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Jacobs, L. L., Polcyn M. J., Mateus O. \á\}vio, Schulp A. S., Gon\{\c c\}alves A. \ó\}nio O., & Morais M. L. (2016).  Post-Gondwana Africa and the vertebrate history of the Angolan Atlantic Coast. Memoirs of Museum Victoria. 74, 343\–\}362. Abstract
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Salminen, J., Dinis J., & Mateus O. (2013).  Preliminary magnetostratigraphy for Jurassic/Cretaceous transition in Porto da Calada, Portugal. In: Veikkolainen, T., Suhonen, K., Näränen, J., Kauristie, K., and Kaasalainen, S. (eds.). XXVI Geofysiikan päivät,. 119-122., May 21-22 2013 in Helsinkisalminen.johanna_gfp2013_portugal_preliminary_magnetostratigraphy_for_jurassic_cretaceous_transition_in.pdf
Salminen, J., Dinis J., & Mateus O. (2014).  Preliminary Magnetostratigraphy for the Jurassic–Cretaceous Transition in Porto da Calada, Portugal. (Rogério Rocha, João Pais, José Carlos Kullberg, Stanley Finney, Ed.).STRATI 2013 First International Congress on Stratigraphy At the Cutting Edge of Stratigraphy. 873-877., Heidelberg New York Dordrecht London: Springer Abstractsalminen_et_al_2014_porto_da_calada_stratigraphy_jk_boundary_in_rocha_et_al_strati_.pdf

We present a stratigraphic log supporting a preliminary magnetostratigraphy of a Tithonian–Berriasian section in Porto da Calada (Portugal). Based on biostratigraphy and reversed and normal magnetostratigraphy, the location of the Tithonian–Berriasian boundary is tentatively located at ca. 52 m, not in disagreement
with former proposals. Due to the occurrence of later remagnetization (diagenesis), the magnetostratigraphic definition of the Tithonian–Berriasian section at the Cabo Espichel (Portugal) location was not able to be established.

Salminen, J., Dinis J., & Mateus O. (2014).  Preliminary Magnetostratigraphy for the Jurassic–Cretaceous Transition in Porto da Calada, Portugal. {STRATI} 2013. 873–877.: Springer Science $\mathplus$ Business Media Abstract
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Milàn, J., Clemmensen L. B., Adolfssen J. S., Estrup E. J., Frobøse N., Klein N., Mateus O., & Wings O. (2012).  A preliminary report on coprolites from the Late Triassic part of the Kap Stewart Formation, Jameson Land, East Greenland. New Mexico Museum of Natural History and Science, Bulletin. 57, 203-205. Abstractmilan_et_al_2012_greenland_coprolites_triassic.pdf

The basal part of the Triassic-Jurassic (Rhaetian-Sinemurian) Kap Stewart Formation, exposed at Jameson Land, East Greenland, yields an extensive coprolite collection from black, parallel-laminated mudstone (“paper shale”), representing an open lacustrine system. Preliminary investigations show three different types of coprolites: elongated cylindrical masses, composed of irregularly wrapped layers; elongated cylindrical masses with constriction marks; and spirally-coiled specimens.

Mateus, O. (2012).  A preliminary report on coprolites from the Late Triassic part of the Kap Stewart Formation, Jameson Land, East Greenland. Bulletin of the New Mexico Museum of Natural History and Science. 57, 203–205., 1, Number NA Abstract

The basal part of the Triassic-Jurassic (Rhaetian-Sinemurian) Kap Stewart Formation, exposed at Jameson Land, East Greenland, yields an extensive coprolite collection from black, parallel-laminated mudstone (“paper shale”), representing an open lacustrine system. Preliminary investigations show three different types of coprolites: elongated cylindrical masses, composed of irregularly wrapped layers; elongated cylindrical masses with constriction marks; and spirally-coiled specimens.

Campos, H., Mateus O., & Moreno-Azanza M. (2017).  Preliminary results on the stratigraphy and taphonomy of multiple bonebeds in the Triassic of Algarve. Abstract book of the XV Encuentro de Jóvenes Investigadores en Paleontología/XV Encontro de Jovens Investigadores em Paleontologia, Lisboa, 428 pp.. 83-87. Abstractcampos_et_al._2017_preliminary_results_pn_the_stratigraphy_and_taphonomy_of_multiple_beds_in_the_triassic_of_algarve..pdf

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Araújo, R., Mateus O., Walen A., & Christiansen N. (2009).  Preparation techniques applied to a stegosaurian Dinosaur from Portugal. Journal of Paleontological Techniques. 5, 1-24., Jan Abstractarajomateusetal2009.preparationtechn.pdfWebsite

General vertebrate paleontological techniques that have been used in the Museum of Lourinhã (Portugal) are presented here, in particular those applied to a stegosaurian dinosaur skeleton, Miragaia longicollum. A monolith jacket technique using polyurethane foam and plaster is presented. Mechanical preparation
techniques combining the use of an electric grinder and airscribes proved effective during the initial phases of preparation on well-preserved bone embedded in hard matrix. We also present a technique to mould monoliths in the early stages of preparation, creating a thin silicone rubber mould in several contiguous
parts. To mould and cast monoliths before removing individual bones has proven valuable for the preservation of taphonomic data and for display purposes. Polyurethane resin combined with plaster is useful for small casts, while polyester resin applied in four layers is the preferred technique for larger casts.
The four layers are composed of: a first thin layer of polyester resin with bone colour; followed by another layer of polyester resin of sediment colour and containing glass microspheres to make it thicker. The third layer is composed of fibre glass chopped strands, and the fourth is composed of fibre glass mats embedded
in plain polyester resin. 3D scanning and digitization techniques where tested for the storage of osteological information of individual bones and proved very promising.

Mateus, O. (2009).  Preparation techniques applied to a stegosaurian Dinosaur from Portugal. Journal of Paleontological Techniques. 5, 1–24., 1, Number NA Abstract
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Araujo, R., Mateus O., Walen A., & Christiansen N. (2009).  Preparation techniques applied to a stegosaurian Dinosaur from Portugal: excavation, laboratory mechanical and chemical preparation, moulding and casting, 3D scanning. Journal of Paleontological Techniques 5: 22.. 22. Abstract
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Jacobs, L.  L., Sousa N., Goncalves A.  O., Mateus O., Polcyn M.  J., & Schulp A.  S. (2020).  Projecto PaleoAngola: Geoheritage and Conservation Paleobiology as Science for Development in Angola. AGU Fall Meeting Abstracts. 2020, SY048-05. Abstractprojecto_paleoangola__geoheritage.pdf

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Hendrickx, C., Mateus O., & Araújo R. (2015).  A proposed terminology of theropod teeth (Dinosauria, Saurischia). Journal of Vertebrate Paleontology. e982797. Abstracthendrickx_et_al_2015_theropod_teeth_svp.pdfWebsite

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Hendrickx, C., Mateus O., & Araujo R. (2015).  A proposed terminology of theropod teeth (Dinosauria, Saurischia). Journal of Vertebrate Paleontology. 35, , Number 5 Abstract
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Mateus, O. (2017).  Que Dinossauros Existiram em Portugal?. : Poster 80x59 cm, as a supplement of newspaper “Correio da Manhã” of 16 September 2017poster_correio_da_manha.jpg
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Rotatori, F. M., Moreno-Azanza M., & Mateus O. (2021).  Re-evaluation of the Holotype of Draconyx loureiroi (Dinosauria, Ornithopoda) with report of new holotype material. PALEODAYS 2021 – XXI Convegno della Società Paleontologica Italiana. 73. Abstractrotatoriet_l_2021_paleodays.pdf

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Rotatori, F. M., Moreno-Azanza M., & Mateus O. (2022).  Reappraisal and new material of the holotype of Draconyx loureiroi (Ornithischia: Iguanodontia) provide insights on the tempo and modo of evolution of thumb-spiked dinosaurs. Zoological Journal of the Linnean Society. , 02 Abstractrotatori_etal2022.pdfWebsite

{The Upper Jurassic Lourinhã Formation is well known for its rich assemblage of fossil vertebrates. In this formation, ornithopod dinosaurs are represented by two iguanodontian species, Eousdryosaurus nanohallucis and Draconyx loureiroi. We recently became aware of unreported material belonging to the holotype of Draconyx loureiroi, consisting of partially articulated manual elements. We here re-describe the holotype specimen ML 357, including the newly discovered material. The specimen was subjected to CT-scanning and its surface data used to assess anatomical characters. Linear measurements of metatarsal III were used to estimate the body length of the specimen. The Draconyx loureiroi holotype was included in two datasets and analysed with maximum parsimony and Bayesian inference approaches to estimate evolutionary rates among Iguanodontia. We present evidence that Draconyx loureiroi is a valid taxon nested in Styracosterna and is clearly diagnosable by a unique combination of characters. Both maximum parsimony and Bayesian inference indicate high evolutionary rates across the Jurassic/Cretaceous transition for the base of Iguanodontia. Length estimation suggests that Draconyx loureiroi was a relatively small, bipedal and possibly cursorial animal. Given its basal phyletic position, we interpret this bauplan was the ancestral condition for Styracosterna, that only later in the Cretaceous evolved into giant quadrupedal forms.}

Castanhinha, R., Araujo R., & Mateus O. (2008).  Reptile Egg Sites From Lourinhã Formation, Late Jurassic, Portugal. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontologia de Vertebrados. , Neuquén, Argentina Abstractcastanhinha_et_al_2008_reptile_egg_sites_from_lourinha_formation_late_jurassic_portugal.pdf

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Castanhinha, R., Araujo R., & Mateus O. (2008).  Reptile Egg Sites From Lourinhã Formation, Late Jurassic, Portugal. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. , Neuquén, Argentina Abstract
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Castanhinha, R., Araujo R., & Mateus O. (2008).  Reptile Egg Sites From Lourinhã Formation, Late Jurassic, Portugal. Livro de Resumos de Tercer Congreso Latinoamericano de Paleontología de Vertebrados. , Neuquén, Argentina Abstract
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Costa, F., Silva T., Fernandes J., Calvo R., & Mateus O. (2017).  Retracing the history of a stegosaurian dinosaur discovery in Portugal and the importance of record-keeping in Palaentology. Abstract book of the XV Encuentro de Jóvenes Investigadores en Paleontología/XV Encontro de Jovens Investigadores em Paleontologia, Lisboa, 428 pp.. 119-124. Abstractcosta_et_al_2017_retracing_the_history_-_2017.pdf

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Sousa, J., & Mateus O. (2018).  A review of Ichthyosauria from Portugal. XVI Annual Meeting of the European Association of Vertebrate Palaeontologists. 179., Caparica, Portugal June 26th-July 1st, 2018 Abstractsousa_e_mateus_2018_eavp_abstract.pdf

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Marzola, M., Mateus O., Milàn J., & Clemmensen L. B. (2018).  A review of Palaeozoic and Mesozoic tetrapods from Greenland. Bulletin of the Geological Society of Denmark. 66, 21–46. Abstractmarzola_et_al_2018_-_review_of_greenlandic_tetrapods.pdf

This article presents a synthesis of Palaeozoic and Mesozoic fossil tetrapods from Greenland, including an updated review of the holotypes and a new photographic record of the main specimens. All fossil tetrapods found are from East Greenland, with at least 30 different known taxa: five stem tetrapods (Acanthostega gunnari, Ichthyostega eigili, I. stensioi, I. watsoni, and Ymeria denticulata) from the Late Devonian of the Aina Dal and Britta Dal Formations; four temnospondyl amphibians (Aquiloniferus kochi, Selenocara groenlandica, Stoschiosaurus nielseni, and Tupilakosaurus heilmani) from the Early Triassic of the Wordie Creek Group; two temnospondyls (Cyclotosaurus naraserluki and Gerrothorax cf. pulcherrimus), one testudinatan (cf. Proganochelys), two stagonolepids (Aetosaurus ferratus and Paratypothorax andressorum), the eudimorphodontid Arcticodactylus, undetermined archosaurs (phytosaurs and both sauropodomorph and theropod dinosaurs), the cynodont Mitredon cromptoni, and three mammals (Haramiyavia clemmenseni, Kuehneotherium, and cf. ?Brachyzostrodon), from the Late Triassic of the Fleming
Fjord Formation; one plesiosaur from the Early Jurassic of the Kap Stewart Formation; one plesiosaur and one ichthyosaur from the Late Jurassic of the Kap Leslie Formation, plus a previously unreported Late Jurassic plesiosaur from Kronprins Christian Land. Moreover, fossil tetrapod trackways are known from the Late Carboniferous (morphotype Limnopus) of the Mesters Vig Formation and at least four different morphologies (such as the crocodylomorph Brachychirotherium, the auropodomorph Eosauropus and Evazoum, and the theropodian Grallator) associated to archosaurian trackmakers are known from the Late Triassic of the Fleming Fjord Formation. The presence of rich fossiliferous tetrapod sites in East Greenland is linked to the presence of well-exposed continental and shallow marine deposits with most finds in terrestrial deposits from the Late Devonian and the Late Triassic.

Marzola, M., Mateus O., Milàn J., & Clemmensen L. B. (2018).  A review of palaeozoic and mesozoic tetrapods from greenland. Bulletin of the Geological Society of Denmark. 66, 21-46. Abstract
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Young, M. T., Hua S., Steel L., Foffa D., Brusatte S. L., Thüring S., Mateus O., Ignacio-Ruiz Omeñaca J., Lepage Y., Havilk P., & Andrade M. B. (2014).  Revision of the Late Jurassic teleosaurid genus Machimosaurus (Crocodylomorpha, Thalattosuchia). Royal Society Open Science. 1(140222), 1-42.young_et_al_2014_machimosaurus_crocodylomorph_revision.pdf
Young, M. T., Hua S., Steel L., Foffa D., Brusatte S. L., Thüring S., Mateus O., Ruiz-Omeñaca J. I., Havlik P., Lepage Y., & De Andrade M. B. (2014).  Revision of the Late Jurassic teleosaurid genus Machimosaurus (Crocodylomorpha, Thalattosuchia). Royal Society Open Science. 1, , Number 2 Abstract
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Mateus, O., & Jacinto J. J. (1997).  Ritmos de Actividade e habitat de Hemidactylus turcicus (Reptilia: Gekkonidae) em Évora, Portugal. Cuadernos de INICE. 74-75, 207-214. Abstract
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Holwerda, F., Tschopp E., & Mateus O. (2014).  Sauropod body fossils in Europe: overview and current issues. XII EAVP Meeting XII Annual Meeting of the European Association of Vertebrate Palaeontologists – Abstract Book. p.77., Torino 24-28 June 2014holwerda_et_al_2014_sauropods_europe_eavp.pdf
Mateus, O. (2009).  The sauropod dinosaur Turiasaurus riodevensis in the Late Jurassic of Portugal. Journal of Vertebrate Paleontology. 29, 144A., Jan Abstractmateus_2009_sauropod_dinosaur_turiasaurus_portugal_svp09abstractspdf.pdfWebsite

A partial sauropod was found in 1996 in Vale Pombas, north of Lourinhã, Central West of Portugal, in the Lourinhã Formation, top of Amoreira Porto Novo member dated as c. 150 M.a. (Early Tithonian, Late Jurassic) and is currently housed at Museum of Lourinhã, in Portugal. The specimen (ML368) comprises a complete tooth with root, anterior chevron and almost complete right forelimb including partial scapula, complete coracoid, humerus, ulna, radius, metacarpals I, III and V, phalanx, and ungual phalanx I. It can be ascribed to Turiasaurus riodevensis, which was previously described from the Villar del Arzobispo
Formation at Riodeva (Teruel, Spain). Characters shared with T. riodevensis holotype include: curvature and asymmetry of tooth crown, expansion of crown, outline of humerus, medial deflection of the proximal end of humerus, shape and prominence of deltopectoral crest, vertical ridge in the distal half of the ulna (considered as diagnostic of Turiasauria), configuration of metacarpals, and bone proportions. It differs from T. riodevensis holotype by the smaller size and the more rectangular ungual phalanx in lateral view. The sediments from which the Riodeva specimen was recovered were previsouly thought to be Tithonian to Berriasian in age. The presence of this species in Portugal, in beds confidently dated as Early
Tithonian, may allow a more precise date for the Riodeva type locality of early Tithonian in age. The humerus of the Portuguese T. riodevensis is 152 cm long. Although shorter than the Spanish specimen (790 mm), it represents a large individual. All adult sauropods recovered in Portugal thus far are very large individuals: Dinheirosaurus (estimated body length is 20-25 m), Lusotitan (humerus length estimated to be 205 cm), Lourinhasaurus (femur length: 174 cm), and Turiasaurus here reported. The lack of of small or medium adult body-size sauropods in the Late Jurassic of Portugal, suggests browsing niches thought to be occupied by smaller forms, could be have been available for other dinosaurs, like the long necked stegosaur Miragaia longicollum.

Mateus, O., & Milan J. (2008).  Sauropod forelimb flexibility deduced from deep manus tracks. 52th Paleontological Association Annual Meeting. 18th-21st December 2008,. 67-68.: University of Glasgow Abstractmateus__milan_2008_palass_sauropod_forelimb_flexibility_deduced_from_deep_manus_tracks.pdf

Sauropods are often considered to have very limited mobility and reduced limb flexibility, mainly due to their giant size and consequent weight. In the Upper Jurassic Lourinhã Formation, central-west Portugal, deep vertical natural casts of sauropod manus tracks are often preserved as the infills of the original tracks. These manus tracks are vertical-walled, with marks of the striations of the skin scales, showing that the movement of the sauropod manus impacting and exiting the mud was totally vertical with no horizontal component of the stride. Some tracks are up to 66 cm deep, which is equivalent to the height of whole sauropod manus. This means that sauropods could lift their anterior feet in a complete vertical manner. Such movement is only possible if there is mobility at elbow and shoulder articulations in a higher degree than previously thought for sauropods. Our vision of sauropod limbs as inflexible columns has to be updated to a more dynamic model for limbs and body.

Mateus, O., & Milan J. (2008).  Sauropod forelimb flexibility deduced from deep manus tracks. (University of, Glasgow, Ed.).52th Paleontological Association Annual Meeting. 18th-21st December 2008. 67-68. Abstract
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Mateus, O. (2009).  The sauropod Turiasaurus riodevensis in the Late Jurassic of Portugal. Journal of Vertebrate Paleontology. 29, 144A., 1, Number 3: Taylor & Francis Abstract

THE SAUROPOD DINOSAUR TURIASAURUS RIODEVENSIS IN THE LATE JURASSIC OF PORTUGAL MATEUS, Octávio, New University of Lisbon (CICEGe-FCT) & Museum of Lourinhã, Lisboa, Portugal A partial sauropod was found in 1996 in Vale Pombas, north of Lourinhã, Central West of Portugal, in the Lourinhã Formation, top of Amoreira Porto Novo member dated as c. 150 M.a. (Early Tithonian, Late Jurassic) and is currently housed at Museum of Lourinhã, in Portugal. The specimen (ML368) comprises a complete tooth with root, anterior chevron and almost complete right forelimb including partial scapula, complete coracoid, humerus, ulna, radius, metacarpals I, III and V, phalanx, and ungual phalanx I. It can be ascribed to Turiasaurus riodevensis, which was previously described from the Villar del Arzobispo Formation at Riodeva (Teruel, Spain). Characters shared with T. riodevensis holotype include: curvature and asymmetry of tooth crown, expansion of crown, outline of humerus, medial deflection of the proximal end of humerus, shape and prominence of deltopectoral crest, vertical ridge in the distal half of the ulna (considered as diagnostic of Turiasauria), configuration of metacarpals, and bone proportions. It differs from T. riodevensis holotype by the smaller size and the more rectangular ungual phalanx in lateral view. The sediments from which the Riodeva specimen was recovered were previsouly thought to be Tithonian to Berriasian in age. The presence of this species in Portugal, in beds confidently dated as Early Tithonian, may allow a more precise date for the Riodeva type locality of early Tithonian in age. The humerus of the Portuguese T. riodevensis is 152 cm long. Although shorter than the Spanish specimen (790 mm), it represents a large individual. All adult sauropods recovered in Portugal thus far are very large individuals: Dinheirosaurus (estimated body length is 20- 25 m), Lusotitan (humerus length estimated to be 205 cm), Lourinhasaurus (femur length: 174 cm), and Turiasaurus here reported. The lack of of small or medium adult body-size sauropods in the Late Jurassic of Portugal, suggests browsing niches thought to be occupied by smaller forms, could be have been available for other dinosaurs, like the long necked stegosaur Miragaia longicollum.

Mateus, O. (2009).  The sauropod Turiasaurus riodevensis in the the Late Jurassic of Portugal. Journal of Vertebrate Paleontology. 29, 144., Number 3 Abstract
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Mateus, O. (2009).  The sauropod Turiasaurus riodevensis in the the Late Jurassic of Portugal. Journal of Vertebrate Paleontology. 29, 144–144., Number 3 Abstract
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Lallensack, J. N., Klein H., Milàn J., Wings O., Mateus O., & Clemmensen L. B. (2017).  Sauropodomorph dinosaur trackways from the Fleming Fjord Formation of East Greenland: Evidence for Late Triassic sauropods. Acta Palaeontologica Polonica. 62(4), 833-843. Abstractlallensack_et_al_2017_-_sauropodomorph_tracks_greenland.pdf

The Late Triassic (Norian–early Rhaetian) Fleming Fjord Formation of central East Greenland preserves a diverse fossil fauna, including both body and trace fossils. Trackways of large quadrupedal archosaurs, although already reported in 1994 and mentioned in subsequent publications, are here described and figured in detail for the first time, based on photogrammetric data collected during fieldwork in 2012. Two trackways can be referred to Eosauropus, while a third, bipedal trackway may be referred to Evazoum, both of which have been considered to represent sauropodomorph dinosaur tracks. Both the Evazoum and the Eosauropus trackways are distinctly larger than other trackways referred to the respective ichnogenera. The trackmaker of the best preserved Eosauropus trackway is constrained using a synapomorphy-based approach. The quadrupedal posture, the entaxonic pes structure, and five weight-bearing digits indicate a derived sauropodiform trackmaker. Other features exhibited by the tracks, including the semi-digitigrade pes and the laterally deflected unguals, are commonly considered synapomorphies of more exclusive clades within Sauropoda. The present trackway documents an early acquisition of a eusauropod-like pes anatomy while retaining a well-developed claw on pedal digit IV, which is reduced in eusauropods. Although unequivocal evidence for sauropod dinosaurs is no older than the Early Jurassic, the present trackway provides evidence for a possible Triassic origin of the group.

Lallensack, J. N., Klein H., Milàn J., Wings O., Mateus O., & Clemmensen L. B. (2017).  Sauropodomorph dinosaur trackways from the Fleming Fjord Formation of East Greenland: evidence for Late Triassic sauropods. Acta Palaeontologica Polonica. 62, 833–843., Number 4 Abstract
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Lallensack,  J. N., Klein  H., Milàn  J., Wings  O., Mateus  O., & Clemmensen  L. B. (2017).  Sauropodomorph dinosaur trackways from the Fleming Fjord Formation of East Greenland: evidence for Late Triassic sauropods. AbstractWebsite
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Cunha, P. P., Mateus O., & Antunes M. T. (2004).  The sedimentology of the Paimogo dinosaur nest site (Portugal, Upper Jurassic). 23 rd IAS Meeting of Sedimentology. 93., Coimbra, Portugal Abstractcunha_p_p_2004_-_the_sedimentology_of_the_paimogo_dinosaur_nest_site_portugal10467.pdf

Sedimentological features of the Paimogo site, 6 km NNW of Lourinhã, western central Portugal are presented. More than one hundred theropod dinosaur eggs (some containing embryo bones) ascribed to Lourinhanosaurus antunesi Mateus 1998, three crocodilian eggs and some other fossils were found at the 32 m2 excavated area of the egg-bearing horizon (Mateus et al., 1998). The stratigraphic position of the site is the Praia Azul member (Lourinhã Formation), roughly corresponding to the Kimmeridgian-Tithonian boundary or, more likely lowermost Tithonian. The maximum flooding surface of the basinal transgressive event where the horizon is located corresponds to the base of the H depositional sequence defined by Pena dos Reis et al. (2000) and probably correlates to the base of sequence Ti1 identified within western European basins (Jacquin et al., 1998), dated as 150 Ma. Possibly during the normal river discharge, the theropods congregated in nesting colonies at the backswamp of an extensive flood plain with small meandering channels and freshwater ponds. There are no evidences that the nest was dug or the eggs buried. The eggs have probably been laid on a flat, shaded, muddy area near the bank of a large pond. It is probable that the eggs have not been actively incubated. The larger number of eggs suggest that they were laid near simultaneously by, at least, six females. The fossil record shows that crocodilians, mammals, gastropods and fish were also present. A flood event occurred when theropod embryos had attained a late stade of ontogenetic development, probably just before hatching. The overflow from a nearby channel flooded the plain, including the area where the eggs had been laid. The sheet flood flowing over the nest resulted into the scattering and breaking up of some dinosaur eggs. Eggshell and embryos skeletal parts fragments were displaced to an adjacent area where, due to hydrodynamic decline, the flow submerged other clutches and moderately dragged their eggs. The flooding caused the drowning of the embryos and covered the eggs with fine-grained sediment, hiding them from predators and scavengers. Hydrodynamic interpretation of the arrangement of the theropod eggs and egg-fragments suggests that the flow came from the NW. When the floodwaters receded, the fine-grained deposits became exposed to subaerial weathering. Although the sediment surface was often wet and small bodies of standing water may still have existed, the sediments were oxidized and plant remains have consequently been destroyed. Some carbonate cementation and redenning resulted from pedogenesis under alternating dry and moist conditions, in a semiarid/ sub-tropical climate under seasonal changing, contrasting conditions. The thick, stratigraphically above and below the nesting horizon mudrocks indicate a long persistence of periodic flooding, alternating with pedogenesis. During the early stages of diagenesis, vertical pressure crushed the eggs. Silt penetration into the inner part of each egg inhibited later flattening during the burial process.

Cunha, P. P., Mateus O., & Antunes M. T. (2004).  The sedimentology of the Paimogo dinosaur nest site (Portugal, Upper Jurassic). 23 rd IAS Meeting of Sedimentology. 93., Coimbra, Portugal Abstract
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Cunha, P. P., Mateus O., & Antunes M. T. (2004).  The sedimentology of the Paimogo dinosaur nest site (Portugal, Upper Jurassic). 23 rd IAS Meeting of Sedimentology. 93–93., Coimbra, Portugal Abstract
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