs of Lourinhã (Portugal). Journal of Vertebrate Paleontology. 19(Suppl. to 3), 62A., Number (Suppl. to 3)mateus_1999_upper_jurassic_lourinha_portugal_svp_1999_abstracts.pdfWebsite
Mateus, O. (1998).  Dinossauros Portugueses. Caderno de resumos do I Congresso de Estudantes de Biologia. 13., Évora Abstract
Mateus, O., Maidment S. C. R., & Christiansen N. A. (2008).  A new specimen aff. Dacentrurus armatus (Dinosauria: Stegosauridae) from the Late Jurassic of Portugal. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontologia de Vertebrados. 157., Neuquén, Argentina Abstractmateus_et_al_2008_a_new_specimen_aff._dacentrurus_armatus_dinosauria_stegosauridae_from_the_late_jurassic_of_portugal.pdf


Mateus, O. (2014).  Cracking dinosaur endothermy: paleophysiology unscrambled. NA, , 1 Abstract

The amniote eggshell functions as a respiratory structure adapted for the optimal transmission of respiratory gasses to and from the embryo according to its physiological requirements. Therefore amniotes with higher oxygen requirements, such as those that sustain higher metabolic rates, can be expected to have eggshells that can maintain a greater gas flux to and from the egg. Studies of extant amniotes have found that eggshells of reduced porosity impose a limit on the metabolic rate of the offspring. Here we show a highly significant relationship between metabolic rates and eggshell porosity in extant amniotes that predicts highly endothermic metabolic rates in dinosaurs. This study finds the eggshell porosity of extant endotherms to be significantly higher than that of extant ectotherms. Eggshell porosity values of dinosaurs are found to be significantly higherthan that of extant ectotherms, but not extant endotherms. Dinosaur eggshells are commonly preserved in the fossil record, and porosity may be readily identified and measured. This provides a simple tool to identify metabolic rates in extinct egg-laying tetrapods whose eggs possessed a mineralized shell

Mateus, O., & Jacinto J. (2008).  Hemidactylus turcicus. Atlas dos Anfíbios e Répteis de Portugal. 130-131., Lisboa: A. Loureiro, N. Ferrand de Almeida, M.A. Carretero, O.S. Paulo. Instituto da Conservação da Natureza e da Biodiversidademateus__jacinto_2008_hemidactylus_turcicus_in_atlas_dos_anfibios_e_repteis_de_portugal.pdf
Mateus, O., & Marzola M. (2014).  Dinosaur taphonomy in the Lourinhã Formation (Late Jurassic, Portugal). 7th International Meeting on Taphonomy and Fossilization, Taphos 2014. 60-61., Ferrara, Italymateus__marzola_2014_lourinha_taphonomy_ferrara_taphonomy_meeting_2014.pdf
Mateus, O. (2012).  Ontogenetical changes in the quadrate of basal tetanurans. Fundamental!. 101–104., 1 Abstract

Although nonavian theropod have received considerable interest in the last years, their ontogeny still remains poorly understood, especially the ontogenetical changes affecting their skull (Rauhutand Fechner, 2005). The quadrate, for instance, is preserved in several embryos and juvenile specimens belonging to many clades of theropods such as the Tyrannosauridae (Carr, 1999), Compsognathidae (Dal Sasso and Maganuco, 2011), Therizinosauroidea (Kúndrat et al., 2007), Oviraptoridae (Norell et al., 1994; Norell et al., 2001; Weishampel et al., 2008) and Troodontidae (Varrichio et al., 2002) but very little is usually said about the anatomy of this bone and no one has ever investigated ontogenetical variation in the nonavian theropod quadrate. The discovery of two quadrates belonging to embryos of the sinraptorid Lourinhanosaurus antunesi from Portugal and five isolated quadrates pertaining to juvenile, subadult and adult specimens of Spinosauridae from Morocco fills this gap and allows some ontogenetic information to be drawn for this bone in these two specific clades of Theropoda.

Mateus, O., Marzola M., Schulp A. S., Jacobs L. L., Polcyn M. J., Pervov V., Gonçalves A. O., & Morais M. L. (2017).  Angolan ichnosite in a diamond mine shows the presence of a large terrestrial mammaliamorph, a crocodylomorph, and sauropod dinosaurs in the Early Cretaceous of Africa. Palaeogeography, Palaeoclimatology, Palaeoecology. 471, 220 - 232. Abstractmateus_et_al_2017_angolan_ichnosite_catoca.pdfWebsite

Abstract We report here new and the first mammaliamorph tracks from the Early Cretaceous of Africa. The tracksite, that also bears crocodylomorph and sauropod dinosaurian tracks, is in the Catoca diamond mine, Lunda Sul Province, Angola. The mammaliamorph tracks have a unique morphology, attributed to Catocapes angolanus ichnogen. et ichnosp. nov. and present an anterolateral projection of digit I and V. The tracks with an average length of 2.7 cm and width of 3.2 cm are the largest mammaliamorph tracks known from the Early Cretaceous unmatched in size in the skeletal fossil record. The crocodylomorph trackways and tracks are attributed to Angolaichnus adamanticus ichnogen. et ichnosp. nov. (‘ichnofamily’ Batrachopodidae) and present a functionally pentadactyl pes, an extremely outwardly rotated handprint, and an unusual tetradactyl and plantigrade manus. One medium-sized sauropod dinosaur trackway preserved skin impressions of a trackmaker with stride length of 1.6 m; a second is that of a small-sized sauropod trackmaker with a pace length of 75 cm.

Mateus, O., Mateus O., Jacobs L. L., Schulp A. S., Polcyn M. J., Tavares T. S., Neto A. B., Morais M. L., & Antunes M. T. (2011).  Angolatitan adamastor, a new sauropod dinosaur and the first record from Angola. Anais da Academia Brasileira de Ciências. Abstract
Mateus, O., Overbeeke M., & Rita F. (2008).  Dinosaur Frauds, Hoaxes and "Frankensteins": How to distinguish fake and genuine vertebrate fossils. Journal of Paleontological Techniques. 2, 1-5. Abstract
Mateus, O., & Milan J. (2010).  First records of crocodyle and pterosaur tracks in the Upper Jurassic of Portugal.. New Mexico Museum of Natural History and Science Bulletin. 51, 83-87., Jan Abstractmateus_and_milan_2010_portugal_first_records_of_crocodyle_and_pterosaur_tracks_in_the_upper_jurassic_of_portugal.pdfWebsite

The Upper Jurassic of Portugal has a rich vertebrate fauna well documented from both body and trace fossils. Although the occurrence of crocodyles and pterosaurs is well documented from body fossils, trace fossils from both groups were unknown until now. Here we describe an isolated crocodyle-like track from Praia da Peralta and pterosaur tracks from the Kimmeridgian of Pedreira do Avelino, Sesimbra (Azóia Fm.) and Porto das Barcas, Lourinhã (Lourinhã Fm.). An enigmatic track suggests the possible presence of a small, tail-dragging tetrapod.
Possible track-makers are suggested based on the known Late Jurassic vertebrate fauna of Portugal.

Mateus, O., Pereira B., Rocha R., & Kullberg J. C. (2018).  Aspiring Geopark Oeste in Portugal: scientific highlights and importance. 8th International Conference on UNESCO Global Geoparks. , 8-14 Sept., Adamello Brenta Geopark, Trentinomateus_et_al_2018_geopark_oeste.pdf
Mateus, O., & Antunes M. T. (2000).  Late Jurassic dinosaurs of Portugal.. Abstracts of the 1st Symposium of European Dinosaurs, p.18. , Dusseldorf, Germany. Abstractmateus__antunes_2000_late_jurassic_dinosaurs_of_portugal_dusseldorf_2000.pdf


Mateus, O. (1999).  Upper Jurassic dinosaurs of Lourinhã (Portugal). Journal of Vertebrate Paleontology. 19, 62., Number (Suppl. to 3) Abstract
Mateus, O., & Milan J. (2008).  Sauropod forelimb flexibility deduced from deep manus tracks. 52th Paleontological Association Annual Meeting. 18th-21st December 2008,. 67-68.: University of Glasgow Abstractmateus__milan_2008_palass_sauropod_forelimb_flexibility_deduced_from_deep_manus_tracks.pdf

Sauropods are often considered to have very limited mobility and reduced limb flexibility, mainly due to their giant size and consequent weight. In the Upper Jurassic Lourinhã Formation, central-west Portugal, deep vertical natural casts of sauropod manus tracks are often preserved as the infills of the original tracks. These manus tracks are vertical-walled, with marks of the striations of the skin scales, showing that the movement of the sauropod manus impacting and exiting the mud was totally vertical with no horizontal component of the stride. Some tracks are up to 66 cm deep, which is equivalent to the height of whole sauropod manus. This means that sauropods could lift their anterior feet in a complete vertical manner. Such movement is only possible if there is mobility at elbow and shoulder articulations in a higher degree than previously thought for sauropods. Our vision of sauropod limbs as inflexible columns has to be updated to a more dynamic model for limbs and body.

Mateus, O., Carrano M. T., & Taquet P. (2012).  Osteology of the embryonic theropods from the Late Jurassic of Paimogo, Portugal. Journal of Vertebrate Paleontology, Program and Abstracts, 2012, p.137. ISSN 1937-2809. 137.mateus_et_al_2012_embryos_paimogo_portugal_svp_2012_abstract_book_meeting_abstracts.pdf.pdf