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2015
Brusatte, S. L., Butler R. J., Mateus O., & Steyer J. S. (2015).  A new species of Metoposaurus from the Late Triassic of Portugal and comments on the systematics and biogeography of metoposaurid temnospondyls. Journal of Vertebrate Paleontology. 35, , Number 3 Abstract
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Xing, L., Lockley M. G., Marty D., Zhang J., Wang Y., Klein H., McCrea R. T., Buckley L. G., Belvedere M., Mateus O., Gierli?ski G. D., Piñuela L., Persons W. S., Wang F., Ran H., Dai H., & Xie X. (2015).  An ornithopod-dominated tracksite from the lower Cretaceous Jiaguan Formation (Barremian-Albian) of Qijiang, South-Central China: New discoveries, ichnotaxonomy, preservation and palaeoecology. PLoS ONE. 10, , Number 10 Abstract
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Tschopp, E., Mateus O., & Benson R. B. J. (2015).  A specimen-level phylogenetic analysis and taxonomic revision of Diplodocidae (Dinosauria, Sauropoda). {PeerJ}. 3, e857.: {PeerJ} AbstractWebsite
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Young, M. T., Hua S., Steel L., Foffa D., Brusatte S. L., Thüring S., Mateus O., Ruiz-Omeñaca J. I., Havlik P., Lepage Y., & de Andrade M. B. (2015).  Addendum to ‘Revision of the Late Jurassic teleosaurid genus Machimosaurus (Crocodylomorpha, Thalattosuchia)’. Royal Society Open Science. 2, , Number 2: The Royal Society Abstractyoung_et_al_2015_addendum_to_revision_of_the_late_jurassic_teleosaurid_genus_machimosaurus_crocodylomorpha_thalattosuchia.pdfWebsite

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Pereira, B. C., Benton M. J., Ruta M., & Mateus O. (2015).  Mesozoic echinoid diversity in Portugal: Investigating fossil record quality and environmental constraints on a regional scale. Palaeogeography, Palaeoclimatology, Palaeoecology. 424, 132 - 146. Abstractpereira_e_al_2015_mesozoic_echinoids_portugal.pdfWebsite

Abstract Several analyses of diversity through geological time use global, synoptic databases, and this practice often makes it difficult to distinguish true signals in changing diversity from regional-scale sampling and/or geological artefacts. Here we investigate how echinoid diversity changed through the Mesozoic of the Lusitanian basin in Portugal based on a comprehensive, revised database, and seek to distinguish biological signal from geological or environmental constraints. The observed diversity pattern is far from having a defined trend, showing many fluctuations that appear to be linked with gaps in the geological record. This study revealed that, independently of the method used, whether correlation tests or model fitting, the diversity signal is not completely explained by the studied sampling proxies. Among the different proxies, marine facies variation in combination with outcrop area best explains the palaeodiversity curve.

Xing, L., Lockley M. G., Marty D., Zhang J., Wang Y., Klein H., McCrea R. T., Buckley L. G., Belvedere M., Mateus O., Gierliński G. D., Piñuela L., Persons, IV S. W., Wang F., Ran H., Dai H., & Xie X. (2015).  An Ornithopod-Dominated Tracksite from the Lower Cretaceous Jiaguan Formation (Barremian–Albian) of Qijiang, South-Central China: New Discoveries, Ichnotaxonomy, Preservation and Palaeoecology. PLoS ONE. 10, e0141059., 10, Number 10: Public Library of Science Abstractlida_et_al_2015_an_ornithopod-dominated_tracksite_from_the.pdfWebsite

The historically-famous Lotus Fortress site, a deep 1.5–3.0-meter-high, 200-meter-long horizonal notch high up in near-vertical sandstone cliffs comprising the Cretaceous Jiaguan Formation, has been known since the 13th Century as an impregnable defensive position. The site is also extraordinary for having multiple tetrapod track-bearing levels, of which the lower two form the floor of part of the notch, and yield very well preserved asseamblages of ornithopod, bird (avian theropod) and pterosaur tracks. Trackway counts indicate that ornithopods dominate (69%) accounting for at least 165 trackmakers, followed by bird (18%), sauropod (10%), and pterosaur (3%). Previous studies designated Lotus Fortress as the type locality of Caririchnium lotus and Wupus agilis both of which are recognized here as valid ichnotaxa. On the basis of multiple parallel trackways both are interpreted as representing the trackways of gregarious species. C. lotus is redescribed here in detail and interpreted to indicate two age cohorts representing subadults that were sometimes bipedal and larger quadrupedal adults. Two other previously described dinosaurian ichnospecies, are here reinterpreted as underprints and considered nomina dubia. Like a growing number of significant tetrapod tracksites in China the Lotus Fortress site reveals new information about the composition of tetrapod faunas from formations in which the skeletal record is sparse. In particular, the site shows the relatively high abundance of Caririchium in a region where saurischian ichnofaunas are often dominant. It is also the only site known to have yielded Wupus agilis. In combination with information from other tracksites from the Jiaguan formation and other Cretaceous formations in the region, the track record is proving increasingly impotant as a major source of information on the vertebrate faunas of the region. The Lotus Fortress site has been developed as a spectacular, geologically-, paleontologically- and a culturally-significant destination within Qijiang National Geological Park.

Brusatte, S. L., Butler R. J., Mateus O., & Steyer S. J. (2015).  A new species of Metoposaurus from the Late Triassic of Portugal and comments on the systematics and biogeography of metoposaurid temnospondyls. Journal of Vertebrate PaleontologyJournal of Vertebrate Paleontology. e912988., 2015: Taylor & Francis Abstractbrusatte_et_al2015metoposaurusportugal.pdfWebsite

ABSTRACTMetoposaurids are a group of temnospondyl amphibians that filled crocodile-like predatory niches in fluvial and lacustrine environments during the Late Triassic. Metoposaurids are common in the Upper Triassic sediments of North Africa, Europe, India, and North America, but many questions about their systematics and phylogeny remain unresolved. We here erect Metoposaurus algarvensis, sp. nov., the first Metoposaurus species from the Iberian Peninsula, based on several new specimens from a Late Triassic bonebed in Algarve, southern Portugal. We describe the cranial and pectoral anatomy of M. algarvensis and compare it with other metoposaurids (particularly other specimens of Metoposaurus from Germany and Poland). We provide a revised diagnosis and species-level taxonomy for the genus Metoposaurus, which is currently represented with certainty by three European species (M. diagnosticus, M. krasiejowensis, M. algarvensis). We also identify cranial characters that differentiate these three species, and may have phylogenetic significance. These include features of the braincase and mandible, which indicate that metoposaurid skulls are more variable than previously thought. The new Portuguese bonebed provides further evidence that metoposaurids congregated in fluvial and lacustrine settings across their geographic range and often succumbed to mass death events. We provide an updated paleogeographic map depicting all known metoposaurid occurrences, which shows that these temnospondyls were globally distributed in low latitudes during the Late Triassic and had a similar, but not identical, paleogeographic range as phytosaurs.http://zoobank.org/urn:lsid:zoobank.org:pub:083C80C6-0AB6-49E1-A636-6A8BDBC06A47ABSTRACTMetoposaurids are a group of temnospondyl amphibians that filled crocodile-like predatory niches in fluvial and lacustrine environments during the Late Triassic. Metoposaurids are common in the Upper Triassic sediments of North Africa, Europe, India, and North America, but many questions about their systematics and phylogeny remain unresolved. We here erect Metoposaurus algarvensis, sp. nov., the first Metoposaurus species from the Iberian Peninsula, based on several new specimens from a Late Triassic bonebed in Algarve, southern Portugal. We describe the cranial and pectoral anatomy of M. algarvensis and compare it with other metoposaurids (particularly other specimens of Metoposaurus from Germany and Poland). We provide a revised diagnosis and species-level taxonomy for the genus Metoposaurus, which is currently represented with certainty by three European species (M. diagnosticus, M. krasiejowensis, M. algarvensis). We also identify cranial characters that differentiate these three species, and may have phylogenetic significance. These include features of the braincase and mandible, which indicate that metoposaurid skulls are more variable than previously thought. The new Portuguese bonebed provides further evidence that metoposaurids congregated in fluvial and lacustrine settings across their geographic range and often succumbed to mass death events. We provide an updated paleogeographic map depicting all known metoposaurid occurrences, which shows that these temnospondyls were globally distributed in low latitudes during the Late Triassic and had a similar, but not identical, paleogeographic range as phytosaurs.http://zoobank.org/urn:lsid:zoobank.org:pub:083C80C6-0AB6-49E1-A636-6A8BDBC06A47

Tschopp, E., Mateus O., & Benson R. B. J. (2015).  A specimen-level phylogenetic analysis and taxonomic revision of Diplodocidae (Dinosauria, Sauropoda). PeerJ. 3, e857., 4 Abstracttschopp_et_al_2015_brontosaurus_peerj-857.pdfWebsite

Diplodocidae are among the best known sauropod dinosaurs. Several species were described in the late 1800s or early 1900s from the Morrison Formation of North America. Since then, numerous additional specimens were recovered in the USA, Tanzania, Portugal, and Argentina, as well as possibly Spain, England, Georgia, Zimbabwe, and Asia. To date, the clade includes about 12 to 15 nominal species, some of them with questionable taxonomic status (e.g., ‘\textit{Diplodocus}’ \textit{hayi} or \textit{Dyslocosaurus polyonychius}), and ranging in age from Late Jurassic to Early Cretaceous. However, intrageneric relationships of the iconic, multi-species genera \textit{Apatosaurus} and \textit{Diplodocus} are still poorly known. The way to resolve this issue is a specimen-based phylogenetic analysis, which has been previously implemented for \textit{Apatosaurus}, but is here performed for the first time for the entire clade of Diplodocidae.The analysis includes 81 operational taxonomic units, 49 of which belong to Diplodocidae. The set of OTUs includes all name-bearing type specimens previously proposed to belong to Diplodocidae, alongside a set of relatively complete referred specimens, which increase the amount of anatomically overlapping material. Non-diplodocid outgroups were selected to test the affinities of potential diplodocid specimens that have subsequently been suggested to belong outside the clade. The specimens were scored for 477 morphological characters, representing one of the most extensive phylogenetic analyses of sauropod dinosaurs. Character states were figured and tables given in the case of numerical characters.The resulting cladogram recovers the classical arrangement of diplodocid relationships. Two numerical approaches were used to increase reproducibility in our taxonomic delimitation of species and genera. This resulted in the proposal that some species previously included in well-known genera like \textit{Apatosaurus} and \textit{Diplodocus} are generically distinct. Of particular note is that the famous genus \textit{Brontosaurus} is considered valid by our quantitative approach. Furthermore, “\textit{Diplodocus}” hayi represents a unique genus, which will herein be called \textit{Galeamopus} gen. nov. On the other hand, these numerical approaches imply synonymization of “\textit{Dinheirosaurus}” from the Late Jurassic of Portugal with the Morrison Formation genus \textit{Supersaurus}. Our use of a specimen-, rather than species-based approach increases knowledge of intraspecific and intrageneric variation in diplodocids, and the study demonstrates how specimen-based phylogenetic analysis is a valuable tool in sauropod taxonomy, and potentially in paleontology and taxonomy as a whole.

2016
Polcyn, M. J., Bardet N., Amaghzaz M., Gon{\c c}alves O. A., Jourani E., Kaddumi H. F., Lindgren J., Mateus O., Meslouhf S., Morais M. L., Pereda-Suberbiola X., Schulp A. S., Vincent P., & Jacobs L. L. (2016).  An extremely derived plioplatecarpine mosasaur from the Maastrichtian of Africa and the Middle East. 5th Triennial Mosasaur Meeting- a global perspective on Mesozoic marine amniotes. 16-20, May 16-20, 2016., Uppsala, Sweden: Museum of Evolutiom, Uppsala University. Abstract

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Polcyn, M. J., Bardet N., Amaghzaz M., Gon\{\c c\}alves O. A., Jourani E., Kaddumi H. F., Lindgren J., Mateus O., Meslouhf S., Morais M. L., Pereda-Suberbiola X., Schulp A. S., Vincent P., & Jacobs L. L. (2016).  An extremely derived plioplatecarpine mosasaur from the Maastrichtian of Africa and the Middle East. 5th Triennial Mosasaur Meeting- a global perspective on Mesozoic marine amniotes. 16-20, May 16-20, 2016., Uppsala, Sweden Abstract
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Polcyn, {M. J. }, Bardet N., Amaghzaz M., Gonçalves {A. O. }, Jourani E., Kaddumi {H. F. }, Lindgren J., Mateus O., Meslouhf S., & Morais {M. L. } (2016).  An extremely derived plioplatecarpine mosasaur from the Maastrichtian of Africa and the Middle East. 32–33. Abstract
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Leal, A. A., Dion{\'ısio A., Braga M. A. S., & Mateus O. (2016).  The long term preservation of Late Jurassic sandstone dinosaur footprints in a museum environment. International Journal of Conservation Science. 7, 627-646. AbstractWebsite

This study focuses on the assessment of the degradation processes occurring in three sandstone infills of fossilized Late Jurassic ornithopod tridactyl footprints, found in 2001 in a coastline cliff in Porto das Barcas (Lourinhã, Portugal) and exhibited in a museum display since 2004. These dinosaur footprints present nowadays severe decay phenomena compromising their physical integrity and are leading gradually to their loss of value. The deterioration patterns were recorded, a map of their distribution was prepared and several samples were collected both in the dinosaur footprints and in the coastline cliff. Different analytical procedures were applied such as XRD, FTIR, FESEM and Ion Chromatography. A microclimatic survey was also performed and air temperature and relative humidity was measured during eight months both indoor and also outdoor. The decay patterns observed are a combination intrinsic and extrinsic factors the stone material, namely swelling of clay minerals in the rock matrix (smectite and chlorite-smectite mixed-layer), presence of salts (mainly chlorides), application of past conservation treatments (poly(vinyl) acetate and epoxy resins) and with the museum’s indoor thermohygrometric conditions (mainly non-stable hygrometric conditions). This scientific knowledge is therefore essential to the sustainable preservation of this paleontological heritage.

Leal, A. A., Dion\{\'ı\}sio A., Braga M. A. S., & Mateus O. (2016).  The long term preservation of Late Jurassic sandstone dinosaur footprints in a museum environment. International Journal of Conservation Science. 7, 627-646. Abstract
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Ceríaco, L. M. P., Gutiérrez E. E., Dubois A., Abdala C. S., Alqarni A. S., Adler K., et al. (2016).  Photography-based taxonomy is inadequate, unnecessary, and potentially harmful for biological sciences. Zootaxa. 4196, 435-445., Number 3 Abstract
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Polcyn, M. J., Bardet N., Amaghzaz M., Gonçalves O. A., Jourani E., Kaddumi H. F., Lindgren J., Mateus O., Meslouhf S., Morais M. L., Pereda-Suberbiola X., Schulp A. S., Vincent P., & Jacobs L. L. (2016).  An extremely derived plioplatecarpine mosasaur from the Maastrichtian of Africa and the Middle East. 5th Triennial Mosasaur Meeting- a global perspective on Mesozoic marine amniotes. 16-20(May 16-20, 2016), May 16-20, 2016., Uppsala, Sweden: Museum of Evolutiom, Uppsala University. Abstractpolcyn_et_al_2016_extremely_derived_mosasaur.pdf

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Leal, A. A., Dionísio A., Braga M. A. S., & Mateus O. (2016).  The long term preservation of Late Jurassic sandstone dinosaur footprints in a museum environment. International Journal of Conservation Science. 7(3), 627-646. AbstractWebsite

This study focuses on the assessment of the degradation processes occurring in three sandstone infills of fossilized Late Jurassic ornithopod tridactyl footprints, found in 2001 in a coastline cliff in Porto das Barcas (Lourinhã, Portugal) and exhibited in a museum display since 2004. These dinosaur footprints present nowadays severe decay phenomena compromising their physical integrity and are leading gradually to their loss of value. The deterioration patterns were recorded, a map of their distribution was prepared and several samples were collected both in the dinosaur footprints and in the coastline cliff. Different analytical procedures were applied such as XRD, FTIR, FESEM and Ion Chromatography. A microclimatic survey was also performed and air temperature and relative humidity was measured during eight months both indoor and also outdoor. The decay patterns observed are a combination intrinsic and extrinsic factors the stone material, namely swelling of clay minerals in the rock matrix (smectite and chlorite-smectite mixed-layer), presence of salts (mainly chlorides), application of past conservation treatments (poly(vinyl) acetate and epoxy resins) and with the museum's indoor thermohygrometric conditions (mainly non-stable hygrometric conditions). This scientific knowledge is therefore essential to the sustainable preservation of this paleontological heritage.

Hendrickx, C., Mateus O., & Buffetaut E. (2016).  Morphofunctional Analysis of the Quadrate of Spinosauridae (Dinosauria: Theropoda) and the Presence of Spinosaurus and a Second Spinosaurine Taxon in the Cenomanian of North Africa.. PLoS ONE. 11, e0144695., 01, Number 1: Public Library of Science Abstracthendrickx_et_al_2016_morphofunctional_analysis_of_the_quadrate_of_spinosauridae_dinosauria.pdfWebsite

Six quadrate bones, of which two almost certainly come from the Kem Kem beds (Cenomanian, Upper Cretaceous) of south-eastern Morocco, are determined to be from juvenile and adult individuals of Spinosaurinae based on phylogenetic, geometric morphometric, and phylogenetic morphometric analyses. Their morphology indicates two morphotypes evidencing the presence of two spinosaurine taxa ascribed to Spinosaurus aegyptiacus and? Sigilmassasaurus brevicollis in the Cenomanian of North Africa, casting doubt on the accuracy of some recent skeletal reconstructions which may be based on elements from several distinct species. Morphofunctional analysis of the mandibular articulation of the quadrate has shown that the jaw mechanics was peculiar in Spinosauridae. In mature spinosaurids, the posterior parts of the two mandibular rami displaced laterally when the jaw was depressed due to a lateromedially oriented intercondylar sulcus of the quadrate. Such lateral movement of the mandibular ramus was possible due to a movable mandibular symphysis in spinosaurids, allowing the pharynx to be widened. Similar jaw mechanics also occur in some pterosaurs and living pelecanids which are both adapted to capture and swallow large prey items. Spinosauridae, which were engaged, at least partially, in a piscivorous lifestyle, were able to consume large fish and may have occasionally fed on other prey such as pterosaurs and juvenile dinosaurs.

Hendrickx, C., Mateus O., & Buffetaut E. (2016).  Morphofunctional Analysis of the Quadrate of Spinosauridae (Dinosauria: Theropoda) and the Presence of Spinosaurus and a Second Spinosaurine Taxon in the Cenomanian of North Africa.. PLoS ONE. 11, e0144695., 01, Number 1: Public Library of Science AbstractWebsite

Six quadrate bones, of which two almost certainly come from the Kem Kem beds (Cenomanian, Upper Cretaceous) of south-eastern Morocco, are determined to be from juvenile and adult individuals of Spinosaurinae based on phylogenetic, geometric morphometric, and phylogenetic morphometric analyses. Their morphology indicates two morphotypes evidencing the presence of two spinosaurine taxa ascribed to Spinosaurus aegyptiacus and? Sigilmassasaurus brevicollis in the Cenomanian of North Africa, casting doubt on the accuracy of some recent skeletal reconstructions which may be based on elements from several distinct species. Morphofunctional analysis of the mandibular articulation of the quadrate has shown that the jaw mechanics was peculiar in Spinosauridae. In mature spinosaurids, the posterior parts of the two mandibular rami displaced laterally when the jaw was depressed due to a lateromedially oriented intercondylar sulcus of the quadrate. Such lateral movement of the mandibular ramus was possible due to a movable mandibular symphysis in spinosaurids, allowing the pharynx to be widened. Similar jaw mechanics also occur in some pterosaurs and living pelecanids which are both adapted to capture and swallow large prey items. Spinosauridae, which were engaged, at least partially, in a piscivorous lifestyle, were able to consume large fish and may have occasionally fed on other prey such as pterosaurs and juvenile dinosaurs.

Ceríaco, L. M. P., Gutiérrez E. E., Dubois A., Abdala C. S., Alqarni A. S., Adler K., et al. (2016).  Photography-based taxonomy is inadequate, unnecessary, and potentially harmful for biological sciences. Zootaxa. 4196(3), 435 - 445., 2016 AbstractWebsite
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2017
Correia, T., Barcelos L., Nunes T., Riff D., & Mateus O. (2017).  On a titanosaur scapula from the Marília Formation (Upper Cretaceous, Bauru Group) in Campina Verde Town. 77. Abstract
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Correia, T., Barcelos L., Nunes T., Riff D., & Mateus O. (2017).  On a titanosaur scapula from the Mar{\'ılia Formation (Upper Cretaceous, Bauru Group) in Campina Verde Town. XXV Congresso Brasileiro de Paleontologia Boletim de Resumos. 77. Abstract
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Russo, J., Mateus O., Marzola M., & Balbino A. (2017).  Two new ootaxa from the late Jurassic: The oldest record of crocodylomorph eggs, from the Lourinhã Formation, Portugal. PLoS ONE. 12, , Number 3 Abstract
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Correia, T., Barcelos L., Nunes T., Riff D., & Mateus O. (2017).  On a titanosaur scapula from the Marília Formation (Upper Cretaceous, Bauru Group) in Campina Verde Town. XXV Congresso Brasileiro de Paleontologia Boletim de Resumos. 77. Abstractthiago_abstract_brazil_2017.pdf

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Russo, J., Mateus O., Marzola M., & Balbino A. (2017).  Two new ootaxa from the late Jurassic: The oldest record of crocodylomorph eggs, from the Lourinhã Formation, Portugal. PLOS ONE. 12, 1-23., 03, Number 3: Public Library of Science Abstractrusso_et_al_2017_two_new_ootaxa.pdfWebsite

The Late Jurassic Lourinhã Formation is known for its abundant remains of dinosaurs, crocodylomorphs and other vertebrates. Among this record are nine localities that have produced either dinosaur embryos, eggs or eggshell fragments. Herein, we describe and identify the first crocodiloid morphotype eggs and eggshells from the Lourinhã Formation, from five occurrences. One clutch from Cambelas, composed of 13 eggs, eggshell fragments from Casal da Rola and Peralta, one crushed egg and eggshells from Paimogo North, and four crushed eggs as well as eggshell fragments from Paimogo South. We observed and confirmed diagnostic morphological characters for crocodiloid eggshells and which are consistent with a crocodylomorph affinity, such as the ellipsoidal shape, wedge-shaped shell units, triangular extinction under cross-polarized light, and tabular ultrastructure. This material is distinctive enough to propose two new ootaxa within the oofamily Krokolithidae, Suchoolithus portucalensis, oogen. and oosp. nov., for the material from Cambelas, the most complete clutch known for crocodiloid eggs, and Krokolithes dinophilus, oosp. nov., for the remaining material. These are the oldest crocodylomorph eggs known, extending the fossil record for this group to the Late Jurassic. Furthermore, except for the clutch from Cambelas, the material was found with theropod eggs and nests, in the other four occurrences, which seem to suggest some form of biological relationship, still unclear at this point.

Moreno-Azanza, M., Bauluz B., Canudo J. I., & Mateus O. (2017).  The conservative structure of the ornithopod eggshell: electron backscatter diffraction characterization of Guegoolithus turolensis from the Early Cretaceous of Spain. Journal of Iberian Geology. 1-9., 2017 Abstractthe_conservative_structure_of_the_ornithopod_eggshell.pdfWebsite

The Spheroolithidae oospecies Guegoolithus turolensis, putatively attributed to non-hadrosauroid styracosterns was first described in the Barremian of the Iberian Basin, and later reported in the Valanginian–Hauterivian of the Cameros Basin, with both occurrences separated by a few hundred kilometres but by over 10 million years.

Moreno-Azanza, M., Bauluz B., Canudo J. I., & Mateus O. (2017).  The conservative structure of the ornithopod eggshell: electron backscatter diffraction characterization of Guegoolithus turolensis from the Early Cretaceous of Spain. Journal of Iberian Geology. 43, 235–243., jun, Number 2: Springer Nature AbstractWebsite
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2018
Tschopp, E., Brinkman D., Henderson J., Turner M. A., & Mateus O. (2018).  Considerations on the replacement of a type species in the case of the sauropod dinosaur Diplodocus Marsh, 1878. Geology of the Intermountain West. 5, 245-262. Abstract
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Tschopp, E., Brinkman D., Henderson J., Turner M. A., & Mateus O. (2018).  Considerations on the replacement of a type species in the case of the sauropod dinosaur Diplodocus Marsh, 1878. Geology of the Intermountain West. 5, 245-262.tschoppetal2018.pdf
2019
Azanza, M. M., Coimbra R., Puértolas-Pascual E., Russo J., Bauluz B., & Mateus O. (2019).  Crystallography of Lourinhanosaurus eggshells (Dinosauria, Theropoda, Allosauroidea). Journal of Vertebrate Paleontology, Program and Abstracts. 156-157. Abstract
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Park, J., Lee Y., Currie P. J., Kobayashi Y., Koppelhus E. B., Barsbold R., Lee S., Kim S., & Mateus O. (2019).  Three new skulls of the Late Cretaceous armored dinosaur Talarurus plicatospineus Maleev, 1952. Journal of Vertebrate Paleontology, Program and Abstracts. 165-166. Abstract
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Azanza, M. M., Coimbra R., Puértolas-Pascual E., Russo J., Bauluz B., & Mateus O. (2019).  Crystallography of Lourinhanosaurus eggshells (Dinosauria, Theropoda, Allosauroidea). Journal of Vertebrate Paleontology, Program and Abstracts. 156-157.moreno_azanza_et_al_2019_svp_abstract.pdf
Belvedere, M., Castanera D., Meyer C. A., Marty D., Mateus O., Silva B. C., Santos V. F., & Cobos A. (2019).  Late Jurassic globetrotters compared: A closer look at large and giant theropod tracks of North Africa and Europe. Journal of African Earth Sciences. 158, 103547. Abstractbelvedere_et_al_2019_jurassic_globetrotters_compared.pdfWebsite

Late Jurassic theropod tracks are very common both in North Africa and Europe. Two recently described ichnotaxa Megalosauripus transjuranicus and Jurabrontes curtedulensis from the Kimmeridgian of Switzerland show the coexistence of two apex predators in the same palaeoenvironment. Similar tracks can be found in tracksites from the Iberian Peninsula and from Morocco. Here, we further explore the similarities among the Swiss ichnotaxa and the other tracks from Germany (Kimmeridgian), Spain (Tithonian-Berriasian), Portugal (Oxfordian-Tithonian) and Morocco (Kimmeridgian) through novel three-dimensional data comparisons. Specimens were grouped in two morphotypes: 1) large and gracile (30 < Foot Length<50 cm) and 2) giant and robust (FL > 50 cm). The analyses show a great morphological overlap among these two morphotypes and the Swiss ichnotaxa (Megalosauripus transjuranicus and Jurabrontes curtedulensis, respectively), even despite the differences in sedimentary environment and age. This suggests a widespread occurrence of similar ichnotaxa along the western margin of Tethys during the Late Jurassic. The new data support the hypothesis of a Gondwana-Laurasia faunal exchange during the Middle or early Late Jurassic, and the presence of migratory routes around the Tethys.

Mateus, O., Callapez P. M., Polcyn M. J., Schulp A. S., Gonçalves A. O., & Jacobs L. L. (2019).  O registo fóssil da biodiversidade em Angola ao longo do tempo: uma perspectiva paleontológica. (Huntley B.J., Russo V., Lages F., Ferrand N., Ed.).Biodiversidade de Angola: Ciência e Conservação - Uma Síntese Moderna. 89-116., Porto: Arte & Ciência Abstractmateus_et_al_2019_paleobiodiversidade_angola.pdf

Este capítulo apresenta uma visão geral da paleobiodiversidade alfa de Angola com base no registo fóssil disponível, o qual se limita às rochas sedimentares, a sua idade variando entre o Pré‑Câmbrico e o pre‑
sente. O período geológico com a maior paleobiodiversidade no registo fóssil angolano é o Cretácico, com mais de 80% do total dos táxones fósseis conhecidos, especialmente moluscos marinhos, sendo estes na sua maioria
amonites. Os vertebrados representam cerca de 15% da fauna conhecida e cerca de um décimo destes são espécies descritas pela primeira vez com base em espécimes de Angola.

Park, J., Lee Y., Currie P. J., Kobayashi Y., Koppelhus E. B., Barsbold R., Lee S., Kim S., & Mateus O. (2019).  Three new skulls of the Late Cretaceous armored dinosaur Talarurus plicatospineus Maleev, 1952. Journal of Vertebrate Paleontology, Program and Abstracts. 165-166.park_et_al_2019_svp_abstract.pdf
2020
Park, J. - Y., Lee Y. - N., Currie P. J., Kobayashi Y., Koppelhus E., Barsbold R., Mateus O., Lee S., & Kim S. - H. (2020).  Additional skulls of Talarurus plicatospineus (Dinosauria: Ankylosauridae) and implications for paleobiogeography and paleoecology of armored dinosaurs. Cretaceous Research. 108, 104340. Abstractpark_et_al_2020_additional_skulls_of_talarurus_plicatospineus_dinosauria_final.pdfWebsite

Three new additional skull specimens of Talarurus plicatospineus have been recovered from the Upper Cretaceous (Cenomanian–Santonian) Bayanshiree Formation, of Bayan Shiree cliffs, eastern Gobi Desert, Mongolia. The skulls feature unique characters such as an anteriorly protruded single internarial caputegulum, around 20 flat or concave nasal-area caputegulae surrounded by a wide sulcus, a vertically oriented elongate loreal caputegulum with a pitted surface, an elongate lacrimal caputegulum positioned above the posterodorsal border of the maxilla, two longitudinally arranged large frontoparietal caputegulae surrounded by smaller rhomboid caputegulae, small but elongate medial supraorbital caputegulae, a posterior supraorbital caputegulum that is four times larger than the anterior one, up to three transverse parallel grooves on the dorsal surface of the posterior supraorbital caputegulum, postocular caputegulae along the ventral to posterior rim of the orbit that extend almost to the anteroventral margin of the squamosal horn, a longitudinal furrow tapering towards the apex of the squamosal horn, a lateral nuchal caputegulum four to five times larger than other nuchal caputegulae, and a pterygovomerine keel with a ventral margin that is dorsally positioned to the alveolar ridge. The phylogenetic analysis result showed that Talarurus is sister to the clade that includes the derived Asian ankylosaurines (Saichania chulsanensis, Tarchia kielanae, and Zaraapelta nomadis). It also shows that there was dispersal of ankylosaurines from Asia into western North America before the Cenomanian. Moreover, the rostral differences between Talarurus and Tsagantegia, another ankylosaur from the same formation, suggest possible niche partitioning between these taxa.

2021
Ezquerro, L., Balauz B., Coimbra R., Nuñez-Lahuerta C., Mateus O., Román T., & Moreno-Azanza M. (2021).  Evidence of sedimentary remodel of Jurassic theropod egg clutches (Lourinhã, Portugal). 18th Conference of the EAVP-Abstract book. 72-73. Abstractezquerro_et_al_2021_sedimentary_remodel.pdf

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Moreno-Azanza, M., Mateus O., Bauluz B., Coimbra R., Ezquerro L., & Núñez-Lahuerta C. (2021).  Hatching in Portugal: a new look to old eggs. XIX Encontro de Jovens Investigadores em Paleontologia. 22. Abstractmoreno-azanza_et_al_2021_eggs_ejip.pdf

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Beccari, V., Mateus O., Wings O., Milàn J., & Clemmensen L. B. (2021).  Issi saaneq gen. et sp. nov.—A New Sauropodomorph Dinosaur from the Late Triassic (Norian) of Jameson Land, Central East Greenland. Diversity. 13, , Number 11 Abstractdiversity-13-00561-v2.pdfWebsite

The Late Triassic (Norian) outcrops of the Malmros Klint Formation, Jameson Land (Greenland) have yielded numerous specimens of non-sauropod sauropodomorphs. Relevant fossils were briefly reported in 1994 and were assigned to Plateosaurus trossingensis. However, continuous new findings of early non-sauropod sauropodomorphs around the globe facilitate comparisons and allow us to now revise this material. Here, the non-sauropod sauropodomorph Issi saaneq gen. et sp. nov. is described based on two almost complete and articulated skulls. The two skulls represent a middle-stage juvenile and a late-stage juvenile or subadult. Issi saaneq differs from all other sauropodomorphs by several unique traits: (1) a small foramen at the medial surface of the premaxilla; (2) an anteroposteriorly elongated dorsoposterior process of the squamosal; (3) a relatively high quadrate relative to rostrum height; (4) a well-developed posterodorsal process of the articular. These features cannot be explained by taphonomy, ontogeny, or intraspecific variation. Issi saaneq shows affinities to Brazilian plateosaurids and the European Plateosaurus, being recovered as the sister clade of the latter in our phylogenetic analysis. It is the northernmost record of a Late Triassic sauropodomorph, and a new dinosaur species erected for Greenland. Issi saaneq broadens our knowledge about the evolution of plateosaurid sauropodomorphs.

Fernandes, A. E., Mateus O., Bauluz B., Coimbra R., Ezquerro L., Núñez-Lahuerta C., Suteu C., & Moreno-Azanza M. (2021).  The Paimogo Dinosaur Egg Clutch Revisited: Using One of Portugal’s Most Notable Fossils to Exhibit the Scientific Method. Geoheritage. 13(3), 66., 2021 Abstractfernandes_et_al-2021-geoheritage.pdfWebsite

Found in the Upper Jurassic outcrops of Lourinhã, Portugal, and first published in 1997, the Paimogo dinosaur egg clutch is one of Portugal’s most remarkable fossils, with over one hundred eggs preserved in association with embryonic bones, of the allosauroid theropod Lourinhanosaurus. However, many questions about it have remained unanswered, even until the present day. After its discovery, this extraordinary fossil became the keystone of a small local museum, greatly kick-starting regional tourism, while also holding the fossils in trust for future generations to study. More than 20 years later, continually sustained paleontological interest from the public has even given rise to both a highly successful dinosaur theme park in the region and an aspiring UNESCO Geopark. Recently, a multidisciplinary team of preparators, paleontologists, sedimentologists, mineralogists, and geochemists revisited an unopened jacket from the original excavation using an array of techniques to address various questions. Studies are ongoing, but the corpus of information obtained and the methodologies utilized to gather data have offered an opportunity to design an exhibit around the history of the Paimogo clutch, highlighting the scientific methods involved, and asserting the importance of preserving geological heritage for the future, when new tools will doubtlessly become available to provide yet another new look at old fossils. Here, we describe our analytical procedures and present an innovative exhibit designed to introduce to the public the latest advances on the research behind an iconic piece of Portuguese geoheritage, increasing its value both as a research item and as an educational resource.

Beccari, V., Pinheiro F. L., Nunes I., Anelli L. E., Mateus O., & Costa F. R. (2021).  Osteology of an exceptionally well-preserved tapejarid skeleton from Brazil: Revealing the anatomy of a curious pterodactyloid clade. PLOS ONE. 16(8), e0254789 - ., 2021/08/25: Public Library of Science Abstractbeccari_et_al_2021.pdfWebsite

A remarkably well-preserved, almost complete and articulated new specimen (GP/2E 9266) of Tupandactylus navigans is here described for the Early Cretaceous Crato Formation of Brazil. The new specimen comprises an almost complete skeleton, preserving both the skull and post-cranium, associated with remarkable preservation of soft tissues, which makes it the most complete tapejarid known thus far. CT-Scanning was performed to allow the assessment of bones still covered by sediment. The specimen can be assigned to Tupa. navigans due to its vertical supra-premaxillary bony process and short and rounded parietal crest. It also bears the largest dentary crest among tapejarine pterosaurs and a notarium, which is absent in other representatives of the clade. The new specimen is here regarded as an adult individual. This is the first time that postcranial remains of Tupa. navigans are described, being also an unprecedented record of an articulated tapejarid skeleton from the Araripe Basin.

2022
Rotatori, F. M., Camilo B., Bertozzo F., Mateus O., & Moreno-Azanza M. (2022).  A basal ankylopollexian dinosaur from the Late Jurassic of Portugal and its implications for iguanodontian diversity. Abstract book of the XIX Annual Conference of the European Association of Vertebrate Palaeontologists, Benevento/Pietraroja, Italy, 27th June-2nd July 2022. 170-171.: PalaeoVertebrata, Special Volume 1- 2022, 224. Doi: 10.18563/pv.eavp Abstractrotatorietal_2022.pdf

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Moreno-Azanza, M., Pérez-Pueyo M., Puértolas-Pascual E., Núñez-Lahuerta C., Mateus O., Bauluz B., Bádenas B., & Canudo J. I. (2022).  Cáscaras de huevo de los últimos cocodrilomorfos del Cretácico (Huesca, España). XXXVII Jornadas de Paleontología SEP - V Congreso Ibérico de Paleontología. p. 119. Abstract2022_moreno-azanza_etal_sep.pdf

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Hendrickx, C., Bell P. R., Pittman M., Milner A. R. C., Cuesta E., O'Connor J., Loewen M., Currie P. J., Mateus O., Kaye T. G., & Delcourt R. (2022).  Morphology and distribution of scales, dermal ossifications, and other non-feather integumentary structures in non-avialan theropod dinosaurs. Biological Reviews. , Number n/a Abstracthendrickxetal.2021.morphologyanddistributionofscales.pdfWebsite

ABSTRACT Modern birds are typified by the presence of feathers, complex evolutionary innovations that were already widespread in the group of theropod dinosaurs (Maniraptoriformes) that include crown Aves. Squamous or scaly reptilian-like skin is, however, considered the plesiomorphic condition for theropods and dinosaurs more broadly. Here, we review the morphology and distribution of non-feathered integumentary structures in non-avialan theropods, covering squamous skin and naked skin as well as dermal ossifications. The integumentary record of non-averostran theropods is limited to tracks, which ubiquitously show a covering of tiny reticulate scales on the plantar surface of the pes. This is consistent also with younger averostran body fossils, which confirm an arthral arrangement of the digital pads. Among averostrans, squamous skin is confirmed in Ceratosauria (Carnotaurus), Allosauroidea (Allosaurus, Concavenator, Lourinhanosaurus), Compsognathidae (Juravenator), and Tyrannosauroidea (Santanaraptor, Albertosaurus, Daspletosaurus, Gorgosaurus, Tarbosaurus, Tyrannosaurus), whereas dermal ossifications consisting of sagittate and mosaic osteoderms are restricted to Ceratosaurus. Naked, non-scale bearing skin is found in the contentious tetanuran Sciurumimus, ornithomimosaurians (Ornithomimus) and possibly tyrannosauroids (Santanaraptor), and also on the patagia of scansoriopterygids (Ambopteryx, Yi). Scales are surprisingly conservative among non-avialan theropods compared to some dinosaurian groups (e.g. hadrosaurids); however, the limited preservation of tegument on most specimens hinders further interrogation. Scale patterns vary among and/or within body regions in Carnotaurus, Concavenator and Juravenator, and include polarised, snake-like ventral scales on the tail of the latter two genera. Unusual but more uniformly distributed patterning also occurs in Tyrannosaurus, whereas feature scales are present only in Albertosaurus and Carnotaurus. Few theropods currently show compelling evidence for the co-occurrence of scales and feathers (e.g. Juravenator, Sinornithosaurus), although reticulate scales were probably retained on the mani and pedes of many theropods with a heavy plumage. Feathers and filamentous structures appear to have replaced widespread scaly integuments in maniraptorans. Theropod skin, and that of dinosaurs more broadly, remains a virtually untapped area of study and the appropriation of commonly used techniques in other palaeontological fields to the study of skin holds great promise for future insights into the biology, taphonomy and relationships of these extinct animals.

Moreno-Azanza, M., Pérez-Pueyo M., Puértolas-Pascual E., Núñez-Lahuerta C., Mateus O., Bauluz B., Bádenas B., & Canudo J. I. (2022).  A new crocodylomorph related ootaxon from the late Maastrichtian of the Southern Pyrenees (Huesca, Spain). Historical Biology. 1-10.: Taylor & Francis Abstracta_new_crocodylomorph_related_ootaxon_from_the_late_maastrichtian_of_the_southern_pyrenees_huesca_spain.pdfWebsite

ABSTRACTCrocodylomorph eggs and eggshells are known as old as the Late Jurassic and are frequent components of most multiootaxic eggshell assemblages. Classified within the oofamily Krokolithidae, thei histo- and ultrastructures are conservative throughout geological time, characterised by inverted-trapezoid-shaped shell units that grow from highly spaced basal knobs and present a diagnostic tabular ultrastructure. Here, we report 327 eggshell fragments from a new fossil site from the Maastrichtian of the Southern Pyrenees, Veracruz 1, and erect a new oogenus and oospecies, Pachykrokolithus excavatum oogen. et oosp. nov. characterised by crocodyloid morphotype and a prominent rugosocavate ornamentation. Eggshells from the slightly older locality of Blasi 2b, previously reported as aff. Krokolithidae, are also assigned to this new ootaxon. Different crocodylomorph taxa coexisted during the Late Cretaceous of the Tremp Basin, hindering the attribution of Pachykrokolithus excavatum oogen. et oosp. nov. to a single clade. Nevertheless, allodaposuchid eusuchians were dominant in this ecosystem, and are the most probable producers of Pachykrokolithus excavatum oogen. et oosp. nov. eggs.

Fernandes, A. F., Beccari V., Kellner A. W. A., & Mateus O. (2022).  A new Gnathosaurine (Archaeopterodactyloidea, Pterosauria) from the Late Jurassic of Portugal. XIX Annual Conference of the European Association of Vertebrate Palaeontologists. 56-57. Abstractfernandes_et_al_2022_pterosaur_eavp_2022_abstract_volume.pdf

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Moreno-Azanza, M., Balauz B., Ezquerro L., Coimbra R., Mateus O., & Nuñez-Lahuerta C. (2022).  Unbroken dinosaur eggs accumulated together do not guarantee autochthony of the fossil assemblage. Journal of Taphonomy. 16(1-4), 129-130. Abstractmoreno-azanza_et_al_2020_unbroken_eggs.pdf

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2023
Rotatori, F. M., Quaranta M., Bertozzo F., Hübner T., Camilo B., Mateus O., & Moreno-Azanza M. (2023).  Hadrosaur-like vascularisation in the dentary of an early diverging iguanodontian dinosaur. Historical Biology. 1-6.: Taylor & Francis AbstractWebsite

ABSTRACTVirtual palaeontology is a growing field, leading palaeontologists to better understand the microanatomy of many extinct species. The application of techniques such as CT and μCT-scanning allows the researchers to study micro-anatomical features in a non-invasive way and make inferences on the palaeobiology of animals. Dinosaurs have been extensively studied using these techniques, with particular focus on the microanatomy of the cranium, whereas relatively little is known of other cranial elements, such as the lower jaw. Here, we aim to fill this gap, describing the microanatomy of the specimen ML 768, an isolated dentary belonging to a dryosaurid iguanodontian dinosaur from the Upper Jurassic of Lourinhã Fm. The dentary ML 768 was subjected to μCT-scanning, and subsequently the data were segmented in Avizo and rendered in Blender. We identified functional and replacement teeth, recognising remnants of old replacement cycles. Furthermore, we mapped a rich neurovascular network present in the dentary and compared it with reference literature. We found that the high vascularisation is shared with other cerapodan dinosaurs with high tooth replacement rates, although homoeostasis may have also played a role in the development of this condition. Further evidence is needed to appreciate the macroevolutionary significance of these findings.

Fernandes, A. E., Beccari V., Kellner A. W. A., & Mateus O. (2023).  A new gnathosaurine (Pterosauria, Archaeopterodactyloidea) from the Late Jurassic of Portugal. PeerJ. 11, e16048. Abstractfernandes_et_al_2023_lusognathus_peerj-16048.pdfWebsite

An incomplete, yet remarkably-sized dentated rostrum and associated partial cervical vertebrae of a pterosaur (ML 2554) were recently discovered from the Late Jurassic (Late Kimmeridgian-Early Tithonian) Lourinhã Formation of Praia do Caniçal, of central west Portugal. This specimen exhibits features such as a spatulated anterior expansion of the rostrum, robust comb-like dentition, and pronounced rims of the tooth alveoli, indicating gnathosaurine affinities. Based on its further unique tooth and dentary morphology, a new genus and species, \textit{Lusognathus almadrava} gen. et spec. nov., is proposed, making this the first named pterosaur species found within Portugal. The presence of this taxon adds yet another element to the fluvio-deltaic lagoonal environment that has been suggested as representative of the Lourinhã Formation in the Late Jurassic, further contributing to the diversity and distribution of gnathosaurines worldwide.