Publications

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Book Chapter
Conference Paper
Estraviz López, D., & Mateus O. (2022).  Bearing the weight: Technical recovery of brown bear fossils (Ursus arctos) from Algar do Vale da Pena, Portugal. XX EJIP. , Cañaveral de León Abstractestraviz-lpezmateus2022bearingtheweighttechnicalrecoveryofbrownbearfossilsursusarctosfromalgardovaledapenaportugal.pdf

The Algar do Vale da Pena is a cave that was discovered during limestone mining works around 1980, at the Natural Park of Serras de Aire e Candeeiros, Portugal. The entrance to the cave, through the collapsed roof, is a 30 meter near-vertical well, inaccessible without means of vertical progression and specialized equipment. Soon after the discovery of the cave, bone material belonging to bears and hundreds of bear claw marks were located inside it. Nevertheless, the technical descent and the extraordinary difficulties of retrieving the calcite flowstones that encased the bones prevented scientists from undertaking excavations for nearly 40 years. Since 2016 a series of expeditions by a team of paleontologists from NOVA University and Museum of Lourinhã recovered more than 20 disarticulated remains from five bone accumulations, separated more than five meters apart, which given the level of erosion and breakage, are para-autochthonous in origin. These are interpreted as belonging to different individuals. Morphological characters, like a relatively elongated third phalanx, the clear separation of paracone-metacone of the first upper molar and the ventral position of the mandibular condyle; as well as numerous morphometric characters, allow to ascribe the remains to Ursus arctos. Nevertheless, the main finding from the cave is a complete skull with an articulated mandible; which denotes absence of transport, encased within a 180 kg calcite flowstone, including other skeletal remains as well, which could not be extracted from the cave until 2021. This block was removed with the help of members of several speleological groups. It promises a wealth of information about fossil brown bears, being the most complete bear fossil ever recovered in Portugal. It is currently being dated trough magnetostratigraphy and prepared. We consider this intervention as an extraordinary example of citizen science applied to paleontology

Crespo, V. D., Sequero López C., Rios M., Estraviz López D., Gamonal A., Martino R., Riccetto M., Callapez P., Legoinha P., & Mateus O. (2023).  Revisiting the classical small mammal assemblage from the Eocene site of Silveirinha: a fresh look at old collections. XI Congresso Nacional de Geologia: Geociências e Desafios globaisAt: Universidade de CoimbraVolume: Livro de Resumos, págs 85-86. , Coimbra Abstractcrespov_etal_xicng2023_silveirinha.pdf

O sítio de Silveirinha é uma das localidades de mamíferos mais conhecidas da Paleontologia do Cenozoico de Portugal e da Europa em geral. Graças à sua rica e diversificada associação de mamíferos, com mais de 30 taxa, foi posicionado no Eocénico inferior (início do Ypresiano, MP7, ca. 55,8 M.a.), sendo o local mais antigo da Europa desta Época, devido à presença única de taxa típicos do Paleocénico superior, juntamente com outras espécies já características do Eocénico inferior. Este estudo irá rever o material de pequenos mamíferos deste sítio, conservado na coleção clássica da Universidade Nova de Lisboa, a fim de fazer uma actualização taxonómica à luz das publicações mais recentes.

Conference Proceedings
Ezquerro, L., Balauz B., Coimbra R., Nuñez-Lahuerta C., Mateus O., Román T., & Moreno-Azanza M. (2021).  Evidence of sedimentary remodel of Jurassic theropod egg clutches (Lourinhã, Portugal). 18th Conference of the EAVP-Abstract book. 72-73. Abstractezquerro_et_al_2021_sedimentary_remodel.pdf

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Ríos, M., Estraviz-López D., Martino R., Lohmann P., Mateus O., & Solounias N. (2022).  The first Sivatheriine (Artiodactyla, Giraffidae) from the Middle Miocene of Portugal. XIX Annual conference of the European Association of Vertebrate Palaeontologists (19th EAVP)At: Benevento, Italy. 167. Abstracteavp_2022_abstractvolume-176.pdf

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Moreno-Azanza, M., Mateus O., Bauluz B., Coimbra R., Ezquerro L., & Núñez-Lahuerta C. (2021).  Hatching in Portugal: a new look to old eggs. XIX Encontro de Jovens Investigadores em Paleontologia. 22. Abstractmoreno-azanza_et_al_2021_eggs_ejip.pdf

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Estraviz López, D., & Mateus O. (2018).  The history of the Quaternary vertebrate paleontology in Portugal. XVI Annual Meeting of the European Association of Vertebrate Palaeontologists. 65., Caparica, Portugal June 26th-July 1st, 2018: Abstract book of the XVI Annual Meeting of the European Association of Vertebrate Palaeontologists, Caparica, Portugal June 26th-July 1st, 2018. Abstractestraviz__mateus_2018_eavp_abstract.pdf

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Estraviz Lopez, D., & Mateus O. (2018).  Paleobiodiversity of Quaternary fossil tetrapods in continental Portugal. 1st Palaeontological Virtual Congress. online. Abstractestraviz-lopez-mateus_palaeovc2018_abstract.pdf

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Moreno-Azanza, M., Ezquerro L., Gasca J. M., Díaz-Berenguer E., Nuñez-Lahuerta C., Pérez-Pueyo M., Puértolas-Pascual E., Sellés A. G., Vila B., & Laita E. (2021).  Two latest Cretaceous egg localities in the external sierras (Southern Pyrenees, Huesca Province, NE Spain). Palaeovertebrata - XVIII annual conference of the European Association of Vertebrate Palaeontologists. 44, 120-121. Abstractmoreno-azanza_et_al_2021_eggs_eavp_abstract.pdf

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Jackson, Y. J., Economos R. C., Jacobs L. L., Mateus O., & Gonçalves A. O. (2020).  When dinosaurs walked through diamonds: constraining the age of Early Cretaceous footprints in volcanic crater sediments. 54th Annual GSA South-Central Section Meeting 2020. , Fort Worth: Geological Society of America Abstracts with Programs. Vol. 52, No. 1jackson_et_al_2020_abstract__when_dinosaurs_walked_through_diamonds__gsa.pdf
Journal Article
Klein, H., Milàn J., Clemmensen L. B., Frobøse N., Mateus O., Klein N., Adolfssen J. S., Estrup E. J., & Wings O. (2016).  Archosaur footprints (cf. Brachychirotherium) with unusual morphology from the Upper Triassic Fleming Fjord Formation (Norian–Rhaetian) of East Greenland. Geological Society, London, Special Publications. 434(1), 71-85. Abstractklein_et_al_2015_archosaur_footprints_cf._brachychirotherium_with_unusual.pdfWebsite

The Ørsted Dal Member of the Upper Triassic Fleming Fjord Formation in East Greenland is well known for its rich vertebrate fauna, represented by numerous specimens of both body and ichnofossils. In particular, the footprints of theropod dinosaurs have been described. Recently, an international expedition discovered several slabs with 100 small chirotheriid pes and manus imprints (pes length 4–4.5 cm) in siliciclastic deposits of this unit. They show strong similarities with Brachychirotherium, a characteristic Upper Triassic ichnogenus with a global distribution. A peculiar feature in the Fleming Fjord specimens is the lack of a fifth digit, even in more deeply impressed imprints. Therefore, the specimens are assigned here tentatively to cf. Brachychirotherium. Possibly, this characteristic is related to the extremely small size and early ontogenetic stage of the trackmaker. The record from Greenland is the first evidence of this morphotype from the Fleming Fjord Formation. Candidate trackmakers are crocodylian stem group archosaurs; however, a distinct correlation with known osteological taxa from this unit is not currently possible. While the occurrence of sauropodomorph plateosaurs in the bone record links the Greenland assemblage more closer to that from the Germanic Basin of central Europe, here the described footprints suggest a Pangaea-wide exchange.Supplementary material: Three-dimensional model of cf. Brachychirotherium pes–manus set (from MGUH 31233b) from the Upper Triassic Fleming Fjord Formation (Norian–Rhaetian) of East Greenland as pdf, ply and jpg files (3D model created by Oliver Wings; photographs taken by Jesper Milàn) is available at https://doi.org/10.6084/m9.figshare.c.2133546

Klein, H., Milàn J., Clemmensen L. B., Frobøse N., Mateus O., Klein N., Adolfssen J. S., Estrup E. J., & Wings O. (2015).  Archosaur footprints (cf. Brachychirotherium) with unusual morphology from the Upper Triassic Fleming Fjord Formation (Norian–Rhaetian) of East Greenland. Geological Society, London, Special Publications. 434, AbstractWebsite

The Ørsted Dal Member of the Upper Triassic Fleming Fjord Formation in East Greenland is well known for its rich vertebrate fauna, represented by numerous specimens of both body and ichnofossils. In particular, the footprints of theropod dinosaurs have been described. Recently, an international expedition discovered several slabs with 100 small chirotheriid pes and manus imprints (pes length 4–4.5 cm) in siliciclastic deposits of this unit. They show strong similarities with Brachychirotherium, a characteristic Upper Triassic ichnogenus with a global distribution. A peculiar feature in the Fleming Fjord specimens is the lack of a fifth digit, even in more deeply impressed imprints. Therefore, the specimens are assigned here tentatively to cf. Brachychirotherium. Possibly, this characteristic is related to the extremely small size and early ontogenetic stage of the trackmaker. The record from Greenland is the first evidence of this morphotype from the Fleming Fjord Formation. Candidate trackmakers are crocodylian stem group archosaurs; however, a distinct correlation with known osteological taxa from this unit is not currently possible. While the occurrence of sauropodomorph plateosaurs in the bone record links the Greenland assemblage more closer to that from the Germanic Basin of central Europe, here the described footprints suggest a Pangaea-wide exchange.Supplementary material: Three-dimensional model of cf. Brachychirotherium pes–manus set (from MGUH 31233b) from the Upper Triassic Fleming Fjord Formation (Norian–Rhaetian) of East Greenland as pdf, ply and jpg files (3D model created by Oliver Wings; photographs taken by Jesper Milàn) is available at https://doi.org/10.6084/m9.figshare.c.2133546

Eberth, D. A., Kobayashi Y., Lee Y. N., Mateus O., Therrien F., Zelenitsky D. K., & Norell M. A. (2009).  Assignment of Yamaceratops dorngobiensis and Associated Redbeds at Shine Us Khudag (Eastern Gobi, Dorngobi Province, Mongolia) to the Redescribed Javkhlant Formation (Upper Cretaceous). Journal of Vertebrate Paleontology. 29, 295-302., Jan: Univ Nova Lisboa, Hokkaido Univ, Museu Lourinha, Amer Museum Nat Hist, Korean Inst Geosci & Mineral Resources, Royal Tyrell Museum, Royal Tyrell Museum, Univ Calgary Abstracteberth_et_al-2009-__assignment_of_yamaceratops_dorngobiensis_and_associated_redbeds_at_shine_us_khudag_eastern_gobi_dorngobi_province_mongolia_to_the_redescribed_javkhlant_formation_upper_cretaceous_javkhlant_fm.pdf

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Eberth, D. A., Kobayashi Y., Lee Y. - N., Mateus O., Therrien F., Zelenitsky D. K., & Norell M. A. (2009).  Assignment of Yamaceratops dorngobiensis and associated redbeds at Shine Us Khudag (eastern Gobi, Dorngobi Province, Mongolia) to the redescribed Javkhlant Formation (Upper Cretaceous). Journal of Vertebrate Paleontology. 29, 295–302., mar, Number 1: Informa {UK} Limited AbstractWebsite
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Hansen, B. B., Milàn J., Clemmensen L. B., Adolfssen J. S., Estrup E. J., Klein N., Mateus O., & Wings O. (2016).  Coprolites from the Late Triassic Kap Stewart Formation, Jameson Land, East Greenland: morphology, classification and prey inclusions. Geological Society, London, Special Publications. 434(1), 49-69. Abstracthansen_et_al_2015_coprolites_from_the_late_triassic_kap_stewart_formation_jameson_land_east_greenland.pdfWebsite

A large collection of vertebrate coprolites from black lacustrine shales in the Late Triassic (Rhaetian–Sinemurian) Kap Stewart Formation, East Greenland is examined with regard to internal and external morphology, prey inclusions, and possible relationships to the contemporary vertebrate fauna. A number of the coprolites were mineralogically examined by X-ray diffraction (XRD), showing the primary mineral composition to be apatite, clay minerals, carbonates and, occasionally, quartz in the form of secondary mineral grains. The coprolite assemblage shows multiple sizes and morphotypes of coprolites, and different types of prey inclusions, demonstrating that the coprolite assemblage originates from a variety of different producers.Supplementary material: A description of the size, shape, structure, texture, contents and preservation of the 328 specimens is available at https://doi.org/10.6084/m9.figshare.c.2134335

Hansen, B. B., Milàn J., Clemmensen L. B., Adolfssen J. S., Estrup E. J., Klein N., Mateus O., & Wings O. (2015).  Coprolites from the Late Triassic Kap Stewart Formation, Jameson Land, East Greenland: morphology, classification and prey inclusions. Geological Society, London, Special Publications. 434, AbstractWebsite

A large collection of vertebrate coprolites from black lacustrine shales in the Late Triassic (Rhaetian–Sinemurian) Kap Stewart Formation, East Greenland is examined with regard to internal and external morphology, prey inclusions, and possible relationships to the contemporary vertebrate fauna. A number of the coprolites were mineralogically examined by X-ray diffraction (XRD), showing the primary mineral composition to be apatite, clay minerals, carbonates and, occasionally, quartz in the form of secondary mineral grains. The coprolite assemblage shows multiple sizes and morphotypes of coprolites, and different types of prey inclusions, demonstrating that the coprolite assemblage originates from a variety of different producers.Supplementary material: A description of the size, shape, structure, texture, contents and preservation of the 328 specimens is available at https://doi.org/10.6084/m9.figshare.c.2134335

Coimbra, R., Moreno-Azanza M., Ezquerro L., Nuñez-Lahuerta C., Gasca J. M., Immenhauser A., Mateus O., & Rocha F. (2023).  Evaluating and comparing geochemical sampling protocols in dinosaur eggshells: refining Cretaceous ecosystem research. Cretaceous Research. 105632. Abstractsingle_file_coimbra_et_al._2023_cretresearch.pdfWebsite

The geochemical signatures of dinosaur eggshells represent well-established proxies in paleoenvironmental and paleobiological research. The variable sampling procedures reported in the literature, however, deserve attention. In order to evaluate the impact of different sampling methodologies on carbon and oxygen isotope and elemental concentrations, grinding was contrasted with drilling to extract powder samples from eggshell fragments collected at several locations. Eggshell data were further contrasted with surface materials, encasing matrix and compared with independent proxies using petrographic and elemental techniques. Iron and manganese elemental concentrations revealed an enrichment sequence depending on the sampling strategy for the same eggshell fragment. This pattern can be mistaken for a variable state of preservation. In contrast, carbon and oxygen isotope values exhibited only subtle differences and lacked clear trends. This suggests that isotope data are less susceptible to different methodological approaches. It is shown that drilling offers a wider range of possibilities compared to grinding (e.g., faster and less destructive). Additionally, drilled powder samples can confidently be used for elemental and isotope analysis, excluding contamination, thus providing a more accurate set of proxy data from eggshell archives.

Mateus, O., Clemmensen L., Klein N., Wings O., Frobøse N., Milàn J., Adolfssen J., & Estrup E. (2014).  The Late Triassic of Jameson Land revisited: new vertebrate findings and the first phytosaur from Greenland. Journal of Vertebrate Paleontology. Program and Abstracts, 2014, 182.mateus_et_al2014-_jameson_land_revisited_-_svp_2014.pdf
Foth, C., Evers S., Pabst B., Mateus O., Flisch A., Patthey M., & Rauhut O. W. M. (2015).  New insights into the lifestyle of Allosaurus (Dinosauria: Theropoda) based on another specimen with multiple pathologies. PeerJ PrePrints. 3, e824v1., 2015 Abstractfoth_et_al_2015_peerj-preprints-824.pdfWebsite

Adult large-bodied theropods are often found with numerous pathologies. A large, almost complete, probably adult Allosaurus specimen from the Howe Stephens Quarry, Morrison Formation (Late Kimmeridgian–Early Tithonian), Wyoming, shows multiple pathologies. Pathologic bones include the left dentary, two cervical vertebrae, one cervical and several dorsal ribs, the left scapula, the left humerus, right ischium, and two left pedal phalanges. These pathologies can be classified as follows: the fifth cervical vertebra, the scapula, several ribs and the ischium are traumatic, and a callus on the shaft of the left pedal phalanx II-2 is traumatic-infectious. Traumatically fractured elements exposed to frequent movement (e.g. the scapula and the ribs) show a tendency to develop pseudarthroses instead of callus healing. The pathologies in the lower jaw and a reduced flexor tubercle of the left pedal phalanx II-2 are most likely traumatic or developmental in origin. The pathologies on the fourth cervical are most likely developmental in origin or idiopathic, that on the left humerus is infectious or idiopathic, whereas left pedal phalanx IV-1 is classified as idiopathic. With exception of the ischium, all traumatic / traumatic-infectious pathologic elements show unambiguous evidences of healing, indicating that the respective pathologies did not cause the death of this individual. Alignment of the scapula and rib pathologies from the left side suggests that all may have been caused by a single traumatic event. The ischial fracture may have been fatal. The occurrence of multiple traumatic pathologies again underlines that large-bodied theropods experienced frequent injuries during life, indicating an active predatory lifestyle, and their survival perhaps supports a gregarious behavior for Allosaurus. Signs of infections are scarce and locally restricted, indicating a successful prevention of the spread of pathogens, as it is the case in extant reptiles (including birds).

Foth, C., Evers S. W., Pabst B., Mateus O., Flisch A., Patthey M., & Rauhut O. W. M. (2015).  New insights into the lifestyle of \\textitAllosaurus (Dinosauria: Theropoda) based on another specimen with multiple pathologies. PeerJ. 3, e940., 5 AbstractWebsite

Adult large-bodied theropods are often found with numerous pathologies. A large, almost complete, probably adult \\textitAllosaurus specimen from the Howe Stephens Quarry, Morrison Formation (Late Kimmeridgian–Early Tithonian), Wyoming, exhibits multiple pathologies. Pathologic bones include the left dentary, two cervical vertebrae, one cervical and several dorsal ribs, the left scapula, the left humerus, the right ischium, and two left pedal phalanges. These pathologies can be classified as follows: the fifth cervical vertebra, the scapula, several ribs and the ischium are probably traumatic, and a callus on the shaft of the left pedal phalanx II-2 is probably traumatic-infectious. Traumatically fractured elements exposed to frequent movement (e.g., the scapula and the ribs) show a tendency to develop pseudarthroses instead of a callus. The pathologies in the lower jaw and a reduced extensor tubercle of the left pedal phalanx II-2 are most likely traumatic or developmental in origin. The pathologies on the fourth cervical are most likely developmental in origin or idiopathic, that on the left humerus could be traumatic, developmental, infectious or idiopathic, whereas the left pedal phalanx IV-1 is classified as idiopathic. With exception of the ischium, all as traumatic/traumatic-infectious classified pathologic elements show unambiguous evidences of healing, indicating that the respective pathologies did not cause the death of this individual. Alignment of the scapula and rib pathologies from the left side suggests that all may have been caused by a single traumatic event. The ischial fracture may have been fatal. The occurrence of multiple lesions interpreted as traumatic pathologies again underlines that large-bodied theropods experienced frequent injuries during life, indicating an active predatory lifestyle, and their survival perhaps supports a gregarious behavior for \\textitAllosaurus. Alternatively, the frequent survival of traumatic events could be also related to the presence of non-endothermic metabolic rates that allow survival based on sporadic food consumption or scavenging behavior. Signs of pathologies consistent with infections are scarce and locally restricted, indicating a successful prevention of the spread of pathogens, as it is the case in extant reptiles (including birds).

Mateus, O., & Estraviz-López D. (2022).  A new theropod dinosaur from the early cretaceous (Barremian) of Cabo Espichel, Portugal: Implications for spinosaurid evolution. PLOS ONE. 17, 1-48., 02, Number 2: Public Library of Science Abstractjournal.pone_.0262614.pdfWebsite

Spinosaurids are some of the most enigmatic Mesozoic theropod dinosaurs due to their unique adaptations to aquatic environments and their relative scarcity. Their taxonomy has proven to be especially problematic. Recent discoveries from Western Europe in general, specifically Iberia, provide some of the best specimens for the understanding of their phylogeny, leading to the description of the spinosaurid Vallibonavenatrix cani and the recognition of the Iberian dinosaur Camarillasaurus cirugedae as one of them. Portuguese associated spinosaurid remains (ML1190) from the Papo Seco Formation (early Barremian) were previously assigned to Baryonyx walkeri but new material recovered in 2020 along with new phylogenetic analyses suggests a different phylogenetic placement, making their revision necessary. Here we show that these remains are not attributable to Baryonyx walkeri, but to a new genus and species, Iberospinus natarioi, gen. et sp. nov. The new taxon is characterized by the presence of a single Meckelian foramen in the Meckelian sulcus, a straight profile of the ventral surface of the dentary and a distal thickening of the acromion process of the pubis between other characters. Iberospinus natarioi is recovered as a sister taxon of the clade formed by Baryonyx and Suchomimus, and outside Spinosaurinae when Vallibonaventrix cani is excluded from the analysis. The description of this taxon reinforces Iberia as a hotspot for spinosaur biodiversity, with several endemic taxa for the region. As expected for the clade, the dentary displays a highly vascularized neurovascular network. The morphometric analysis of parts of the skeleton (pedal phalanx and caudal vertebrae, among others) shows an intermediate condition between basal tetanurans and spinosaurines.

Fernandes, A. E., Mateus O., Bauluz B., Coimbra R., Ezquerro L., Núñez-Lahuerta C., Suteu C., & Moreno-Azanza M. (2021).  The Paimogo Dinosaur Egg Clutch Revisited: Using One of Portugal’s Most Notable Fossils to Exhibit the Scientific Method. Geoheritage. 13(3), 66., 2021 Abstractfernandes_et_al-2021-geoheritage.pdfWebsite

Found in the Upper Jurassic outcrops of Lourinhã, Portugal, and first published in 1997, the Paimogo dinosaur egg clutch is one of Portugal’s most remarkable fossils, with over one hundred eggs preserved in association with embryonic bones, of the allosauroid theropod Lourinhanosaurus. However, many questions about it have remained unanswered, even until the present day. After its discovery, this extraordinary fossil became the keystone of a small local museum, greatly kick-starting regional tourism, while also holding the fossils in trust for future generations to study. More than 20 years later, continually sustained paleontological interest from the public has even given rise to both a highly successful dinosaur theme park in the region and an aspiring UNESCO Geopark. Recently, a multidisciplinary team of preparators, paleontologists, sedimentologists, mineralogists, and geochemists revisited an unopened jacket from the original excavation using an array of techniques to address various questions. Studies are ongoing, but the corpus of information obtained and the methodologies utilized to gather data have offered an opportunity to design an exhibit around the history of the Paimogo clutch, highlighting the scientific methods involved, and asserting the importance of preserving geological heritage for the future, when new tools will doubtlessly become available to provide yet another new look at old fossils. Here, we describe our analytical procedures and present an innovative exhibit designed to introduce to the public the latest advances on the research behind an iconic piece of Portuguese geoheritage, increasing its value both as a research item and as an educational resource.

Ceríaco, L. M. P., Gutiérrez E. E., Dubois A., Abdala C. S., Alqarni A. S., Adler K., et al. (2016).  Photography-based taxonomy is inadequate, unnecessary, and potentially harmful for biological sciences. Zootaxa. 4196(3), 435 - 445., 2016 AbstractWebsite
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Ceríaco, L. M. P., Gutiérrez E. E., Dubois A., Abdala C. S., Alqarni A. S., Adler K., et al. (2016).  Photography-based taxonomy is inadequate, unnecessary, and potentially harmful for biological sciences. Zootaxa. 4196, 435-445., Number 3 Abstract
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Milàn, J., Clemmensen L. B., Adolfssen J. S., Estrup E. J., Frobøse N., Klein N., Mateus O., & Wings O. (2012).  A preliminary report on coprolites from the Late Triassic part of the Kap Stewart Formation, Jameson Land, East Greenland. New Mexico Museum of Natural History and Science, Bulletin. 57, 203-205. Abstractmilan_et_al_2012_greenland_coprolites_triassic.pdf

The basal part of the Triassic-Jurassic (Rhaetian-Sinemurian) Kap Stewart Formation, exposed at Jameson Land, East Greenland, yields an extensive coprolite collection from black, parallel-laminated mudstone (“paper shale”), representing an open lacustrine system. Preliminary investigations show three different types of coprolites: elongated cylindrical masses, composed of irregularly wrapped layers; elongated cylindrical masses with constriction marks; and spirally-coiled specimens.

Mateus, O., Estraviz López D., & Madeira Mateus S. G. (2022).  Type specimens alone have a strong correlation with taxa record by geological epoch: the case study of the fossil vertebrates named from {Portuguese} types. Comunicações Geológicas. 109, 57–64. Abstractmateus-et-al_2022_type_specimens.pdfWebsite

ABSTRACT: Type specimens (holotypes, neotypes, syntypes, etc.) are of crucial importance because they are the only tangible evidence of the nomenclatural act that supports the understanding of paleobiodiversity. The list of the vertebrate species whose type specimen is based on fossils from Portugal is presented here. We counted 206 species, of which there are 45 bony fishes, 39 Late Jurassic Mammaliamorpha, 33 Cenozoic mammals, 28 non-avian dinosaurs, 25 non-dinosaur reptiles, 11 Cenozoic Aves, 22 Chondrichthyes and three other vertebrates. Except for the Quaternary fossil record, the type specimens can be used as a shortcut for measuring the fossil record and paleobiodiversity through geological time and rock units because they correlate in 95% with the fossil record by epoch in the case-study of fossil vertebrates of Portugal.

Moreno-Azanza, M., Balauz B., Ezquerro L., Coimbra R., Mateus O., & Nuñez-Lahuerta C. (2022).  Unbroken dinosaur eggs accumulated together do not guarantee autochthony of the fossil assemblage. Journal of Taphonomy. 16(1-4), 129-130. Abstractmoreno-azanza_et_al_2020_unbroken_eggs.pdf

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Guillaume, A. R. D., Ezquerro L., Moreno-Azanza M., & Mateus O. (2023).  Vertebrate microfossil assemblages from the Lourinhã Formation: a sneak peek on the paleoecology of the Late Jurassic in Portugal. The Anatomical Record. 306(#S1), 118-121.: Conference: 14th Symposium on Mesozoic Terrestrial Ecosystems and BiotaAt … Abstractguillaume_et_al_2023_vertebrate_microfossil.pdf

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Clemmensen, L. B., Milàn J., Adolfssen J. S., Estrup E. J., Frobøse N., Klein N., Mateus O., & Wings O. (2015).  The vertebrate-bearing Late Triassic Fleming Fjord Formation of central East Greenland revisited: stratigraphy, palaeoclimate and new palaeontological data. Geological Society, London, Special Publications. 434(1), 31-47. Abstractclemmensenetal2015greenland.pdfWebsite

In Late Triassic (Norian–Rhaetian) times, the Jameson Land Basin lay at 40° N on the northern part of the supercontinent Pangaea. This position placed the basin in a transition zone between the relatively dry interior of the supercontinent and its more humid periphery. Sedimentation in the Jameson Land Basin took place in a lake–mudflat system and was controlled by orbitally forced variations in precipitation. Vertebrate fossils have consistently been found in these lake deposits (Fleming Fjord Formation), and include fishes, dinosaurs, amphibians, turtles, aetosaurs and pterosaurs. Furthermore, the fauna includes mammaliaform teeth and skeletal material. New vertebrate fossils were found during a joint vertebrate palaeontological and sedimentological expedition to Jameson Land in 2012. These new finds include phytosaurs, a second stem testudinatan specimen and new material of sauropodomorph dinosaurs, including osteologically immature individuals. Phytosaurs are a group of predators common in the Late Triassic, but previously unreported from Greenland. The finding includes well-preserved partial skeletons that show the occurrence of four individuals of three size classes. The new finds support a late Norian–early Rhaetian age for the Fleming Fjord Formation, and add new information on the palaeogeographical and palaeolatitudinal distribution of Late Triassic faunal provinces.

Clemmensen, L. B., Milàn J., Adolfssen J. S., Estrup E. J., Frobøse N., Klein N., Mateus O., & Wings O. (2015).  The vertebrate-bearing Late Triassic Fleming Fjord Formation of central East Greenland revisited: stratigraphy, palaeoclimate and new palaeontological data. Geological Society, London, Special Publications. 434, AbstractWebsite

In Late Triassic (Norian–Rhaetian) times, the Jameson Land Basin lay at 40° N on the northern part of the supercontinent Pangaea. This position placed the basin in a transition zone between the relatively dry interior of the supercontinent and its more humid periphery. Sedimentation in the Jameson Land Basin took place in a lake–mudflat system and was controlled by orbitally forced variations in precipitation. Vertebrate fossils have consistently been found in these lake deposits (Fleming Fjord Formation), and include fishes, dinosaurs, amphibians, turtles, aetosaurs and pterosaurs. Furthermore, the fauna includes mammaliaform teeth and skeletal material. New vertebrate fossils were found during a joint vertebrate palaeontological and sedimentological expedition to Jameson Land in 2012. These new finds include phytosaurs, a second stem testudinatan specimen and new material of sauropodomorph dinosaurs, including osteologically immature individuals. Phytosaurs are a group of predators common in the Late Triassic, but previously unreported from Greenland. The finding includes well-preserved partial skeletons that show the occurrence of four individuals of three size classes. The new finds support a late Norian–early Rhaetian age for the Fleming Fjord Formation, and add new information on the palaeogeographical and palaeolatitudinal distribution of Late Triassic faunal provinces.

Jackson, Y., Economos R., Jacobs L., Mateus O., & Gonçalves A. O. (2021).  When Dinosaurs Walked Through Diamonds: Constraining the Age of Early Cretaceous Footprints in Volcanic Crater Sediments. SMU Journal of Undergraduate Research. 6(1), : DOI: https://doi.org/ 10.25172/jour.6.1.1 Available at: https://scholar.smu … Abstractwhen_dinosaurs_walked_through_diamonds.pdf

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